91 lines
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91 lines
11 KiB
XML
<document id="030072D72ABB2F6127FB8C14A25E0A69" ID-DOI="http://dx.doi.org/10.3897/zookeys.34.268" ID-GBIF-Dataset="7485373c-0219-4ed7-bb5c-ac664acf6775" checkinTime="1427905504904" checkinUser="donat" docAuthor="Alberto Ballerio & Munetoshi Maruyama" docDate="2010" docId="038951095C0F3D1466B6FA90AAA1FDF7" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="Zookeys 34" docTitle="Madrasostes clypeale Paulian 1993" docType="treatment" docVersion="9" lastPageNumber="84" masterDocId="FFB029715C083D1C663AFFC4A802FFD6" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageNumber="84" updateTime="1732486157858" updateUser="ExternalLinkService">
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<mods:mods id="3C7B0962138FBEC2272F10AA72C65A61" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2844BC7136B3C35B1B874C68CAE6B073">
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<mods:title id="240EC7125EF579797B8B935ABDDC7622">The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:name id="9D9A3E1ED6D900015F451D52984A918F" type="personal">
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<mods:role id="0CCAE3F55FD04E61FFF88620278AF916">
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<mods:roleTerm id="68183F507920C6FB15C2D4CD2A8001A7">Author</mods:roleTerm>
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<mods:namePart id="F193AEDDBFE330E5805A742EA999E436">Alberto Ballerio</mods:namePart>
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<mods:name id="2A9DFE843BC8C0D35C5EC64F61A167DE" type="personal">
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<mods:role id="750FF68D7E6D1E85A2E0E388638AF78F">
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<mods:roleTerm id="688C7808957B0120B521A5428954AB60">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2B19FD1FF1E2122DC2BB53B366A8F6E3">Munetoshi Maruyama</mods:namePart>
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<mods:typeOfResource id="2C4ABF4590DB4443CC912D28E9D6BF10">text</mods:typeOfResource>
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<mods:titleInfo id="8A826F733C6D53B6A490647AEFA8A900">
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<mods:title id="B045E43093F8D3B3A5237A851B2BF920">Zookeys</mods:title>
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</mods:titleInfo>
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<mods:part id="BB2CF8C62FEE5DEB6D388F38DAE4A2C8">
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<mods:date id="1C632BAC7F77776564BCF4A770FB3849">2010</mods:date>
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<mods:detail id="C155CF0E48004ECE94C3262A272D2FA3" type="volume">
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<mods:number id="52DFEE7F694890F5EDC94A58C4F90F79">34</mods:number>
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</mods:detail>
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<mods:extent id="DAB0BEC6F94A75C5D76E67DE1D51456E" unit="page">
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<mods:start id="E857A707E8D3E1DCEA39E4EAD0C8CBCA">77</mods:start>
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<mods:end id="2FE6EDA1DBF5065B586B584E3F7FE574">104</mods:end>
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<mods:classification id="4423367663B2DD8B7712CA870C5A40AA">journal article</mods:classification>
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<mods:identifier id="7EB04B815087406A0CC744488130C20A" type="DOI">http://dx.doi.org/10.3897/zookeys.34.268</mods:identifier>
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<treatment id="038951095C0F3D1466B6FA90AAA1FDF7" ID-DOI="http://doi.org/10.5281/zenodo.6210769" ID-GBIF-Taxon="114923677" ID-Zenodo-Dep="6210769" LSID="urn:lsid:plazi:treatment:038951095C0F3D1466B6FA90AAA1FDF7" httpUri="http://treatment.plazi.org/id/038951095C0F3D1466B6FA90AAA1FDF7" lastPageId="8" lastPageNumber="84" pageId="7" pageNumber="84" scope_family="Hybosoridae" scope_order="Coleoptera" scope_superFamily="Scarabaeoidea">
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<subSubSection id="C33AB3945C0F3D1B66B6FA90AB28FAB8" box="[140,810,1363,1390]" pageId="7" pageNumber="84" type="nomenclature">
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<paragraph id="8B9FE01F5C0F3D1B66B6FA90AB28FAB8" box="[140,810,1363,1390]" pageId="7" pageNumber="84">
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<taxonomicName id="4C209B9C5C0F3D1B66B6FA90AA2DFABB" box="[140,559,1363,1390]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="clypeale">
