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<document id="EA69821A5C008133865AA4FC708DA83E" ID-DOI="10.24199/j.mmv.2024.83.01" ID-ISSN="1447-2554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718999175370" checkinUser="felipe" docAuthor="Mah, Christopher L." docDate="2023" docId="03889522DD70FFA0FF4AFDCBFD5E889A" docLanguage="en" docName="MemMusVictoria.83.1-35.pdf" docOrigin="Memoirs of Museum Victoria 83" docSource="http://dx.doi.org/10.24199/j.mmv.2024.83.01" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Atheraster umbo Mah, 2023, n. sp." docType="treatment" docVersion="1" lastPageNumber="13" masterDocId="FFB1ED5ADD79FFACFFD9FFE2FF8F8D61" masterDocTitle="New Goniasteridae and in situ observations significant to deep-sea coral predation" masterLastPageNumber="35" masterPageNumber="1" pageNumber="10" updateTime="1719015880144" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3734F533BE264D8C863821E2018E9507">New Goniasteridae and in situ observations significant to deep-sea coral predation</mods:title>
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<treatment id="03889522DD70FFA0FF4AFDCBFD5E889A" LSID="urn:lsid:plazi:treatment:03889522DD70FFA0FF4AFDCBFD5E889A" httpUri="http://treatment.plazi.org/id/03889522DD70FFA0FF4AFDCBFD5E889A" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<subSubSection id="C33B77BFDD70FFA5FF4AFDCBFC858F06" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9E2434DD70FFA5FF4AFDCBFE0C8F5F" blockId="9.[147,387,553,575]" box="[147,387,553,575]" pageId="9" pageNumber="10">
<heading id="D0D69358DD70FFA5FF4AFDCBFE0C8F5F" bold="true" box="[147,387,553,575]" fontSize="9" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B955F826DD70FFA5FF4AFDCBFE0C8F5F" bold="true" box="[147,387,553,575]" pageId="9" pageNumber="10">
<taxonomicName id="4C215FB7DD70FFA5FF4AFDCBFECA8F5E" ID-CoL="C6D56" authorityName="Mah" authorityYear="2023" box="[147,325,553,575]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="umbo" status="sp. nov.">
<emphasis id="B955F826DD70FFA5FF4AFDCBFECA8F5E" bold="true" box="[147,325,553,575]" italics="true" pageId="9" pageNumber="10">Atheraster umbo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A266455DDD70FFA5FE92FDCBFE0C8F5F" box="[331,387,553,574]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9E2434DD70FFA5FF60FDB6FC858F06" blockId="9.[147,778,596,642]" box="[185,778,596,615]" pageId="9" pageNumber="10">urn:lsid:zoobank.org:act:049A6DE1-0722-4252-A363-</paragraph>
</subSubSection>
<subSubSection id="C33B77BFDD70FFA0FF4AFD8DFD5E889A" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9E2434DD70FFA5FF4AFD8DFEA88FE3" blockId="9.[147,778,596,642]" box="[147,295,623,642]" pageId="9" pageNumber="10">D373EE1C1E25</paragraph>
<paragraph id="8B9E2434DD70FFA5FF4AFD78FEC48FCE" blockId="9.[147,331,665,688]" box="[147,331,665,688]" pageId="9" pageNumber="10">
<figureCitation id="131A38B1DD70FFA5FF4AFD78FE9B8FD1" box="[147,276,666,688]" captionStart="Figure 4" captionStartId="10.[140,200,1830,1849]" captionTargetBox="[212,1368,227,1813]" captionTargetId="figure-16@10.[212,1368,227,1813]" captionTargetPageId="10" captionText="Figure 4. Atheraster umbo n. sp. holotype, NMV F307974: a, abactinal surface; b, abactinal surface/marginal plates detail; c, abactinal surface detail; d, lateral view showing marginal plate surfaces e, actinal surface; f, actinal surface detail including pedicellariae, furrow spines. Pedicellariae circled. Scale: a, e=10.0 mm; b, d, f=5.0 mm; c=3.0 mm." pageId="9" pageNumber="10">Figures 4af</figureCitation>
,
<figureCitation id="131A38B1DD70FFA5FEC7FD7BFEC48FCE" box="[286,331,665,687]" captionStart="Figure 5" captionStartId="11.[147,207,1830,1849]" captionTargetBox="[147,1447,422,1814]" captionTargetId="figure-70@11.[147,1447,422,1814]" captionTargetPageId="11" captionText="Figure 5. Atheraster umbo n. sp. variation paratype, NMV F84959: a, abactinal surface; b, abactinal surface detail; c, abactinal arm surface detail; d, lateral view showing marginal plate surfaces e, actinal surface; f, actinal surface detail including pedicellariae, furrow spines. Scale: a, e=10.0 mm; b, c, d, f=5.0 mm." pageId="9" pageNumber="10">5af</figureCitation>
</paragraph>
<paragraph id="8B9E2434DD70FFA5FF4AFD24FEB88E77" blockId="9.[147,778,710,790]" pageId="9" pageNumber="10">
<emphasis id="B955F826DD70FFA5FF4AFD24FE8C8FBA" box="[147,259,710,731]" italics="true" pageId="9" pageNumber="10">Etymology.</emphasis>
The species epithet,
<taxonomicName id="4C215FB7DD70FFA5FE3AFD24FD948FBA" authorityName="Mah" authorityYear="2023" box="[483,539,710,731]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="umbo">
<emphasis id="B955F826DD70FFA5FE3AFD24FD948FBA" box="[483,539,710,731]" italics="true" pageId="9" pageNumber="10">umbo</emphasis>
</taxonomicName>
, is Latin for a boss or swelling, alluding to the tumescent projections present on the marginal plates.
