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<document id="D81848B84437F38A18AF06A762226917" ID-DOI="10.15298/rusentj.30.3.15" ID-ISSN="0132-8069" ID-Zenodo-Dep="13178900" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722594922894" checkinUser="felipe" docAuthor="Krivosheina, M. G. &amp; Ozerov, A. L." docDate="2021" docId="0388A40719008C4D5BF1B03DFA8FC698" docLanguage="en" docName="RussEntomolJ.30.3.345-360.pdf" docOrigin="Russian Entomological Journal 30 (3)" docSource="http://dx.doi.org/10.15298/rusentj.30.3.15" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Ephydra villosa Krivosheina et Ozerov 2021, sp.n." docType="treatment" docVersion="2" lastPageNumber="358" masterDocId="FFB1DC7F190D8C405B13B76BFF9DC336" masterDocTitle="A review of the shore-fly genus Ephydra Fallén, 1810 (Diptera: Ephydridae) of Russia" masterLastPageNumber="360" masterPageNumber="345" pageNumber="358" updateTime="1722635640315" updateUser="ExternalLinkService">
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<mods:title id="A5F7F79A5AA496D3760081E0953AB4A1">A review of the shore-fly genus Ephydra Fallén, 1810 (Diptera: Ephydridae) of Russia</mods:title>
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<mods:namePart id="92018D3DC105A3A9E9178F593FA82BE0">Krivosheina, M. G.</mods:namePart>
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<mods:namePart id="F8EDDCDC7BEC4788EBD233C2FF25D927">Ozerov, A. L.</mods:namePart>
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<treatment id="0388A40719008C4D5BF1B03DFA8FC698" LSID="urn:lsid:plazi:treatment:0388A40719008C4D5BF1B03DFA8FC698" httpUri="http://treatment.plazi.org/id/0388A40719008C4D5BF1B03DFA8FC698" lastPageNumber="358" pageId="13" pageNumber="358">
<subSubSection id="C33B469A19008C4D5BF1B03DFD56C458" box="[226,715,1878,1902]" pageId="13" pageNumber="358" type="nomenclature">
<paragraph id="8B9E151119008C4D5BF1B03DFD56C458" blockId="13.[226,715,1878,1934]" box="[226,715,1878,1902]" pageId="13" pageNumber="358">
<heading id="D0D6A27D19008C4D5BF1B03DFD56C458" box="[226,715,1878,1902]" fontSize="10" level="2" pageId="13" pageNumber="358" reason="4">
<taxonomicName id="4C216E9219008C4D5BF1B03DFD17C458" ID-CoL="9J76B" authority="Krivosheina et Ozerov" authorityName="Krivosheina et Ozerov" authorityYear="2021" box="[226,650,1878,1902]" class="Insecta" family="Ephydridae" genus="Ephydra" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="358" phylum="Arthropoda" rank="species" species="villosa" status="sp. nov.">
<emphasis id="B955C90319008C4D5BF1B03DFE10C458" box="[226,397,1878,1902]" italics="true" pageId="13" pageNumber="358">Ephydra villosa</emphasis>
Krivosheina et Ozerov
</taxonomicName>
,
<emphasis id="B955C90319008C4D5985B03DFD56C458" bold="true" box="[662,715,1878,1902]" pageId="13" pageNumber="358">
<taxonomicNameLabel id="A266747819008C4D5985B03DFD56C458" box="[662,715,1878,1902]" pageId="13" pageNumber="358" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33B469A19008C4D5A6AB01DFDD6C4B8" box="[377,587,1910,1934]" pageId="13" pageNumber="358" type="description">
<paragraph id="8B9E151119008C4D5A6AB01DFDD6C4B8" blockId="13.[226,715,1878,1934]" box="[377,587,1910,1934]" pageId="13" pageNumber="358">