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<emphasis id="B9543C0D5C0F3D1B66B6FA90AA2DFABB" bold="true" box="[140,559,1363,1390]" pageId="7" pageNumber="84">
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<emphasis id="B9543C0D5C0F3D1B66B6FA90A983FAB8" bold="true" box="[140,385,1364,1390]" italics="true" pageId="7" pageNumber="84">Madrasostes clypeale</emphasis>
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Paulian, 1993
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</emphasis>
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</taxonomicName>
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(
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<figureCitation id="131BFC9A5C0F3D1B6404FA97AA83FAB8" box="[574,641,1363,1390]" captionStart="Figure 2" captionStartId="15.[140,211,1558,1580]" captionTargetBox="[140,1107,172,1515]" captionTargetId="figure@15.[140,1108,165,1521]" captionTargetPageId="15" captionText="Figure 2. Ceratocanthinae of Ulu Gombak. A Madrasostes depressum habitus dorsal B Madrasostes boucomonti habitus dorsal C Madrasostes agostii sp. n. habitus dorsal D Madrasostes clypeale male habitus dorsal E Madrasostes malyanum habitus dorsal F Madrasostes mirificum sp. n. habitus dorsal G habitus lateral of male H fore body of female I male protibia dorsal view J male protibia lateral view." pageId="7" pageNumber="84">Fig. 2</figureCitation>
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D,
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<figureCitation id="131BFC9A5C0F3D1B649CFA97AAE5FAB8" box="[678,743,1363,1390]" captionStart="Figure 6" captionStartId="21.[140,211,1206,1228]" captionTargetBox="[140,1107,165,1186]" captionTargetId="figure@21.[140,1108,165,1187]" captionTargetPageId="21" captionText="Figure 6. Photos of alive specimens taken in habitat at Ulu Gombak (photos © M. Maruyama, 2008). A Madrasostes variolosum B Madrasostes agostii C Madrasostes clypeale male D idem female E Eusphaeropeltis sp. b F Eusphaeropeltis sp. c." pageId="7" pageNumber="84">Fig. 6</figureCitation>
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C, D)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33AB3945C0F3D1B6683FAB3AB45FA47" box="[185,839,1398,1425]" pageId="7" pageNumber="84" type="description">
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<paragraph id="8B9FE01F5C0F3D1B6683FAB3AB45FA47" box="[185,839,1398,1425]" pageId="7" pageNumber="84">Known distribution: Indonesia (Sumatra) (Paulian 1993).</paragraph>
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</subSubSection>
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<subSubSection id="C33AB3945C0F3D146683FA5EAAA1FDF7" lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84" type="materials_examined">
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<paragraph id="8B9FE01F5C0F3D146683FA5EAAA1FDF7" lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84">
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New material examined: West Malaysia, Perak, 25 km NE of Ipoh, Banjaran Titi Wangsa Mounts, Gunung Korbu, 1400–1800 mt., 11–31.I.1999, leg. P. Cechovsky (ABCB); West Malaysia, Pearak, Penang Island, above botanical garden, 250 mt., 12.XI.1999, #4a, leg. Cuccodoro & Loebl, sifting forest leaf litter (MHNG, ABCB). Remarks. First record for West Malaysia (Perak and Selangor). The amazing sexual dimorphism of this species has already been described (Ballerio 2006). An examination of the microsculpturing of pronotum through scanning electron microscopy revealed that the pronotal punctures bottom surface is areolate (
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<figureCitation id="131BFC9A5C003D14656BFF62AB90FF17" box="[849,914,166,193]" captionStart="Figure 5" captionStartId="20.[140,211,1093,1115]" captionTargetBox="[140,1107,165,1070]" captionTargetId="figure@20.[140,1108,165,1073]" captionText="Figure 5. SEM photographs of: A Madrasostes clypeale: male pronotum B detail of male pronotum puncture C female pronotum D detail of female pronotum punctures E and F Madrasostes mirificum sp. n. mouthparts: labium G epipharynx H mandibles I detail of galear brush J maxilla." pageId="8" pageNumber="85">Fig. 5</figureCitation>
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A, B). The larger punctures of females have the areolate surface limited to a portion of the bottom and slightly raised (