</paragraph>
<paragraph id="8B9E2434DD70FFA5FF4AFCCFFD6C88ED" blockId="9.[146,781,813,1420]" pageId="9" pageNumber="10">
<emphasis id="B955F826DD70FFA5FF4AFCCFFF708E23" box="[147,255,813,834]" italics="true" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Body stout, strongly stellate (R/r=3.2), arms elongate, triangular, interradial arcs weakly curved to straight. Abactinal plates irregular, bare, extending to arm tip. Arm plates variably flat to strongly convex, acutely large relative to adjacent disk plates. Pedicellariae abundant, 412, along depression in each interradius. Five plates across proximally on arms decreasing to a single series to terminus. Marginal plates, 4044 per interradius, 2022 per arm side, interradially with 210 small granules/tubercles, but otherwise smooth and bare. Marginal plates along the arms with single distinct tumescence. Granules coarse, nearly tubercular, most abundant (115) interradially on marginal plates, decreasing then absent along arm surface. Actinal plates with distinct alveolar pedicellariae flanked on either side by bisected hemispherical granules. Furrow spines 515, but mostly seven or eight, with a prominent single tong-like pedicellaria with quadrate valves on each adambulacral plate. Spine, large thick, 23 times the thickness of the furrow spines, identical in length to the longest of the furrow spines, present at oblique angle to pedicellariae. Oral plates, each with large, paddle-shaped pedicellaria per plate, thus two per interradius, but not symmetrically positioned.
</paragraph>
<paragraph id="8B9E2434DD70FFA5FF4AFA41FD8E8B98" blockId="9.[147,779,1442,1903]" pageId="9" pageNumber="10">
<emphasis id="B955F826DD70FFA5FF4AFA41FE8B88D9" box="[147,260,1443,1464]" italics="true" pageId="9" pageNumber="10">Comments.</emphasis>
Although lacking well-developed spines, the tumescent structures on the marginal plates display a strongly arched shape and pattern that appears very similar to spines in other
<taxonomicName id="4C215FB7DD70FFA5FF04FA18FE228B71" box="[221,429,1530,1552]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="B955F826DD70FFA5FF04FA18FEC78B6E" box="[221,328,1530,1551]" italics="true" pageId="9" pageNumber="10">Atheraster</emphasis>
species.
</taxonomicName>
This species marginal plate ornamentation pattern parallels
<taxonomicName id="4C215FB7DD70FFA5FE3EF9FAFDCB8B4D" authorityName="Mah" authorityYear="2023" box="[487,580,1559,1581]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="pheos" status="n. sp.">
<emphasis id="B955F826DD70FFA5FE3EF9FAFDCB8B4D" box="[487,580,1559,1581]" italics="true" pageId="9" pageNumber="10">A. pheos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A266455DDD70FFA5FD89F9FAFD058B4C" box="[592,650,1560,1581]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
in showing interradial superomarginal plates with 210 minute surficial accessory structures (i.e. granules/tubercles) relative to the larger, single arched tumescence present on superomarginal arm plates present on each arm.
<taxonomicName id="4C215FB7DD70FFA5FE01F96EFD0B8BC0" authorityName="Mah" authorityYear="2023" box="[472,644,1676,1697]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="pheos">
<emphasis id="B955F826DD70FFA5FE01F96EFD0B8BC0" box="[472,644,1676,1697]" italics="true" pageId="9" pageNumber="10">Atheraster pheos</emphasis>
</taxonomicName>
also displays many small spines, 610 on each interradial superomarginal, which are succeeded by a single conical spine on each superomarginal plate along the arm.