<figureCitation id="131A099419008C4D5A6AB01DFE56C4B8" box="[377,459,1910,1934]" captionStart="Figs 715" captionStartId="4.[226,263,1757,1776]" captionTargetBox="[224,728,1366,1684]" captionTargetId="figure-15@4.[207,757,1346,1698]" captionTargetPageId="4" captionText="Figs 715. Aedeagus of Ephydra spp., lateral view: 7 — E. afghanica Dahl; 8 — E. attica Becker; 9 — E. glauca Meigen; 10 — E. japonica Miyagi; 11 — E. macellaria Egger; 12 — E. pseudomurina Krivosheina; 13 — E. riparia Fallén; 14 — E. sholtzi Becker; 15 — E. villosa sp.n. Рис. 715. Эдеагус Ephydra spp., сбоку: 7 — E. afghanica Dahl; 8 — E. attica Becker; 9 — E. glauca Meigen; 10 — E. japonica Miyagi; 11 — E. macellaria Egger; 12 — E. pseudomurina Krivosheina; 13 — E. riparia Fallén; 14 — E. sholtzi Becker; 15 — E. villosa sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178908" httpUri="https://zenodo.org/record/13178908/files/figure.png" pageId="13" pageNumber="358">Figs 15</figureCitation>
,
<figureCitation id="131A099419008C4D5AC4B01DFE69C4B8" box="[471,500,1910,1934]" captionStart="Figs 1624" captionStartId="6.[224,261,1523,1542]" captionTargetBox="[196,545,724,1418]" captionTargetPageId="6" captionText="Figs 1624. Gonite of Ephydra spp., lateral view: 16 — E. afghanica Dahl; 17 — E. attica Becker; 18 — E. glauca Meigen; 19 — E. japonica Miyagi; 20 — E. macellaria Egger; 21 — E. pseudomurina Krivosheina; 22 — E. riparia Fallén; 23 — E. sholtzi Becker; 24 — E. villosa sp.n. Рис. 1624. Гонит Ephydra spp., сбоку: 16 — E. afghanica Dahl; 17 — E. attica Becker; 18 — E. glauca Meigen; 19 — E. japonica Miyagi; 20 — E. macellaria Egger; 21 — E. pseudomurina Krivosheina; 22 — E. riparia Fallén; 23 — E. sholtzi Becker; 24 — E. villosa sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178910" httpUri="https://zenodo.org/record/13178910/files/figure.png" pageId="13" pageNumber="358">24</figureCitation>
,
<figureCitation id="131A099419008C4D5913B01DFD80C4B8" box="[512,541,1910,1934]" captionStart="Figs 3133" captionStartId="9.[221,257,1173,1192]" captionTargetBox="[202,1370,331,1139]" captionTargetId="figure-8@9.[632,975,314,1085]" captionTargetPageId="9" captionText="Figs 3133. Epandrium and surstyli of Ephydra spp., dorsal view: 31 — E. riparia Fallén; 32 — E. sholtzi Becker; 33 — E. villosa sp.n. Рис. 3133. Эпандриум и сурстили Ephydra spp., сверху: 31 — E. riparia Fallén; 32 — E. sholtzi Becker; 33 — E. villosa sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178918" httpUri="https://zenodo.org/record/13178918/files/figure.png" pageId="13" pageNumber="358">33</figureCitation>
,
<figureCitation id="131A099419008C4D593AB01DFDDAC4B8" box="[553,583,1910,1934]" captionStart="Figs 4042" captionStartId="12.[226,262,1217,1236]" captionTargetBox="[1071,1378,340,1175]" captionTargetId="figure-16@12.[1058,1399,314,1195]" captionTargetPageId="12" captionText="Figs 4042. Epandrium and surstyli of Ephydra spp., lateral view: 40 — E. riparia Fallén; 41 — E. sholtzi Becker; 42 — E. villosa sp.n. Рис. 4042. Эпандриум и сурстили Ephydra spp., сбоку: 40 — E. riparia Fallén; 41 — E. sholtzi Becker; 42 — E. villosa sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178924" httpUri="https://zenodo.org/record/13178924/files/figure.png" pageId="13" pageNumber="358">42</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B469A19008C4D5BF1B0D9FED8CB38" pageId="13" pageNumber="358" type="materials_examined">
<paragraph id="8B9E151119008C4D5BF1B0D9FED8CB38" blockId="13.[189,775,1967,2062]" pageId="13" pageNumber="358">
<materialsCitation id="3B491F4C19008C4D5BF1B0D9FEDCCB38" collectingDate="1962-07-19" collectionCode="ZISP, ZMUM" country="Russia" latitude="53.017" location="Sakhyurta" longLatPrecision="64" longitude="106.883" municipality="Baikal Lake" pageId="13" pageNumber="358" specimenCount="2" specimenCount-male="2" stateProvince="Irkutsk" typeStatus="holotype">
MATERIAL.
