treatments-xml/data/03/86/A4/0386A43AEF63A2228B85FF53FCB5F80C.xml
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<document id="F7B3834AB4B9014A397C8E27F16E6D8F" ID-CLB-Dataset="32564" ID-DOI="10.11646/zootaxa.4300.3.1" ID-GBIF-Dataset="e96ae80b-2946-4418-8940-07dd60d7728e" ID-ISSN="1175-5326" ID-Zenodo-Dep="838438" ID-ZooBank="554D8B46-D396-42FA-9604-6DA9DFA3EFE7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1501748083066" checkinUser="plazi" docAuthor="Molineri, Carlos &amp; Salles, Frederico F." docDate="2017" docId="0386A43AEF63A2228B85FF53FCB5F80C" docLanguage="en" docName="zootaxa.4300.3.1.pdf" docOrigin="Zootaxa 4300 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Campsurus pereirae Molineri &amp; Salles 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="321" masterDocId="FFBFDC42EF77A2368B12FFC4FF88FFB5" masterDocTitle="Review of selected species of Campsurus Eaton 1868 (Ephemeroptera: Polymitarcyidae), with description of eleven new species and a key to male imagos of the genus" masterLastPageNumber="354" masterPageNumber="301" pageNumber="321" updateTime="1698466192200" updateUser="ExternalLinkService">
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<mods:title id="FF94721D0054639239495DC1FFF12752">Review of selected species of Campsurus Eaton 1868 (Ephemeroptera: Polymitarcyidae), with description of eleven new species and a key to male imagos of the genus</mods:title>
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<mods:namePart id="0DEB4467403B01C9FE5E7F3DA0394CC6">Molineri, Carlos</mods:namePart>
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<mods:namePart id="8FDCDB4043D6ADAEA76B301BE2D3A2ED">Salles, Frederico F.</mods:namePart>
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<mods:title id="9F313706A2D95CD19D428A1CFDA451DA">Zootaxa</mods:title>
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<mods:date id="E38FC45F1A488DB117ACE9DB5678DE20">2017</mods:date>
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<treatment id="0386A43AEF63A2228B85FF53FCB5F80C" ID-DOI="http://doi.org/10.5281/zenodo.6033482" ID-GBIF-Taxon="132630402" ID-Zenodo-Dep="6033482" LSID="urn:lsid:plazi:treatment:0386A43AEF63A2228B85FF53FCB5F80C" httpUri="http://treatment.plazi.org/id/0386A43AEF63A2228B85FF53FCB5F80C" lastPageNumber="321" pageId="20" pageNumber="321">
<subSubSection id="C33546A7EF63A2228B85FF53FE18FF66" pageId="20" pageNumber="321" type="nomenclature">
<paragraph id="8B90152CEF63A2228B85FF53FE79FF07" blockId="20.[151,497,151,211]" box="[151,497,151,178]" pageId="20" pageNumber="321">
<heading id="D0D8A240EF63A2228B85FF53FE79FF07" bold="true" box="[151,497,151,178]" fontSize="11" level="1" pageId="20" pageNumber="321" reason="1">
<emphasis id="B95BC93EEF63A2228B85FF53FE79FF07" bold="true" box="[151,497,151,178]" pageId="20" pageNumber="321">
<taxonomicName id="4C2F6EAFEF63A2228B85FF53FE04FF04" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[151,396,151,177]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pereirae" status="sp. nov.">
<emphasis id="B95BC93EEF63A2228B85FF53FE04FF04" bold="true" box="[151,396,151,177]" italics="true" pageId="20" pageNumber="321">Campsurus pereirae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A2228A81FF5CFE79FF07" box="[403,497,152,178]" pageId="20" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B90152CEF63A2228B85FF7EFE18FF66" blockId="20.[151,497,151,211]" box="[151,400,186,211]" pageId="20" pageNumber="321">
(
<figureCitation id="131409A9EF63A2228B8DFF7EFEAFFF66" box="[159,295,186,211]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Figs. 7174</figureCitation>