</paragraph>
<paragraph id="8B9E2434DD70FFA5FF60F8E3FBF18C54" blockId="9.[147,779,1442,1903]" lastBlockId="9.[816,1449,229,599]" pageId="9" pageNumber="10">
The
<specimenCount id="9D27EFBDDD70FFA5FF37F8E3FED38A76" box="[238,348,1793,1815]" count="3" pageId="9" pageNumber="10" type="generic" typeStatus="type">three type</specimenCount>
specimens, which were collected from distinctly different localities (the Cocos/Keeling Islands and Tasmanian/southeast Pacific), show the same arched or tumescent marginal plates. Additional specimens could show these to be damaged or be some sort of incipient spine, but examination of material on hand suggests they are a diagnostic character of this distinct species.
</paragraph>
<paragraph id="8B9E2434DD70FFA5FC8EFEDEFB668C82" blockId="9.[816,1449,229,599]" pageId="9" pageNumber="10">
This species invites comparison with two Pacific
<taxonomicName id="4C215FB7DD70FFA5FCE8FEBBFC7F8C0E" box="[817,1008,345,367]" pageId="9" pageNumber="10">
<emphasis id="B955F826DD70FFA5FCE8FEBBFC128C0F" box="[817,925,345,366]" italics="true" pageId="9" pageNumber="10">Circeaster</emphasis>
species
</taxonomicName>
,
<taxonomicName id="4C215FB7DD70FFA5FC24FEBBFB6E8C0E" authority="Mah 2006" authorityName="Mah" authorityYear="2006" box="[1021,1249,345,367]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="sandrae">
<emphasis id="B955F826DD70FFA5FC24FEBBFBE18C0F" box="[1021,1134,345,366]" italics="true" pageId="9" pageNumber="10">C. sandrae</emphasis>
<bibRefCitation id="EFB059C5DD70FFA5FBAFFEBBFB6E8C0E" author="Mah, C. L." box="[1142,1249,345,367]" pageId="9" pageNumber="10" pagination="917 - 954" refId="ref23487" refString="Mah, C. L. 2006. Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema 28 (4): 917 - 954." type="journal article" year="2006">Mah 2006</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C215FB7DD70FFA5FACDFEBBFCE68CED" authority="Mah 2006" authorityName="Mah" authorityYear="2006" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="pullus">
<emphasis id="B955F826DD70FFA5FACDFEBBFAFE8C0F" box="[1300,1393,345,366]" italics="true" pageId="9" pageNumber="10">C. pullus</emphasis>
<bibRefCitation id="EFB059C5DD70FFA5FAA0FEBBFCE68CED" author="Mah, C. L." pageId="9" pageNumber="10" pagination="917 - 954" refId="ref23487" refString="Mah, C. L. 2006. Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema 28 (4): 917 - 954." type="journal article" year="2006">Mah 2006</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C215FB7DD70FFA5FCA9FE94FB958CEA" authorityName="Mah" authorityYear="2023" box="[880,1050,374,395]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="umbo" status="sp. nov.">
<emphasis id="B955F826DD70FFA5FCA9FE94FB958CEA" box="[880,1050,374,395]" italics="true" pageId="9" pageNumber="10">Atheraster umbo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A266455DDD70FFA5FBC6FE94FBDA8CEA" box="[1055,1109,374,395]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
differs in lacking any granulation on the marginal plates surface, displaying large strongly convex abactinal arm plate and in showing a large number of abactinal granules around the madreporite.
</paragraph>
<paragraph id="8B9E2434DD70FFA5FC8EFE08FAF28F36" blockId="9.[816,1449,229,599]" pageId="9" pageNumber="10">
Although no direct association was observed, the
<collectingCountry id="F33664A4DD70FFA5FAA0FE08FC168F7C" name="New Caledonia" pageId="9" pageNumber="10">New Caledonia</collectingCountry>
specimen, IE-2013-7118, was collected from a station that also included members of the
<taxonomicName id="4C215FB7DD70FFA5FB2BFDC6FACC8F5B" box="[1266,1347,548,570]" class="Anthozoa" family="Isididae" kingdom="Animalia" order="Alcyonacea" pageId="9" pageNumber="10" phylum="Cnidaria" rank="family">Isididae</taxonomicName>
(bamboo corals), which goniasterid predators feed upon frequently.
</paragraph>
<paragraph id="8B9E2434DD70FFA5FCE8FD8FFBA28FDD" blockId="9.[817,1448,621,701]" pageId="9" pageNumber="10">
<emphasis id="B955F826DD70FFA5FCE8FD8FFC228FE3" box="[817,941,621,642]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C215FB7DD70FFA5FCE8FD8FFC278FE3" authorityName="Mah" authorityYear="2023" box="[817,936,621,642]" pageId="9" pageNumber="33" rank="subFamily" status="nov. subfamily" subFamily="Occurrence">Occurrence</taxonomicName>
.