<typeStatus id="549AABB319008C4D5A4FB0D9FE31C4F3" box="[348,428,1970,1989]" pageId="13" pageNumber="358" type="holotype">Holotype</typeStatus>
<specimenCount id="9D27DE9819008C4D5AA2B0C4FE58C4F3" box="[433,453,1967,1989]" count="1" pageId="13" pageNumber="358" type="male"></specimenCount>
,
<collectingCountry id="F336558119008C4D5ADCB0D9FD91C4F3" box="[463,524,1970,1989]" name="Russia" pageId="13" pageNumber="358">Russia</collectingCountry>
:
<collectingRegion id="49E5DBF319008C4D5907B0D9FD0DC4F3" box="[532,656,1970,1989]" country="Russia" name="Irkutsk" pageId="13" pageNumber="358">Irkutsk Oblast</collectingRegion>
,
<collectingMunicipality id="6BFA8F6B19008C4D598AB0D9FC9EC4F3" box="[665,771,1970,1989]" pageId="13" pageNumber="358">Baikal Lake</collectingMunicipality>
,
<location id="8EFE43CA19008C4D5BAEB0A1FE88C4EB" LSID="urn:lsid:plazi:treatment:0388A40719008C4D5BF1B03DFA8FC698:8EFE43CA19008C4D5BAEB0A1FE88C4EB" box="[189,277,1994,2013]" country="Russia" latitude="53.017" longLatPrecision="64" longitude="106.883" municipality="Baikal Lake" name="Sakhyurta" pageId="13" pageNumber="358" stateProvince="Irkutsk">Sakhyurta</location>
env., MRS [=river station Malomorskaya) (
<geoCoordinate id="EE1573D619008C4D59A0B0A1FC9BC4EB" box="[691,774,1994,2013]" degrees="53.017" direction="north" orientation="latitude" pageId="13" pageNumber="358" precision="55" value="53.017">53.017ºN</geoCoordinate>
<geoCoordinate id="EE1573D619008C4D5BAEB089FE81C4C3" box="[189,284,2018,2037]" degrees="106.883" direction="east" orientation="longitude" pageId="13" pageNumber="358" precision="55" value="106.883">106.883ºE</geoCoordinate>
)],
<date id="FF9F33D119008C4D5A23B089FE06C4C3" box="[304,411,2018,2037]" pageId="13" pageNumber="358" value="1962-07-19">
<collectingDate id="EFDBCA3919008C4D5A23B089FE06C4C3" box="[304,411,2018,2037]" pageId="13" pageNumber="358" value="1962-07-19">19.VII.1962</collectingDate>
</date>
, Gorodkov (
<collectionCode id="ED308DD419008C4D5914B089FDA5C4C3" box="[519,568,2018,2037]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="13" pageNumber="358">ZISP</collectionCode>
).
<typeStatus id="549AABB319008C4D5955B088FD0FC4C0" box="[582,658,2019,2038]" pageId="13" pageNumber="358" type="paratype">Paratype</typeStatus>
<specimenCount id="9D27DE9819008C4D5985B08BFD37C4C0" box="[662,682,2016,2038]" count="1" pageId="13" pageNumber="358" type="male"></specimenCount>
with same label (
<collectionCode id="ED308DD419008C4D5BE7B090FEA1CB38" box="[244,316,2043,2062]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="13" pageNumber="358">ZMUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B469A19008C4D5841B650FA8FC698" pageId="13" pageNumber="358" type="description">
<paragraph id="8B9E151119008C4D5841B650FC3FC246" blockId="13.[812,1399,315,1454]" pageId="13" pageNumber="358">
DESCRIPTION. Medium sized flies, body length
<quantity id="4CD9B8F419008C4D583FB633FC03C246" box="[812,926,344,368]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="13" pageNumber="358" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">55.5 mm</quantity>
.
</paragraph>
<paragraph id="8B9E151119008C4D5841B61DFC57C16D" blockId="13.[812,1399,315,1454]" pageId="13" pageNumber="358">
<emphasis id="B955C90319008C4D5841B61DFC11C2B8" box="[850,908,374,398]" italics="true" pageId="13" pageNumber="358">Head</emphasis>
golden-brown pollinose except metallic shining frontal vitta and upper face, with greenish tinge; fronto-orbital plate brownish pollinose. 3 long fronto-orbital setae, anterior interfrontal strong, about 1/2 of ocellar. Postocular setae strong. Palpus dark grey. Antenna blackish, brownish dusted; arista short pubescent on dorsal surface on basal 1/2. Gena and parafacial golden dusted.
</paragraph>
<paragraph id="8B9E151119008C4D5841B50AFBC0C185" blockId="13.[812,1399,315,1454]" pageId="13" pageNumber="358">
<emphasis id="B955C90319008C4D5841B50AFC02C14F" box="[850,927,609,633]" italics="true" pageId="13" pageNumber="358">Thorax</emphasis>
black, brownish pollinose. Scutum subshining with bluish-greenish tinger. Acrostichals setulose in two rows, intra-alars 2+2.
</paragraph>
<paragraph id="8B9E151119008C4D5841B5D2FC0AC03D" blockId="13.[812,1399,315,1454]" pageId="13" pageNumber="358">
<emphasis id="B955C90319008C4D5841B5D2FC19C1E6" box="[850,900,697,720]" italics="true" pageId="13" pageNumber="358">Legs</emphasis>
. Femora and tibia black, densely greyish dust- ed; tarsi of all legs yellowish, but tarsomeres 4 and 5 darkened.
</paragraph>
<paragraph id="8B9E151119008C4D5841B47AFB30C778" blockId="13.[812,1399,315,1454]" pageId="13" pageNumber="358">
<emphasis id="B955C90319008C4D5841B47AFC2AC01F" box="[850,951,785,809]" italics="true" pageId="13" pageNumber="358">Abdomen</emphasis>
black, grey pollinose, subshining. Tergite 5 approximately equal in length to tergite 4. Male genitalia small, as long as tergite 5. Epandrium oval in dorsal (posterior) view (
<figureCitation id="131A099419008C4D5F27B402FB18C0B7" box="[1076,1157,873,897]" captionStart="Figs 3133" captionStartId="9.[221,257,1173,1192]" captionTargetBox="[202,1370,331,1139]" captionTargetId="figure-8@9.[632,975,314,1085]" captionTargetPageId="9" captionText="Figs 3133. Epandrium and surstyli of Ephydra spp., dorsal view: 31 — E. riparia Fallén; 32 — E. sholtzi Becker; 33 — E. villosa sp.n. Рис. 3133. Эпандриум и сурстили Ephydra spp., сверху: 31 — E. riparia Fallén; 32 — E. sholtzi Becker; 33 — E. villosa sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178918" httpUri="https://zenodo.org/record/13178918/files/figure.png" pageId="13" pageNumber="358">Fig. 33</figureCitation>
), wider in cercal part, covered with long setae; surstyli relatively broad and curved, covered with many long setae. Epandrium curved in lateral view (
<figureCitation id="131A099419008C4D5F38B4AAFBE3C0EF" box="[1067,1150,961,985]" captionStart="Figs 4042" captionStartId="12.[226,262,1217,1236]" captionTargetBox="[1071,1378,340,1175]" captionTargetId="figure-16@12.[1058,1399,314,1195]" captionTargetPageId="12" captionText="Figs 4042. Epandrium and surstyli of Ephydra spp., lateral view: 40 — E. riparia Fallén; 41 — E. sholtzi Becker; 42 — E. villosa sp.n. Рис. 4042. Эпандриум и сурстили Ephydra spp., сбоку: 40 — E. riparia Fallén; 41 — E. sholtzi Becker; 42 — E. villosa sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178924" httpUri="https://zenodo.org/record/13178924/files/figure.png" pageId="13" pageNumber="358">Fig. 42</figureCitation>
); surstylus is at obtuse angle to epandrium, with short subapical tooth. Aedeagus long and narrow with pointed tip in lateral view (
<figureCitation id="131A099419008C4D5827B372FC14C706" box="[820,905,1048,1073]" captionStart="Figs 715" captionStartId="4.[226,263,1757,1776]" captionTargetBox="[224,728,1366,1684]" captionTargetId="figure-15@4.[207,757,1346,1698]" captionTargetPageId="4" captionText="Figs 715. Aedeagus of Ephydra spp., lateral view: 7 — E. afghanica Dahl; 8 — E. attica Becker; 9 — E. glauca Meigen; 10 — E. japonica Miyagi; 11 — E. macellaria Egger; 12 — E. pseudomurina Krivosheina; 13 — E. riparia Fallén; 14 — E. sholtzi Becker; 15 — E. villosa sp.n. Рис. 715. Эдеагус Ephydra spp., сбоку: 7 — E. afghanica Dahl; 8 — E. attica Becker; 9 — E. glauca Meigen; 10 — E. japonica Miyagi; 11 — E. macellaria Egger; 12 — E. pseudomurina Krivosheina; 13 — E. riparia Fallén; 14 — E. sholtzi Becker; 15 — E. villosa sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178908" httpUri="https://zenodo.org/record/13178908/files/figure.png" pageId="13" pageNumber="358">Fig. 15</figureCitation>
). Gonite with broad basal part and slender pointed tip in lateral view (
<figureCitation id="131A099419008C4D5F43B35DFB3DC778" box="[1104,1184,1078,1102]" captionStart="Figs 1624" captionStartId="6.[224,261,1523,1542]" captionTargetBox="[196,545,724,1418]" captionTargetPageId="6" captionText="Figs 1624. Gonite of Ephydra spp., lateral view: 16 — E. afghanica Dahl; 17 — E. attica Becker; 18 — E. glauca Meigen; 19 — E. japonica Miyagi; 20 — E. macellaria Egger; 21 — E. pseudomurina Krivosheina; 22 — E. riparia Fallén; 23 — E. sholtzi Becker; 24 — E. villosa sp.n. Рис. 1624. Гонит Ephydra spp., сбоку: 16 — E. afghanica Dahl; 17 — E. attica Becker; 18 — E. glauca Meigen; 19 — E. japonica Miyagi; 20 — E. macellaria Egger; 21 — E. pseudomurina Krivosheina; 22 — E. riparia Fallén; 23 — E. sholtzi Becker; 24 — E. villosa sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178910" httpUri="https://zenodo.org/record/13178910/files/figure.png" pageId="13" pageNumber="358">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="8B9E151119008C4D5841B338FADFC6A6" blockId="13.[812,1399,315,1454]" pageId="13" pageNumber="358">
NOTE. The species is close to
<taxonomicName id="4C216E9219008C4D5FB6B33FFCF9C7BE" authority="Wirth, 1975" authorityName="Wirth" authorityYear="1975" class="Insecta" family="Ephydridae" genus="Ephydra" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="358" phylum="Arthropoda" rank="species" species="tibetensis">
<emphasis id="B955C90319008C4D5FB6B33FFAB4C75D" box="[1189,1321,1107,1131]" italics="true" pageId="13" pageNumber="358">E. tibetensis</emphasis>
<bibRefCitation id="EFB068E019008C4D5E22B338FCF9C7BE" author="Wirth W. W." pageId="13" pageNumber="358" pagination="11 - 44" refId="ref13912" refString="Wirth W. W. 1975. A revision of the brine flies of the genus Ephydra of the Old World (Diptera: Ephydridae) // Entomologica Scandinavica. Vol. 6. No. 1. P. 11 - 44." type="book chapter" year="1975">Wirth, 1975</bibRefCitation>
</taxonomicName>
, differing from it in extremely developed setae and hairs on body and the structure of male genitalia; surstylus with well separated subapical tooth in lateral view, aedeagus with long apical narrow part, gonite significantly narrowed apically and curved at the very tip. From
<taxonomicName id="4C216E9219008C4D5889B26FFB96C62D" authorityName="Becker" authorityYear="1896" box="[922,1035,1283,1307]" class="Insecta" family="Ephydridae" genus="Ephydra" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="358" phylum="Arthropoda" rank="species" species="scholtzi">
<emphasis id="B955C90319008C4D5889B26FFB96C62D" box="[922,1035,1283,1307]" italics="true" pageId="13" pageNumber="358">E. scholtzi</emphasis>
</taxonomicName>
the new species differs in strong postocular setae and from
<taxonomicName id="4C216E9219008C4D5F40B24AFB20C60F" authorityName="Meigen" authorityYear="1830" box="[1107,1213,1313,1337]" class="Insecta" family="Ephydridae" genus="Ephydra" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="358" phylum="Arthropoda" rank="species" species="glauca">
<emphasis id="B955C90319008C4D5F40B24AFB20C60F" box="[1107,1213,1313,1337]" italics="true" pageId="13" pageNumber="358">E. glauca</emphasis>
</taxonomicName>
in weaker pollinosity and the presence of subshining surfaces on upper face, scutum and abdomen. From both species it differs in the development of 2+2 intra-alar setae.
</paragraph>
<paragraph id="8B9E151119008C4D5841B2FDFA8FC698" blockId="13.[812,1399,315,1454]" box="[850,1298,1430,1454]" pageId="13" pageNumber="358">
DISRTIBUTION.
<collectingCountry id="F336558119008C4D5F08B2FDFBFBC698" box="[1051,1126,1430,1454]" name="Russia" pageId="13" pageNumber="358">Russia</collectingCountry>
:
<collectingRegion id="49E5DBF319008C4D5F62B2FDFA90C698" box="[1137,1293,1430,1454]" country="Russia" name="Irkutsk" pageId="13" pageNumber="358">Irkutsk Oblast</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>