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<figureCitation id="131BFC9A5C003D14670CFF29A97AFED1" box="[310,376,237,263]" captionStart="Figure 5" captionStartId="20.[140,211,1093,1115]" captionTargetBox="[140,1107,165,1070]" captionTargetId="figure@20.[140,1108,165,1073]" captionText="Figure 5. SEM photographs of: A Madrasostes clypeale: male pronotum B detail of male pronotum puncture C female pronotum D detail of female pronotum punctures E and F Madrasostes mirificum sp. n. mouthparts: labium G epipharynx H mandibles I detail of galear brush J maxilla." pageId="8" pageNumber="85">Fig. 5</figureCitation>
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C, D, E). Each puncture is often preceded by a short fine seta. It was not possible to detect any pore inside the punctures, so that the dirt which usually fills the punctures comes probably from outside. The species closest to
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<emphasis id="B9543C0D5C003D1465E3FEF0AC51FE98" box="[985,1107,307,334]" italics="true" pageId="8" pageNumber="85">M. clypeale</emphasis>
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is
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<taxonomicName id="4C209B9C5C003D14669DFE92AA0BFEA7" authority="Paulian, 1987" authorityName="Paulian" authorityYear="1987" box="[167,521,342,369]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="thai">
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<emphasis id="B9543C0D5C003D14669DFE92A95FFEA7" box="[167,349,342,369]" italics="true" pageId="8" pageNumber="85">Madrasostes thai</emphasis>
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Paulian, 1987
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</taxonomicName>
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, known only by the male holotype from Thailand (Chanthaburi province, well north of the isthmus of Kra, which is the boundary between the Sundaland and Indochinese subregions, see Corlett 2009). Both
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<emphasis id="B9543C0D5C003D1465E0FE5AAC51FE61" box="[986,1107,413,439]" italics="true" pageId="8" pageNumber="85">M. clypeale</emphasis>
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and
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<taxonomicName id="4C209B9C5C003D146684FE05A910FE0D" box="[190,274,448,475]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.bestMatchDistance="47" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="thai">
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<emphasis id="B9543C0D5C003D146684FE05A910FE0D" box="[190,274,448,475]" italics="true" pageId="8" pageNumber="85">M. thai</emphasis>
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</taxonomicName>
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represent a distinctive group within
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<taxonomicName id="4C209B9C5C003D146489FE04AB31FE0D" box="[691,819,448,475]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="genus">
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<emphasis id="B9543C0D5C003D146489FE04AB31FE0D" box="[691,819,448,475]" italics="true" pageId="8" pageNumber="85">Madrasostes</emphasis>
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</taxonomicName>
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and their current generic placement in the genus
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<taxonomicName id="4C209B9C5C003D1467A1FE27AA19FE28" box="[411,539,483,510]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="genus">
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<emphasis id="B9543C0D5C003D1467A1FE27AA19FE28" box="[411,539,483,510]" italics="true" pageId="8" pageNumber="85">Madrasostes</emphasis>
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</taxonomicName>
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must be regarded as provisional in the framework of a revision of the
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<taxonomicName id="4C209B9C5C003D14675CFDC3A9FBFDF7" box="[358,505,519,545]" class="Insecta" family="Hybosoridae" genus="Perignamptus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="genus">
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<emphasis id="B9543C0D5C003D14675CFDC3A9FBFDF7" box="[358,505,519,545]" italics="true" pageId="8" pageNumber="85">Perignamptus</emphasis>
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</taxonomicName>
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generic group.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |