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<document id="D35F6052BABBBA6E2C8559EE664004AE" ID-DOI="10.5281/zenodo.5329313" ID-ISSN="0374-1036" ID-Zenodo-Dep="5329313" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630162999482" checkinUser="felipe" docAuthor="Hisamatsu, Sadatomo" docDate="2011" docId="0388D148EA088738FE73FEE5FE556149" docLanguage="en" docName="ActaEntMusNatPra.2011.51.2.551-585.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 51 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Kateretidae Erichson 1846" docType="treatment" docVersion="4" lastPageNumber="555" masterDocId="FFB1A930EA0A873CFF97FFF6FFB1633A" masterDocTitle="A review of the Japanese Kateretidae fauna (Coleoptera: Cucujoidea)" masterLastPageNumber="585" masterPageNumber="551" pageNumber="553" updateTime="1699169862046" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A3A5ADBBBEF2FB7CEEC5DFE9655CE66D">A review of the Japanese Kateretidae fauna (Coleoptera: Cucujoidea)</mods:title>
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<mods:namePart id="5F9E823C132F528AD7204C8C176A345D">Hisamatsu, Sadatomo</mods:namePart>
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<treatment id="0388D148EA088738FE73FEE5FE556149" ID-DOI="http://doi.org/10.5281/zenodo.5397241" ID-Zenodo-Dep="5397241" LSID="urn:lsid:plazi:treatment:0388D148EA088738FE73FEE5FE556149" httpUri="http://treatment.plazi.org/id/0388D148EA088738FE73FEE5FE556149" lastPageId="4" lastPageNumber="555" pageId="2" pageNumber="553">
<subSubSection id="C33B33D5EA08873EFE73FEE5FCA86211" box="[484,793,275,299]" pageId="2" pageNumber="553" type="nomenclature">
<paragraph id="8B9E605EEA08873EFE73FEE5FCA86211" blockId="2.[484,793,275,299]" box="[484,793,275,299]" pageId="2" pageNumber="553">
<heading id="D0D6D732EA08873EFE73FEE5FCA86211" bold="true" box="[484,793,275,299]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="553" reason="3">
<taxonomicName id="4C211BDDEA08873EFE73FEE5FCA86211" ID-CoL="BL3" ID-ENA="295809" authority="Erichson, 1846" authorityName="Erichson" authorityYear="1846" box="[484,793,275,299]" class="Insecta" family="Kateretidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="family">
<emphasis id="B955BC4CEA08873EFE73FEE5FCA86211" bold="true" box="[484,793,275,299]" pageId="2" pageNumber="553">Kateretidae Erichson, 1846</emphasis>
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<subSubSection id="C33B33D5EA08873EFF00FEB7FC9E626E" box="[151,815,321,340]" pageId="2" pageNumber="553" type="type_taxon">
<paragraph id="8B9E605EEA08873EFF00FEB7FC9E626E" blockId="2.[151,815,321,340]" box="[151,815,321,340]" pageId="2" pageNumber="553">
<emphasis id="B955BC4CEA08873EFF00FEB7FEB2626E" bold="true" box="[151,259,321,340]" pageId="2" pageNumber="553">
<typeStatus id="549ADEFCEA08873EFF00FEB7FF75626E" box="[151,196,321,340]" pageId="2" pageNumber="553">Type</typeStatus>
genus.
</emphasis>
<taxonomicName id="4C211BDDEA08873EFE9FFEB4FE7C626E" ID-CoL="57PN" authority="Herbst, 1793" authorityName="Herbst" authorityYear="1793" box="[264,461,321,340]" class="Insecta" family="Kateretidae" genus="Kateretes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA08873EFE9FFEB4FEE8626E" box="[264,345,322,340]" italics="true" pageId="2" pageNumber="553">Kateretes</emphasis>
<bibRefCitation id="EFB01DAFEA08873EFEC9FEB7FE7C626E" author="HERBST J. F. W." box="[350,461,321,340]" pageId="2" pageNumber="553" refId="ref18722" refString="HERBST J. F. W. 1793. Natursystem aller bekannten in- und auslandischen Insecten, als eine Fortsetzung der von Buffonschen Naturgeschichte. Der Kafer vierte Theil. Paulischen Buchhandlung, Berlin, xvi + 392 pp., 16 pls." type="book" year="1793">Herbst, 1793</bibRefCitation>
</taxonomicName>
(subsequent designation by
<bibRefCitation id="EFB01DAFEA08873EFD28FEB7FC94626E" author="ICZN" box="[703,805,321,340]" pageId="2" pageNumber="553" pagination="82 - 86" refId="ref19159" refString="ICZN 1999: Opinion 1916. Brachypterinae Zwick, 1973 (Insecta, Plecoptera): spelling emended to Brachypterainae, so removing the homonymy with Brachypterinae Erichson, [1845] (Insecta, Coleoptera); Kateretidae Erichson in Agassiz, [1846]: given precedence over Brachypterinae Erichson. Bulletin of Zoological Nomenclature 56: 82 - 86." type="journal article" year="1999">ICZN 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33B33D5EA08873FFF00FE9AFDD662A9" lastPageId="3" lastPageNumber="554" pageId="2" pageNumber="553" type="diagnosis">
<paragraph id="8B9E605EEA08873FFF00FE9AFC936209" blockId="2.[151,1127,364,453]" lastBlockId="3.[113,1089,219,500]" lastPageId="3" lastPageNumber="554" pageId="2" pageNumber="553">
<emphasis id="B955BC4CEA08873EFF00FE9AFEBD62BE" bold="true" box="[151,268,364,388]" pageId="2" pageNumber="553">Diagnosis.</emphasis>
Length 1.3–6.0 mm. Head prognathous; subantennal grooves and subantennal ridges absent; frontoclypeal suture short or absent; maxilla with galea and lacinia; antennae 11-jointed, last 2 or 3 segments forming a loose club (without club in
<taxonomicName id="4C211BDDEA08873FFCEAFE58FF1B63CE" authority="Kirejtshuk, 1989" authorityName="Kirejtshuk" authorityYear="1989" class="Insecta" family="Kateretidae" genus="Sibirhelus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="3" lastPageNumber="554" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA08873EFCEAFE58FC5862FF" box="[893,1001,430,453]" italics="true" pageId="2" pageNumber="553">Sibirhelus</emphasis>
<bibRefCitation id="EFB01DAFEA08873FFC78FE5AFF1B63CE" author="KIREJTSHUK A. G." lastPageId="3" lastPageNumber="554" pageId="2" pageNumber="553" pagination="145 - 149" refId="ref19515" refString="KIREJTSHUK A. G. 1989: Novye palearkticheskie rod i vidy sem. Kateretidae (Coleoptera) i zamechaniya po sinonimii. (New Palaearctic genus and species of the family Kateretidae (Coleoptera) and notes on the synonymy). Zoologicheskii Zhurnal 68: 145 - 149 (in Russian, English title and summary)." type="journal article" year="1989">Kirejtshuk, 1989</bibRefCitation>
</taxonomicName>
). Procoxal cavities incompletely closed posteriorly. Male genitalia with separated parameres; aedeagus asymmetrical. Larvae without urogomphi and preogomphi (see
<bibRefCitation id="EFB01DAFEA09873FFC40FF0AFF58620E" author="JELINEK J. &amp; CLINE A. R." pageId="3" pageNumber="554" pagination="386 - 390" refId="ref19386" refString="JELINEK J. &amp; CLINE A. R. 2010: Chapter 10.25. Kateretidae Erichson in Agassiz, 1846. Pp. 386 - 390. In: LESCHEN R. A. B., BEUTEL R. G. &amp; LAWRENCE J. F. (eds.): Handbook of Zoology, Arthropoda: Insecta, Coleoptera, Beetles Morphology and Systematics. De Gruyter, Berlin, New York, 786 pp." type="book chapter" year="2010">JELÍNEK &amp; CLINE 2010</bibRefCitation>
for detailed diagnostic features of adults and larvae).
</paragraph>
<caption id="DF5E30D6EA08873EFF00FA24FCAB650F" ID-DOI="http://doi.org/10.5281/zenodo.5329315" ID-Zenodo-Dep="5329315" httpUri="https://zenodo.org/record/5329315/files/figure.png" pageId="2" pageNumber="553" startId="2.[151,182,1490,1509]" targetBox="[275,997,488,1467]" targetPageId="2">
<paragraph id="8B9E605EEA08873EFF00FA24FCAB650F" blockId="2.[151,1126,1490,1589]" pageId="2" pageNumber="553">
Fig. 1. Habitus and labels of
<taxonomicName id="4C211BDDEA08873EFE07FA24FE4366DF" authorityName="Erichson" authorityYear="1846" box="[400,498,1490,1509]" class="Insecta" family="Kateretidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="family">Kateretidae</taxonomicName>
spp. A–B –
<taxonomicName id="4C211BDDEA08873EFDC8FA25FC1066DF" authority="Hisamatsu, 1985" authorityName="Hisamatsu" authorityYear="1985" box="[607,929,1490,1509]" class="Insecta" family="Kateretidae" genus="Kateretes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B955BC4CEA08873EFDC8FA25FCB966DF" box="[607,776,1491,1509]" italics="true" pageId="2" pageNumber="553">Kateretes japonicus</emphasis>
<bibRefCitation id="EFB01DAFEA08873EFC99FA24FC1066DF" author="HISAMATSU S." box="[782,929,1490,1509]" pageId="2" pageNumber="553" pagination="28 - 31" refId="ref18861" refString="HISAMATSU S. 1985: Nitidulidae Latreille, 1802. Pp. 174 - 197, pls. 28 - 31. In: KUROSAWA Y., HISAMATSU S. &amp; SASAJI H. (eds.): The Coleoptera of Japan in Color, 3. Hoikusha, Osaka, x + 500 incl. 72 pls (in Japanese)." type="book chapter" year="1985">Hisamatsu, 1985</bibRefCitation>
</taxonomicName>
: A – holotype, male; B – ditto, labels; C–D –
<taxonomicName id="4C211BDDEA08873EFEDDFA18FE66653A" authorityName="S-T.Hisamatsu" authorityYear="2006" box="[330,471,1518,1536]" class="Insecta" family="Kateretidae" genus="Kateretes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="takagii">
<emphasis id="B955BC4CEA08873EFEDDFA18FE66653A" box="[330,471,1518,1536]" italics="true" pageId="2" pageNumber="553">Kateretes takagii</emphasis>
</taxonomicName>
S-T.
<bibRefCitation id="EFB01DAFEA08873EFD95FA1BFD20653A" author="HISAMATSU S-T." box="[514,657,1517,1536]" pageId="2" pageNumber="553" pagination="243 - 247" refId="ref18957" refString="HISAMATSU S-T. 2006: A new species of the genus Kateretes Herbst (Coleoptera, Kateretidae) from Japan. Japanese Journal of Systematic Entomology 12: 243 - 247." type="journal article" year="2006">Hisamatsu, 2006</bibRefCitation>
:C – holotype, male; D – ditto, labels; E–F –
<taxonomicName id="4C211BDDEA08873EFC69FA18FED76520" authority="Reitter, 1892" authorityName="Reitter" authorityYear="1892" class="Insecta" family="Kateretidae" genus="Platamartus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="jakowlewi">
<emphasis id="B955BC4CEA08873EFC69FA18FF5F6520" italics="true" pageId="2" pageNumber="553">Platamartus jakowlewi</emphasis>
<bibRefCitation id="EFB01DAFEA08873EFF63F9F1FED76520" author="REITTER E." box="[244,358,1543,1562]" pageId="2" pageNumber="553" pagination="151 - 154" refId="ref20576" refString="REITTER E. 1892: Siebenter Beitrag zur Coleopterenfauna des russischen Reiches. Wiener Entomologische Zeitung 11: 151 - 154." type="journal article" year="1892">Reitter, 1892</bibRefCitation>
</taxonomicName>
: E – male and F – female (both Tochigi Pref.); G–H –
<taxonomicName id="4C211BDDEA08873EFCA8F9FEFEA6650F" authority="(Reitter, 1878)" baseAuthorityName="Reitter" baseAuthorityYear="1878" class="Insecta" family="Kateretidae" genus="Heterhelus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="japonicus" subGenus="Heterhelus">
<emphasis id="B955BC4CEA08873EFCA8F9FEFC2D6520" box="[831,924,1544,1562]" italics="true" pageId="2" pageNumber="553">Heterhelus</emphasis>
(
<emphasis id="B955BC4CEA08873EFC3EF9FEFBB76520" box="[937,1030,1544,1562]" italics="true" pageId="2" pageNumber="553">Heterhelus</emphasis>
)
<emphasis id="B955BC4CEA08873EFB84F9FEFBD76520" box="[1043,1126,1544,1562]" italics="true" pageId="2" pageNumber="553">japonicus</emphasis>
(
<bibRefCitation id="EFB01DAFEA08873EFF0AF9D4FEA1650F" author="REITTER E." box="[157,272,1570,1589]" pageId="2" pageNumber="553" pagination="165 - 194" refId="ref20493" refString="REITTER E. 1878: Coleopterorum species novae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch Botanischen Gesellschaft in Wien 27: 165 - 194." type="journal article" year="1878">Reitter, 1878</bibRefCitation>
)
</taxonomicName>
: G – lectotype, male; H – ditto, labels. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B9E605EEA09873FFFE6FECDFDD662A9" blockId="3.[113,1089,219,500]" pageId="3" pageNumber="554">
<emphasis id="B955BC4CEA09873FFFE6FECDFF5E6269" bold="true" box="[113,239,315,339]" pageId="3" pageNumber="554">Bionomics.</emphasis>
Both larval and adult
<taxonomicName id="4C211BDDEA09873FFE71FECDFDD06269" authorityName="Erichson" authorityYear="1846" box="[486,609,315,339]" class="Insecta" family="Kateretidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="family">Kateretidae</taxonomicName>
are anthophagous, developing in flowers of angiosperms (
<bibRefCitation id="EFB01DAFEA09873FFE9FFEAAFE4A624E" author="JELINEK J. &amp; CLINE A. R." box="[264,507,348,372]" pageId="3" pageNumber="554" pagination="386 - 390" refId="ref19386" refString="JELINEK J. &amp; CLINE A. R. 2010: Chapter 10.25. Kateretidae Erichson in Agassiz, 1846. Pp. 386 - 390. In: LESCHEN R. A. B., BEUTEL R. G. &amp; LAWRENCE J. F. (eds.): Handbook of Zoology, Arthropoda: Insecta, Coleoptera, Beetles Morphology and Systematics. De Gruyter, Berlin, New York, 786 pp." type="book chapter" year="2010">JELÍNEK &amp; CLINE 2010</bibRefCitation>
). Larvae are monophagous or oligophagous, typically requiring specific plants for their development.
</paragraph>
</subSubSection>
<subSubSection id="C33B33D5EA098738FFE6FE6DFE556149" lastPageId="4" lastPageNumber="555" pageId="3" pageNumber="554" type="distribution">
<paragraph id="8B9E605EEA098738FFE6FE6DFCF4628E" blockId="3.[113,1089,219,500]" lastBlockId="4.[151,1127,219,628]" lastPageId="4" lastPageNumber="555" pageId="3" pageNumber="554">
<emphasis id="B955BC4CEA09873FFFE6FE6DFEB56289" bold="true" box="[113,260,411,435]" pageId="3" pageNumber="554">Distribution.</emphasis>
<taxonomicName id="4C211BDDEA09873FFE9DFE6DFE346289" authorityName="Erichson" authorityYear="1846" box="[266,389,411,435]" class="Insecta" family="Kateretidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="family">Kateretidae</taxonomicName>
comprises about 95 species within 14 genera worldwide (
<bibRefCitation id="EFB01DAFEA09873FFC65FE6AFEBA62EE" author="JELINEK J. &amp; CLINE A. R." pageId="3" pageNumber="554" pagination="386 - 390" refId="ref19386" refString="JELINEK J. &amp; CLINE A. R. 2010: Chapter 10.25. Kateretidae Erichson in Agassiz, 1846. Pp. 386 - 390. In: LESCHEN R. A. B., BEUTEL R. G. &amp; LAWRENCE J. F. (eds.): Handbook of Zoology, Arthropoda: Insecta, Coleoptera, Beetles Morphology and Systematics. De Gruyter, Berlin, New York, 786 pp." type="book chapter" year="2010">JELÍNEK &amp; CLINE 2010</bibRefCitation>
). There are two described fossil genera,
<taxonomicName id="4C211BDDEA09873FFD29FE4BFBF162EE" authority="Kirejtshuk &amp; Nel, 2008" authorityName="Kirejtshuk &amp; Nel" authorityYear="2008" box="[702,1088,443,468]" class="Insecta" family="Kateretidae" genus="Eoceniretes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA09873FFD29FE4BFC8A62EE" box="[702,827,445,468]" italics="true" pageId="3" pageNumber="554">Eoceniretes</emphasis>
<bibRefCitation id="EFB01DAFEA09873FFCD5FE4DFBF162EE" author="KIREJTSHUK A. G. &amp; NEL A." box="[834,1088,443,468]" pageId="3" pageNumber="554" pagination="419 - 442" refId="ref19686" refString="KIREJTSHUK A. G. &amp; NEL A. 2008: New beetles of the suborder Polyphaga from the Lowermost Eocene French amber (Insecta: Coleoptera). Annales de la Societe Entomologique de France, Nouvelle Serie 44: 419 - 442." type="journal article" year="2008">Kirejtshuk &amp; Nel, 2008</bibRefCitation>
</taxonomicName>
from Lowermost Eocene French amber and
<taxonomicName id="4C211BDDEA09873FFDF6FE2BFBB362CE" authority="Kirejtshuk &amp; Azar, 2008" authorityName="Kirejtshuk &amp; Azar" authorityYear="2008" box="[609,1026,475,500]" class="Insecta" family="Kateretidae" genus="Lebanoretes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA09873FFDF6FE2BFD5462CE" box="[609,741,477,500]" italics="true" pageId="3" pageNumber="554">Lebanoretes</emphasis>
<bibRefCitation id="EFB01DAFEA09873FFD78FE2DFBB362CE" author="KIREJTSHUK A. G. &amp; AZAR D." box="[751,1026,475,500]" pageId="3" pageNumber="554" pagination="15 - 46" refId="ref19650" refString="KIREJTSHUK A. G. &amp; AZAR D. 2008: New taxa of beetles (Insecta, Coleoptera) from Lebanese amber with evolutionary and systematic comments. Alavesia 2: 15 - 46." type="journal article" year="2008">Kirejtshuk &amp; Azar, 2008</bibRefCitation>
</taxonomicName>
from Lower Cretaceous Lebanese amber (
<bibRefCitation id="EFB01DAFEA0E8738FDBEFF2AFCF063CE" author="KIREJTSHUK A. G. &amp; AZAR D." box="[553,833,220,244]" pageId="4" pageNumber="555" pagination="15 - 46" refId="ref19650" refString="KIREJTSHUK A. G. &amp; AZAR D. 2008: New taxa of beetles (Insecta, Coleoptera) from Lebanese amber with evolutionary and systematic comments. Alavesia 2: 15 - 46." type="journal article" year="2008">KIREJTSHUK &amp; AZAR 2008</bibRefCitation>
,
<bibRefCitation id="EFB01DAFEA0E8738FCD8FF2AFBEA63CE" author="KIREJTSHUK A. G. &amp; NEL A." box="[847,1115,220,244]" pageId="4" pageNumber="555" pagination="419 - 442" refId="ref19686" refString="KIREJTSHUK A. G. &amp; NEL A. 2008: New beetles of the suborder Polyphaga from the Lowermost Eocene French amber (Insecta: Coleoptera). Annales de la Societe Entomologique de France, Nouvelle Serie 44: 419 - 442." type="journal article" year="2008">KIREJTSHUK &amp; NEL 2008</bibRefCitation>
). Members of Katereridae are mainly distributed in the Holarctic Region, but some taxa are distributed in other geographical regions – Neotropical Region:
<taxonomicName id="4C211BDDEA0E8738FCA5FEEBFB8A6209" authority="Reitter, 1875" authorityName="Reitter" authorityYear="1875" box="[818,1083,283,308]" class="Insecta" family="Nitidulidae" genus="Cercometes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA0E8738FCA5FEEBFC1F620E" box="[818,942,285,308]" italics="true" pageId="4" pageNumber="555">Cercometes</emphasis>
<bibRefCitation id="EFB01DAFEA0E8738FC24FEEDFB8A6209" author="REITTER E." box="[947,1083,283,307]" pageId="4" pageNumber="555" pagination="1 - 88" refId="ref20461" refString="REITTER E. 1875: Die europaischen Nitidularien mit kurzer Charakteristik der Gattungen und Bemerkungen uber schwierige Arten verzeichnet. Deutsche Entomologische Zeitschrift 19 (3): 1 - 88." type="journal article" year="1875">Reitter, 1875</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C211BDDEA0E8738FF00FECBFE59626E" authority="Jelinek, 1979" authorityName="Jelinek" authorityYear="1979" box="[151,488,315,340]" class="Insecta" family="Kateretidae" genus="Neobrachypterus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA0E8738FF00FECBFEFE626E" box="[151,335,317,340]" italics="true" pageId="4" pageNumber="555">Neobrachypterus</emphasis>
Jelínek, 1979
</taxonomicName>
; Oriental Region: some species of
<taxonomicName id="4C211BDDEA0E8738FCF6FECBFF7E624E" authority="Kugelann, 1794" authorityName="Kugelann" authorityYear="1794" class="Insecta" family="Kateretidae" genus="Brachypterus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA0E8738FCF6FECBFC40626E" box="[865,1009,317,340]" italics="true" pageId="4" pageNumber="555">Brachypterus</emphasis>
Kugelann, 1794
</taxonomicName>
,
<taxonomicName id="4C211BDDEA0E8738FF4BFEABFDE26249" authority="Jacquelin du Val, 1858" authorityName="Jacquelin du Val" authorityYear="1858" box="[220,595,347,372]" class="Insecta" family="Kateretidae" genus="Heterhelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA0E8738FF4BFEABFEE0624E" box="[220,337,349,372]" italics="true" pageId="4" pageNumber="555">Heterhelus</emphasis>
Jacquelin du Val, 1858
</taxonomicName>
, and
<taxonomicName id="4C211BDDEA0E8738FD18FEABFC39624E" authority="Herbst, 1793" authorityName="Herbst" authorityYear="1793" box="[655,904,347,372]" class="Insecta" family="Kateretidae" genus="Kateretes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA0E8738FD18FEABFD45624E" box="[655,756,349,372]" italics="true" pageId="4" pageNumber="555">Kateretes</emphasis>
<bibRefCitation id="EFB01DAFEA0E8738FD6BFEADFC39624E" author="HERBST J. F. W." box="[764,904,347,372]" pageId="4" pageNumber="555" refId="ref18722" refString="HERBST J. F. W. 1793. Natursystem aller bekannten in- und auslandischen Insecten, als eine Fortsetzung der von Buffonschen Naturgeschichte. Der Kafer vierte Theil. Paulischen Buchhandlung, Berlin, xvi + 392 pp., 16 pls." type="book" year="1793">Herbst, 1793</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C211BDDEA0E8738FC0EFEABFF6162AE" authority="Jelinek, 1978" authorityName="Jelinek" authorityYear="1978" class="Insecta" family="Kateretidae" genus="Kateretes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="B955BC4CEA0E8738FC0EFEABFBBA624E" box="[921,1035,349,372]" italics="true" pageId="4" pageNumber="555">K. ornatus</emphasis>
<bibRefCitation id="EFB01DAFEA0E8738FB83FEADFF6162AE" author="JELINEK J." pageId="4" pageNumber="555" pagination="171 - 218" refId="ref19251" refString="JELINEK J. 1978: Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Nitidulidae. Entomologica Basiliensia 3: 171 - 218." type="journal article" year="1978">Jelínek, 1978</bibRefCitation>
</taxonomicName>
); and Australian Region:
<taxonomicName id="4C211BDDEA0E8738FE74FE8BFCAB62AE" authority="Kirejtshuk, 2000" authorityName="Kirejtshuk" authorityYear="2000" box="[483,794,379,404]" class="Insecta" family="Kateretidae" genus="Jelinekiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA0E8738FE74FE8BFDEF62AE" box="[483,606,381,404]" italics="true" pageId="4" pageNumber="555">Jelinekiella</emphasis>
Kirejtshuk, 2000
</taxonomicName>
(
<taxonomicName id="4C211BDDEA0E8738FCBDFE8BFBEE62A9" authority="Kirejtshuk, 1986" authorityName="Kirejtshuk" authorityYear="1986" box="[810,1119,379,404]" class="Insecta" family="Kateretidae" genus="Jelinekiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B955BC4CEA0E8738FCBDFE8BFC1062AE" box="[810,929,381,404]" italics="true" pageId="4" pageNumber="555">J. aterrima</emphasis>
Kirejtshuk, 1986
</taxonomicName>
) and
<taxonomicName id="4C211BDDEA0E8738FF53FE6BFD8C628E" authority="Blackburn, 1892" authorityName="Blackburn" authorityYear="1892" box="[196,573,411,436]" class="Insecta" family="Kateretidae" genus="Notobrachypterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="B955BC4CEA0E8738FF53FE6BFE34628E" box="[196,389,413,436]" italics="true" pageId="4" pageNumber="555">Notobrachypterus</emphasis>
Blackburn, 1892
</taxonomicName>
(
<bibRefCitation id="EFB01DAFEA0E8738FDDBFE6AFC8B628E" author="JELINEK J. &amp; CLINE A. R." box="[588,826,412,436]" pageId="4" pageNumber="555" pagination="386 - 390" refId="ref19386" refString="JELINEK J. &amp; CLINE A. R. 2010: Chapter 10.25. Kateretidae Erichson in Agassiz, 1846. Pp. 386 - 390. In: LESCHEN R. A. B., BEUTEL R. G. &amp; LAWRENCE J. F. (eds.): Handbook of Zoology, Arthropoda: Insecta, Coleoptera, Beetles Morphology and Systematics. De Gruyter, Berlin, New York, 786 pp." type="book chapter" year="2010">JELÍNEK &amp; CLINE 2010</bibRefCitation>
).
</paragraph>
<caption id="DF5E30D6EA09873FFFE6FA26FC1D650E" ID-DOI="http://doi.org/10.5281/zenodo.5329319" ID-Zenodo-Dep="5329319" httpUri="https://zenodo.org/record/5329319/files/figure.png" pageId="3" pageNumber="554" startId="3.[113,144,1488,1507]" targetBox="[232,958,532,1461]" targetPageId="3">
<paragraph id="8B9E605EEA09873FFFE6FA26FC1D650E" blockId="3.[113,1089,1488,1588]" pageId="3" pageNumber="554">
Fig. 2. Habitus and labels of
<taxonomicName id="4C211BDDEA09873FFEFEFA26FE7A66D9" authorityName="Erichson" authorityYear="1846" box="[361,459,1488,1507]" class="Insecta" family="Kateretidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="family">Kateretidae</taxonomicName>
spp. A–B –
<taxonomicName id="4C211BDDEA09873FFDA1FA27FC7366D9" authority="(Reitter, 1878)" baseAuthorityName="Reitter" baseAuthorityYear="1878" box="[566,962,1488,1507]" class="Insecta" family="Kateretidae" genus="Heterhelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="morio" subGenus="Heterhelus">
<emphasis id="B955BC4CEA09873FFDA1FA27FD2266D9" box="[566,659,1489,1507]" italics="true" pageId="3" pageNumber="554">Heterhelus</emphasis>
(
<emphasis id="B955BC4CEA09873FFD37FA27FD4C66D9" box="[672,765,1489,1507]" italics="true" pageId="3" pageNumber="554">Heterhelus</emphasis>
)
<emphasis id="B955BC4CEA09873FFC9EFA27FC8D66D9" box="[777,828,1489,1507]" italics="true" pageId="3" pageNumber="554">morio</emphasis>
(
<bibRefCitation id="EFB01DAFEA09873FFCDFFA26FC0A66D9" author="REITTER E." box="[840,955,1488,1507]" pageId="3" pageNumber="554" pagination="165 - 194" refId="ref20493" refString="REITTER E. 1878: Coleopterorum species novae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch Botanischen Gesellschaft in Wien 27: 165 - 194." type="journal article" year="1878">Reitter, 1878</bibRefCitation>
)
</taxonomicName>
: A – lectotype, male; B – ditto, labels; C –
<taxonomicName id="4C211BDDEA09873FFEC3FA1AFD0C66C4" authority="(Heer, 1841)" baseAuthorityName="Heer" baseAuthorityYear="1841" box="[340,701,1515,1534]" class="Insecta" family="Kateretidae" genus="Heterhelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="solani" subGenus="Boreades">
<emphasis id="B955BC4CEA09873FFEC3FA1AFE0066C4" box="[340,433,1516,1534]" italics="true" pageId="3" pageNumber="554">Heterhelus</emphasis>
(
<emphasis id="B955BC4CEA09873FFE2AFA1AFDBC66C4" box="[445,525,1516,1534]" italics="true" pageId="3" pageNumber="554">Boreades</emphasis>
)
<emphasis id="B955BC4CEA09873FFD8FFA1AFDFD66C4" box="[536,588,1516,1534]" italics="true" pageId="3" pageNumber="554">solani</emphasis>
(
<bibRefCitation id="EFB01DAFEA09873FFDC1FA1DFD0766C4" author="HEER O." box="[598,694,1515,1534]" pageId="3" pageNumber="554" pagination="361 - 652" refId="ref18647" refString="HEER O. 1841: Fauna Coleopterorum Helvetica. Pars 1., fasciculus 3. Impensis Orelii, Fuesslini et Sociorum, Turici, 361 - 652 pp." type="book chapter" year="1841">Heer, 1841</bibRefCitation>
)
</taxonomicName>
, male (Hokkaido); D –
<taxonomicName id="4C211BDDEA09873FFC16FA1AFF416523" authority="(Reitter, 1900)" baseAuthorityName="Reitter" baseAuthorityYear="1900" class="Insecta" family="Kateretidae" genus="Sibirhelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B955BC4CEA09873FFC16FA1AFBF066C4" box="[897,1089,1516,1534]" italics="true" pageId="3" pageNumber="554">Sibirhelus corpulentus</emphasis>
(
<bibRefCitation id="EFB01DAFEA09873FFFE0F9F3FF586523" author="REITTER E." box="[119,233,1541,1561]" pageId="3" pageNumber="554" pagination="225 - 232" refId="ref20598" refString="REITTER E. 1900: Beschreibung und Abbildung von neun Coleopteren der palaearctischen Fauna. Wiener Entomologische Zeitung 19: 225 - 232, pl. 2." type="journal article" year="1900">Reitter, 1900</bibRefCitation>
)
</taxonomicName>
, male (Iwate Pref.); E –
<taxonomicName id="4C211BDDEA09873FFE56F9F0FCA16523" authority="(Fabricius, 1792)" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[449,784,1541,1561]" class="Insecta" family="Kateretidae" genus="Brachypterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="urticae">
<emphasis id="B955BC4CEA09873FFE56F9F0FDC76522" box="[449,630,1542,1560]" italics="true" pageId="3" pageNumber="554">Brachypterus urticae</emphasis>
(
<bibRefCitation id="EFB01DAFEA09873FFD16F9F3FCBB6523" author="FABRICIUS J. C." box="[641,778,1541,1561]" pageId="3" pageNumber="554" refId="ref18075" refString="FABRICIUS J. C. 1792: Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonymis, locis, descriptionibus, observationibus. Tom I. Pars I. Christ. Gottl. Proft, Hafniae, xx + 330 pp." type="book" year="1792">Fabricius, 1792</bibRefCitation>
)
</taxonomicName>
, male (Hokkaido); F–G –
<taxonomicName id="4C211BDDEA09873FFC66F9F0FE016509" authority="Hisamatsu, 1985" authorityName="Hisamatsu" authorityYear="1985" class="Insecta" family="Kateretidae" genus="Brachypterolus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="shimoyamai">
<emphasis id="B955BC4CEA09873FFC66F9F0FEA96509" italics="true" pageId="3" pageNumber="554">Brachypterolus shimoyamai</emphasis>
<bibRefCitation id="EFB01DAFEA09873FFE89F9D6FE016509" author="HISAMATSU S." box="[286,432,1568,1587]" pageId="3" pageNumber="554" pagination="28 - 31" refId="ref18861" refString="HISAMATSU S. 1985: Nitidulidae Latreille, 1802. Pp. 174 - 197, pls. 28 - 31. In: KUROSAWA Y., HISAMATSU S. &amp; SASAJI H. (eds.): The Coleoptera of Japan in Color, 3. Hoikusha, Osaka, x + 500 incl. 72 pls (in Japanese)." type="book chapter" year="1985">Hisamatsu, 1985</bibRefCitation>
</taxonomicName>
: F – holotype, male; G – ditto, labels. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B9E605EEA0E8738FF00FE4AFE556149" blockId="4.[151,1127,219,628]" pageId="4" pageNumber="555">
<emphasis id="B955BC4CEA0E8738FF00FE4AFF6262EE" bold="true" box="[151,211,444,468]" pageId="4" pageNumber="555">Note.</emphasis>
Detailed descriptions of the family were recently provided (e.g.,
<bibRefCitation id="EFB01DAFEA0E8738FC3DFE4AFBD362EE" author="KIREJTSHUK A. G." box="[938,1122,444,468]" pageId="4" pageNumber="555" pagination="210 - 216" refId="ref19575" refString="KIREJTSHUK A. G. 1992: Family Kateretidae Erichson, 1846. Pp. 210 - 216. In: LEHR P. A. (ed.): Opredelitel' nasekomykh Dal'nego Vostoka SSSR. [Identification key to the insects of the Far East of the USSR. Volume 3. Part 2.]. Nauka Pren, St. Petersburg, Russia, 704 pp (in Russian)." type="book chapter" year="1992">KIREJTSHUK 1992</bibRefCitation>
,
<bibRefCitation id="EFB01DAFEA0E8738FF00FE2AFE9962CE" author="AUDISIO P." box="[151,296,476,500]" pageId="4" pageNumber="555" refId="ref17691" refString="AUDISIO P. 1993: Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. Vol. 32. Calderini Edizione, Bologna, xvi + 971 pp." type="book" year="1993">AUDISIO 1993</bibRefCitation>
,
<bibRefCitation id="EFB01DAFEA0E8738FEA5FE2AFDAF62CE" author="JELINEK J. &amp; CLINE A. R." box="[306,542,476,500]" pageId="4" pageNumber="555" pagination="386 - 390" refId="ref19386" refString="JELINEK J. &amp; CLINE A. R. 2010: Chapter 10.25. Kateretidae Erichson in Agassiz, 1846. Pp. 386 - 390. In: LESCHEN R. A. B., BEUTEL R. G. &amp; LAWRENCE J. F. (eds.): Handbook of Zoology, Arthropoda: Insecta, Coleoptera, Beetles Morphology and Systematics. De Gruyter, Berlin, New York, 786 pp." type="book chapter" year="2010">JELÍNEK &amp; CLINE 2010</bibRefCitation>
). There are some comprehensive studies on Palaearctic fauna (KIRK- SPRIGGS 1996,
<bibRefCitation id="EFB01DAFEA0E8738FE57FE0AFDC06129" author="SPORNRAFT K." box="[448,625,507,532]" pageId="4" pageNumber="555" pagination="20 - 77" refId="ref20983" refString="SPORNRAFT K. 1967: 50 - Familie Nitidulidae. Pp. 20 - 77. In: FREUDE H., HARDE K. W. &amp; LOHSE G. A. (eds.): Die Kafer Mitteleuropas. Vol. 7. Goecke &amp; Evers Verlag, Krefeld, 310 pp." type="book chapter" year="1967">SPORNRAFT 1967</bibRefCitation>
,
<bibRefCitation id="EFB01DAFEA0E8738FDE9FE0AFC87612E" author="KIREJTSHUK A. G." box="[638,822,508,532]" pageId="4" pageNumber="555" pagination="210 - 216" refId="ref19575" refString="KIREJTSHUK A. G. 1992: Family Kateretidae Erichson, 1846. Pp. 210 - 216. In: LEHR P. A. (ed.): Opredelitel' nasekomykh Dal'nego Vostoka SSSR. [Identification key to the insects of the Far East of the USSR. Volume 3. Part 2.]. Nauka Pren, St. Petersburg, Russia, 704 pp (in Russian)." type="book chapter" year="1992">KIREJTSHUK 1992</bibRefCitation>
,
<bibRefCitation id="EFB01DAFEA0E8738FCD5FE0AFC67612E" author="AUDISIO P." box="[834,982,508,532]" pageId="4" pageNumber="555" refId="ref17691" refString="AUDISIO P. 1993: Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. Vol. 32. Calderini Edizione, Bologna, xvi + 971 pp." type="book" year="1993">AUDISIO 1993</bibRefCitation>
), and a comprehensive treatment of the New World fauna is currently underway (Cline &amp; Audisio, pers. comm. 2010). Some larvae were illustrated by
<bibRefCitation id="EFB01DAFEA0E8738FD14FDCAFC02616E" author="BOVING A. G. &amp; CRAIGHEAD F. C." box="[643,947,572,596]" pageId="4" pageNumber="555" pagination="1 - 351" refId="ref17898" refString="BOVING A. G. &amp; CRAIGHEAD F. C. 1931: An illustrated synopsis of the principal larval forms of the order Coleoptera. Entomologica Americana 11: 1 - 351." type="journal article" year="1931">BÖVING &amp; CRAIGHEAD (1931)</bibRefCitation>
,
<bibRefCitation id="EFB01DAFEA0E8738FC2AFDCAFBD3616E" author="HAYASHI N." box="[957,1122,572,596]" pageId="4" pageNumber="555" pagination="1 - 97" refId="ref18610" refString="HAYASHI N. 1978: A contribution to the knowledge of the larvae of Nitidulidae occurring in Japan (Coleoptera: Cucujoidea). Insecta Matsumurana, N. S. 14: 1 - 97." type="book chapter" year="1978">HAYASHI (1978)</bibRefCitation>
, and CLINE &amp; AUDISIO (unpubl.).
</paragraph>
</subSubSection>
</treatment>
</document>
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