</emphasis>
<collectingCountry id="F33664A4DD70FFA5FC6AFD8FFB268FE2" box="[947,1193,621,643]" name="Cocos (Keeling) Islands" pageId="9" pageNumber="10">Cocos (Keeling) Islands</collectingCountry>
Territory, Indian Ocean,
<collectingCountry id="F33664A4DD70FFA5FCE8FD68FC1F8FC1" box="[817,912,650,672]" name="Australia" pageId="9" pageNumber="10">Australia</collectingCountry>
, Lord Howe Rise, Southeast Cape,
<collectingRegion id="49E5EAD6DD70FFA5FAD1FD68FAE48FC1" box="[1288,1387,650,672]" country="Australia" name="Tasmania" pageId="9" pageNumber="10">Tasmania</collectingRegion>
,
<collectingCountry id="F33664A4DD70FFA5FAA0FD68FC178FDC" name="New Caledonia" pageId="9" pageNumber="10">New Caledonia</collectingCountry>
1896
<quantity id="4CD989D1DD70FFA5FC05FD45FBA78FDD" box="[988,1064,679,701]" metricMagnitude="3" metricUnit="m" metricValue="1.589" pageId="9" pageNumber="10" unit="m" value="1589.0">1589 m</quantity>
.
</paragraph>
<paragraph id="8B9E2434DD70FFA5FCE8FD31FA288E67" blockId="9.[816,1449,723,1903]" pageId="9" pageNumber="10">
<emphasis id="B955F826DD70FFA5FCE8FD31FC228F89" box="[817,941,723,744]" italics="true" pageId="9" pageNumber="10">Description.</emphasis>
Body stout, strongly stellate (R/r=2.13.2), arms elongate, triangular, interradial arcs weakly curved to straight.
</paragraph>
<paragraph id="8B9E2434DD70FFA5FC8EFCEFFBA58B51" blockId="9.[816,1449,723,1903]" pageId="9" pageNumber="10">
Abactinal surface, weakly arched, plates abutted, polygonal, extending from disk to arm tip. Arm and disk plates round to irregularly polygonal in shape but relatively similar in size. The
<typeStatus id="549A9A96DD70FFA5FBDDFC86FBD08E1B" box="[1028,1119,868,890]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
, from Ningaloo, has arm plates only slightly larger than those on disk with gradual transition, but a minority of arm plates, variably 14 per arm, are strongly convex, rising well above the body surface. Tasmanian specimens show arm plates up to three or four times the size of disk plates, most arm plates with strongly convex surface, appearing nearly hemispherical. Arm plate surfaces all smooth, devoid of granules with gradually decreasing number of abactinal plates, 35 at arm base, decreasing to a single row adjacent to the terminus. Superomarginals with two or three pairs abutted at arm tip on the Tasmanian specimens, but plates continue to terminus on the Ningaloo specimen. Disk plates mostly bare save for dense aggregation of granules surrounding the madreporite. Pedicellariae present in abundance on the Ningaloo specimen, but largely absent from the surface of the Tasmanian ones. Paddle-shaped pedicellariae with flat, round valves present on a minority of disk plates, each pedicellaria within a distinct alveolus or pit. Pedicellariae are most abundant, 412, in a distinct valley-like depression present in each interradius. Each plate surrounded by 930 polygonal to quadrate granules in close-set series. Each abactinal plate central region flat to weakly concave. Madreporite convex, round in shape with well-developed sulci. Surrounding region with large, coarse granules, 112 per plate.
</paragraph>
<paragraph id="8B9E2434DD70FFA0FC8EF9D5FDB78C79" blockId="9.[816,1449,723,1903]" lastBlockId="12.[139,772,229,1149]" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">Marginal plates, 4044 (2022 along each arm), present in each interradius (arm tip to arm tip, forming a wide boundary around body periphery. Interradial marginal plates with a laterally-directed facing becoming more dorsal facing along arms. Disk superomarginal plates bare, smooth save for 210, distinct, spherical granules or tubercles, leaving concave pitting on surface, centrally present on each interradial plate. Ornamentation on superomarginal plate surface along arm with a single distinct tumescence, variably arched with a tip becoming more rounded distally, forming a distinctly crenulated periphery along the arm. Pedicellariae alveolar, 14, tong-like with triangular blades on marginal plate surface. Superomarginal plate surface weakly concave, no granules on superomarginal plate surface. Each superomarginal plate with a rounded edge, each plate round in cross-section. Marginal plates with 40100 peripheral granules, quadrate to polygonal. Inferomarginals with 120 distinct, coarse granules, highest interradially, with 1015 on central inferomarginal plate surfaces decreasing to a single granule at arm base. Pedicellariae, 16, mostly 34, alveolar with triangular valves decreasing distally and absent along arms.</paragraph>
<caption id="DF5E74BCDD73FFA6FF55F8C4FC8C8A0F" pageId="10" pageNumber="11" startId="10.[140,200,1830,1849]" targetBox="[212,1368,227,1813]" targetPageId="10" targetType="figure">
<paragraph id="8B9E2434DD73FFA6FF55F8C4FC8C8A0F" blockId="10.[140,1441,1830,1902]" pageId="10" pageNumber="11">
Figure 4.
<taxonomicName id="4C215FB7DD73FFA6FF3DF8C4FEF68A58" authorityName="Mah" authorityYear="2023" box="[228,377,1830,1849]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="umbo" status="sp. nov.">
<emphasis id="B955F826DD73FFA6FF3DF8C4FEF68A58" box="[228,377,1830,1849]" italics="true" pageId="10" pageNumber="11">Atheraster umbo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A266455DDD73FFA6FEA6F8C5FE3F8A5B" box="[383,432,1831,1850]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
holotype, NMV F307974: a, abactinal surface; b, abactinal surface/marginal plates detail; c, abactinal surface detail; d, lateral view showing marginal plate surfaces e, actinal surface; f, actinal surface detail including pedicellariae, furrow spines. Pedicellariae circled. Scale: a, e=10.0 mm; b, d, f=5.0 mm; c=3.0 mm.
</paragraph>
</caption>
<caption id="DF5E74BCDD72FFA7FF4AF8C4FEEE8A0F" pageId="11" pageNumber="12" startId="11.[147,207,1830,1849]" targetBox="[147,1447,422,1814]" targetPageId="11" targetType="figure">
<paragraph id="8B9E2434DD72FFA7FF4AF8C4FEEE8A0F" blockId="11.[147,1448,1830,1902]" pageId="11" pageNumber="12">
Figure 5.
<taxonomicName id="4C215FB7DD72FFA7FF31F8C4FEF48A58" authorityName="Mah" authorityYear="2023" box="[232,379,1830,1849]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="12" phylum="Echinodermata" rank="species" species="umbo" status="sp. nov.">
<emphasis id="B955F826DD72FFA7FF31F8C4FEF48A58" box="[232,379,1830,1849]" italics="true" pageId="11" pageNumber="12">Atheraster umbo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A266455DDD72FFA7FE59F8C5FE208A5B" box="[384,431,1831,1850]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
variation paratype, NMV F84959: a, abactinal surface; b, abactinal surface detail; c, abactinal arm surface detail; d, lateral view showing marginal plate surfaces e, actinal surface; f, actinal surface detail including pedicellariae, furrow spines. Scale: a, e=10.0 mm; b, c, d, f=5.0 mm.
</paragraph>
</caption>
<paragraph id="8B9E2434DD75FFA0FF6BFEFDFD068F5B" blockId="12.[139,772,229,1149]" pageId="12" pageNumber="13">Actinal plates in three full series arranged in chevrons, individual plates, irregularly polygonal to round. Each plate with distinctly large, alveolar, paddle-shaped pedicellariae, approximately 1.0 mm in length with quadrate tips, flanked on either side of paired valves by bisected hemispherical granules. Single actinal plate series continues to proximal arm region, where it ends. Actinal plates covered by large round tubercles, coarse granules abundant, closely arranged adjacent to inferomarginal plates. Each actinal plate with coarse granules, 6 to 15 present around periphery.</paragraph>
<paragraph id="8B9E2434DD75FFA0FF6BFDA3FDFC8E19" blockId="12.[139,772,229,1149]" pageId="12" pageNumber="13">Furrow spines 58, quadrate in cross-section with rounded tips. Furrow spines 1215 on adambulacral plate adjacent to the oral plate, these furrow spines flattened to spatulate with quadrate-shaped tips. Most adambulacral plates have a prominent, alveolar, paddle-shaped pedicellaria. Remaining surface of each adambulacral surface with a single, large blunt spine on distal side of plate, this spine 23 times the width of the furrow spines, extending to approximate height of tallest furrow spine. One to five thick subambulacral spines, short to granular in size on remainder of plate, but especially on side in contact with actinal plates.</paragraph>
<paragraph id="8B9E2434DD75FFA0FF6BFC9DFD01891C" blockId="12.[139,772,229,1149]" pageId="12" pageNumber="13">Oral plates, each with a large, paddle-shaped pedicellaria approximately 2.0 mm in length, thus two per interradius but unevenly present per interradius. Furrow spines on oral plates, 1825, closely arranged, each flattened with tallest plates projecting into mouth. Each oral plate with 810 granular spines with angular tips, quadrate to polygonal in cross-section, along the central fossae on the side of each plate. Peripheral oral plate with 57 angular granular spines along edge adjacent to contact with adambulacral plates.</paragraph>
<paragraph id="8B9E2434DD75FFA0FF55FB70FEA4886F" blockId="12.[140,772,1170,1531]" pageId="12" pageNumber="13">
<emphasis id="B955F826DD75FFA0FF55FB70FECA89C4" box="[140,325,1170,1189]" italics="true" pageId="12" pageNumber="13">Material examined.</emphasis>
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<emphasis id="B955F826DD75FFA0FE8BFB70FE3F89C4" bold="true" box="[338,432,1170,1189]" pageId="12" pageNumber="13">
<typeStatus id="549A9A96DD75FFA0FE8BFB70FE2389C4" box="[338,428,1170,1189]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
.
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<collectionCode id="ED30BCF1DD75FFA0FE66FB70FE7A89C4" box="[447,501,1170,1189]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="12" pageNumber="13" type="Museum">NMV</collectionCode>
<specimenCode id="DB878C4FDD75FFA0FDDDFB70FDDB89C4" box="[516,596,1170,1189]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="12" pageNumber="13" type="Museum">F307974</specimenCode>
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,
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,
<collectingCountry id="F33664A4DD75FFA0FE73FB4EFD8E89DE" box="[426,513,1196,1215]" name="Australia" pageId="12" pageNumber="13">Australia</collectingCountry>
.
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,
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, 1896
<quantity id="4CD989D1DD75FFA0FF1DFB25FE8389BB" box="[196,268,1223,1242]" metricMagnitude="3" metricUnit="m" metricValue="1.589" pageId="12" pageNumber="13" unit="m" value="1589.0">
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.
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et al. aboard
<collectionCode id="ED30BCF1DD75FFA0FD91FB25FDE989BB" box="[584,614,1223,1242]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="13">RV</collectionCode>
<taxonomicName id="4C215FB7DD75FFA0FDB6FB25FD5489B8" box="[623,731,1223,1241]" class="Actinopterygii" family="Nemichthyidae" genus="Investigator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B955F826DD75FFA0FDB6FB25FD5489B8" box="[623,731,1223,1241]" italics="true" pageId="12" pageNumber="13">Investigator</emphasis>
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, IN
<date id="FF9F02F4DD75FFA0FF55FB03FF698995" box="[140,230,1249,1268]" pageId="12" pageNumber="13" value="2022-05-08">
<collectingDate id="EFDBFB1CDD75FFA0FF55FB03FF698995" box="[140,230,1249,1268]" pageId="12" pageNumber="13" value="2022-05-08">2022 V08</collectingDate>
</date>
IOT 2,
<collectorName id="26D441E2DD75FFA0FEF6FB03FDA68995" box="[303,553,1249,1268]" pageId="12" pageNumber="13">Marine Invertebrates Team</collectorName>
,
<date id="FF9F02F4DD75FFA0FDEAFB03FD478995" box="[563,712,1249,1268]" pageId="12" pageNumber="13" value="2022-10-14" valueMax="2022-10-15" valueMin="2022-10-14">
<collectingDate id="EFDBFB1CDD75FFA0FDEAFB03FD478995" box="[563,712,1249,1268]" pageId="12" pageNumber="13" value="2022-10-14" valueMax="2022-10-15" valueMin="2022-10-14">1415 Oct 2022</collectingDate>
</date>
. 1 wet spec,
<collectionCode id="ED30BCF1DD75FFA0FF66FB1EFF42886E" box="[191,205,1276,1295]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
=7.7, r=2.4
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</paragraph>
<paragraph id="8B9E2434DD75FFA0FF6BFAF4FE8A8816" blockId="12.[140,772,1170,1531]" pageId="12" pageNumber="13">
<materialsCitation id="3B492E69DD75FFA0FF6BFAF4FD708822" collectionCode="NMV" elevation="1375" latitude="-44.27" location="84959. 84 km SSE of South East" longLatPrecision="682" longitude="147.33" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Cape, Tasmania" typeStatus="paratype">
<emphasis id="B955F826DD75FFA0FF6BFAF4FE968848" bold="true" box="[178,281,1302,1321]" pageId="12" pageNumber="13">
<typeStatus id="549A9A96DD75FFA0FF6BFAF4FE9A8848" box="[178,277,1302,1321]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED30BCF1DD75FFA0FEFCFAF4FED48848" box="[293,347,1302,1321]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="12" pageNumber="13" type="Museum">NMV</collectionCode>
F
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<quantity id="4CD989D1DD75FFA0FEADFAF7FE738849" box="[372,508,1301,1321]" metricMagnitude="7" metricUnit="m" metricValue="8.495984" pageId="12" pageNumber="13" unit="km" value="84959.84">84959. 84 km</quantity>
SSE of South East
</location>
<collectingRegion id="49E5EAD6DD75FFA0FD16FAF4FF6C8822" isEnumeration="true" pageId="12" pageNumber="13">Cape, Tasmania</collectingRegion>
on J1 seamount.
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,
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,
<quantity id="4CD989D1DD75FFA0FD96FAD2FD408822" box="[591,719,1328,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="12" pageNumber="13" unit="m" value="1375.0" valueMax="1450.0" valueMin="1300.0">
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</quantity>
, coll
</materialsCitation>
.
<materialsCitation id="3B492E69DD75FFA0FF55FAA8FF72883C" box="[140,253,1354,1373]" collectionCode="T" collectorName="Stranks" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Cape, Tasmania" typeStatus="paratype">
<collectionCode id="ED30BCF1DD75FFA0FF55FAA8FF14883C" box="[140,155,1354,1373]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="13">T</collectionCode>
.N.
<collectorName id="26D441E2DD75FFA0FF61FAA8FF72883C" box="[184,253,1354,1373]" pageId="12" pageNumber="13">Stranks</collectorName>
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<materialsCitation id="3B492E69DD75FFA0FEDBFAA8FD70883C" box="[258,767,1354,1373]" collectingDate="1997-01-27" collectionCode="CSIRO, R, V" location="Southern Surveyor." pageId="12" pageNumber="13" specimenCount="1" stateProvince="Cape, Tasmania" typeStatus="paratype">
<collectionCode id="ED30BCF1DD75FFA0FEDBFAA8FECA883C" box="[258,325,1354,1373]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="12" pageNumber="13" type="Museum">CSIRO</collectionCode>
,
<collectionCode id="ED30BCF1DD75FFA0FE94FAA8FED3883C" box="[333,348,1354,1373]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
/
<collectionCode id="ED30BCF1DD75FFA0FEBDFAA8FEFB883C" box="[356,372,1354,1373]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="13" type="Museum">V</collectionCode>
<emphasis id="B955F826DD75FFA0FEA0FAA8FDA6883C" box="[377,553,1354,1373]" italics="true" pageId="12" pageNumber="13">
<location id="8EFE72EFDD75FFA0FEA0FAA8FDA6883C" LSID="urn:lsid:plazi:treatment:03889522DD70FFA0FF4AFDCBFD5E889A:8EFE72EFDD75FFA0FEA0FAA8FDA6883C" box="[377,553,1354,1373]" name="Southern Surveyor." pageId="12" pageNumber="13" stateProvince="Cape, Tasmania">Southern Surveyor.</location>
</emphasis>
<date id="FF9F02F4DD75FFA0FDF4FAA8FD17883C" box="[557,664,1354,1373]" pageId="12" pageNumber="13" value="1997-01-27">
<collectingDate id="EFDBFB1CDD75FFA0FDF4FAA8FD17883C" box="[557,664,1354,1373]" pageId="12" pageNumber="13" value="1997-01-27">27 Jan 1997</collectingDate>
</date>
. 1 wet spec
</materialsCitation>
, R=6.2, r=2.0.
</paragraph>
<paragraph id="8B9E2434DD75FFA0FF6BFA9DFE2F88A7" blockId="12.[140,772,1170,1531]" pageId="12" pageNumber="13">
<materialsCitation id="3B492E69DD75FFA0FF6BFA9DFEB488F3" box="[178,315,1407,1426]" collectionCode="NMV" pageId="12" pageNumber="13" specimenCode="F159690" specimenCount="1">
<collectionCode id="ED30BCF1DD75FFA0FF6BFA9DFF6788F3" box="[178,232,1407,1426]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="12" pageNumber="13" type="Museum">NMV</collectionCode>
<specimenCode id="DB878C4FDD75FFA0FF35FA9DFEB488F3" box="[236,315,1407,1426]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="12" pageNumber="13" type="Museum">F159690</specimenCode>
</materialsCitation>
<materialsCitation id="3B492E69DD75FFA0FE99FA9DFDAD88CD" collectorName="Coll. Geoscience" country="Australia" elevation="1532" latitude="-27.413683" location="Lord Howe Rise" longLatPrecision="1" longitude="161.70035" pageId="12" pageNumber="13" specimenCount="1">
<location id="8EFE72EFDD75FFA0FE99FA9DFE5A88F3" LSID="urn:lsid:plazi:treatment:03889522DD70FFA0FF4AFDCBFD5E889A:8EFE72EFDD75FFA0FE99FA9DFE5A88F3" box="[320,469,1407,1426]" country="Australia" latitude="-27.413683" longLatPrecision="1" longitude="161.70035" name="Lord Howe Rise" pageId="12" pageNumber="13">Lord Howe Rise</location>
,
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,
<geoCoordinate id="EE1542F3DD75FFA0FD5EFA9DFD7088F3" box="[647,767,1407,1426]" degrees="161.70035" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="161.70035">161.70035° E</geoCoordinate>
,
<quantity id="4CD989D1DD75FFA0FF55FA7BFF5B88CD" box="[140,212,1433,1452]" metricMagnitude="3" metricUnit="m" metricValue="1.5319999999999998" pageId="12" pageNumber="13" unit="m" value="1532.0">
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.
<collectorName id="26D441E2DD75FFA0FF39FA7BFEF088CD" box="[224,383,1433,1452]" pageId="12" pageNumber="13">Coll. Geoscience</collectorName>
<collectingCountry id="F33664A4DD75FFA0FE5EFA7BFE5488CD" box="[391,475,1433,1452]" name="Australia" pageId="12" pageNumber="13">Australia</collectingCountry>
aboard
</materialsCitation>
<materialsCitation id="3B492E69DD75FFA0FDF5FA7BFEAC88A6" collectingDate="2007-10-17" collectionCode="R, V" country="Australia" pageId="12" pageNumber="13" specimenCount="1">
<collectionCode id="ED30BCF1DD75FFA0FDF5FA7BFDB488CD" box="[556,571,1433,1452]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
/
<collectionCode id="ED30BCF1DD75FFA0FD9BFA7BFDDD88CD" box="[578,594,1433,1452]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="13" type="Museum">V</collectionCode>
<emphasis id="B955F826DD75FFA0FD85FA78FD3488CD" box="[604,699,1434,1452]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C215FB7DD75FFA0FD85FA78FD3888CD" box="[604,695,1434,1452]" class="Arachnida" family="Uloboridae" genus="Tangaroa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Tangaroa</taxonomicName>
,
</emphasis>
<date id="FF9F02F4DD75FFA0FD1AFA7BFF3488A7" pageId="12" pageNumber="13" value="2007-10-17">
<collectingDate id="EFDBFB1CDD75FFA0FD1AFA7BFF3488A7" pageId="12" pageNumber="13" value="2007-10-17">17 Oct 2007</collectingDate>
</date>
. 1 wet spec
</materialsCitation>
, R=3.9, r=1.8.
</paragraph>
<paragraph id="8B9E2434DD75FFA0FF6BFA2CFD5E889A" blockId="12.[140,772,1170,1531]" pageId="12" pageNumber="13">
<materialsCitation id="3B492E69DD75FFA0FF6BFA2CFDD9889A" collectionCode="MNHN" country="New Caledonia" elevation="1390" latitude="-23.028334" location="New Caledonia" longLatPrecision="125" longitude="168.155" pageId="12" pageNumber="13" specimenCode="IE-2013-7118" specimenCount="1">
<collectionCode id="ED30BCF1DD75FFA0FF6BFA2CFF778880" box="[178,248,1486,1505]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="13">MNHN</collectionCode>
<specimenCode id="DB878C4FDD75FFA0FF25FA2CFEFB8880" box="[252,372,1486,1505]" collectionCode="IE-" pageId="12" pageNumber="13">IE-2013-7118</specimenCode>
<collectingCountry id="F33664A4DD75FFA0FEA1FA2CFD8B8881" box="[376,516,1485,1505]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
,
<geoCoordinate id="EE1542F3DD75FFA0FDD2FA2CFDD38880" box="[523,604,1486,1505]" degrees="23" direction="south" minutes="1.7" orientation="latitude" pageId="12" pageNumber="13" precision="92" value="-23.028334">23º 1.7'S</geoCoordinate>
,
<geoCoordinate id="EE1542F3DD75FFA0FDBCFA2FFD4B8880" box="[613,708,1485,1505]" degrees="168" direction="east" minutes="9.3" orientation="longitude" pageId="12" pageNumber="13" precision="92" value="168.155">168º 9.3' E</geoCoordinate>
,
<quantity id="4CD989D1DD75FFA0FD12FA2CFF5E889A" metricMagnitude="3" metricUnit="m" metricValue="1.39" metricValueMax="1.4069999999999998" metricValueMin="1.373" pageId="12" pageNumber="13" unit="m" value="1390.0" valueMax="1407.0" valueMin="1373.0">
<elevation id="000CC307DD75FFA0FD12FA2CFF5E889A" metricMagnitude="3" metricUnit="m" metricValue="1.39" metricValueMax="1.4069999999999998" metricValueMin="1.373" pageId="12" pageNumber="13" unit="m" value="1390.0" valueMax="1407.0" valueMin="1373.0">1373 1407 m</elevation>
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. Coll. KANADEEP 2, CP 5108. 1 wet spec
</materialsCitation>
, R=5.3, r=1.6.
</paragraph>
</subSubSection>
</treatment>
</document>