,
<figureCitation id="131409A9EF63A2228A20FF7FFED8FF66" box="[306,336,187,211]" captionStart="FIGURES 85 93" captionStartId="24.[151,264,1909,1932]" captionTargetBox="[168,1418,342,1885]" captionTargetId="figure@24.[163,1418,337,1885]" captionTargetPageId="24" captionText="FIGURES 85 93. Campsurus povilla sp. nov., male imago: 85, genitalia, v. v.; 86, detail of pedestals, v. v.; 87, detail of penes, v. v. Nymph: 88, left mandibular tusk, detail of inner tubercles; 89, left mandible, occlusal view (the basal part and molae were displaced during slide preparation; 90, same, detail of apex; 91, right mandible, occlusal view. Female abdominal sternum VIII with detail of socket: 92, C. povilla sp. nov.; 93, C. zunigae sp. nov." httpUri="https://zenodo.org/record/838456/files/figure.png" pageId="20" pageNumber="321">93</figureCitation>
,
<figureCitation id="131409A9EF63A2228A4FFF7EFE00FF66" box="[349,392,186,211]" captionStart="FIGURES 167 170" captionStartId="44.[151,264,1960,1983]" captionTargetBox="[257,1322,202,1910]" captionTargetId="figure@44.[246,1329,193,1922]" captionTargetPageId="44" captionText="FIGURES 167 170. Male imago, general view, dorsal: 167, C. dasilvai sp. nov.; 168, C. zunigae sp. nov.; 169, C. pereirae sp. nov.; 170, C. janae sp. nov." httpUri="https://zenodo.org/record/838480/files/figure.png" pageId="20" pageNumber="321">169</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33546A7EF63A2228B85FEC6FE26FA2B" pageId="20" pageNumber="321" type="materials_examined">
<paragraph id="8B90152CEF63A2228B85FEC6FAEFFE8A" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="321">
<materialsCitation id="3B471F71EF63A2228B85FEC6FAEFFE8A" ID-GBIF-Occurrence="1585352604" collectingDate="2012-02-26" collectionCode="APM, CZNC" collectorName="Parque Nacional das Emas &amp; LF Sgarbi &amp; Santos, E" country="Brazil" latitude="-18.26111" longLatPrecision="21" longitude="-52.888336" pageId="20" pageNumber="321" specimenCount="2" specimenCount-male="2" stateProvince="Goias" typeStatus="holotype">
<emphasis id="B95BC93EEF63A2228B85FEC6FEC2FEAE" bold="true" box="[151,330,258,283]" pageId="20" pageNumber="321">
<typeStatus id="5494AB8EEF63A2228B85FEC6FF5AFEAE" box="[151,210,258,283]" pageId="20" pageNumber="321">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5494AB8EEF63A2228A44FEC6FE36FEAE" box="[342,446,258,283]" pageId="20" pageNumber="321" type="holotype">Holotype</typeStatus>
<specimenCount id="9D29DEA5EF63A2228AD8FEC6FE57FEAE" box="[458,479,258,283]" pageId="20" pageNumber="321" type="male"></specimenCount>
i from
<collectingCountry id="F33855BCEF63A2228924FEC7FD16FEAE" box="[566,670,259,283]" name="Brazil" pageId="20" pageNumber="321">BRAZIL</collectingCountry>
,
<collectingRegion id="49EBDBCEEF63A22289BCFEC6FD7AFEAE" box="[686,754,258,283]" country="Brazil" name="Goias" pageId="20" pageNumber="321">Goiás</collectingRegion>
,
<collectorName id="26DA70FAEF63A2228811FEC7FBC9FEAF" box="[771,1089,258,283]" pageId="20" pageNumber="321">Parque Nacional das Emas</collectorName>
,
<collectingDate id="EFD5CA04EF63A2228F43FEC6FB4DFEAE" box="[1105,1221,258,283]" pageId="20" pageNumber="321" value="2012-02-26">
<date id="FF9133ECEF63A2228F43FEC6FB4DFEAE" box="[1105,1221,258,283]" pageId="20" pageNumber="321" value="2012-02-26">26.ii.2012</date>
</collectingDate>
,
<geoCoordinate id="EE1B73EBEF63A2228FC4FEC7FAEAFEAE" box="[1238,1378,258,283]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="15" value="-18.26111">S 18°15'40&quot;</geoCoordinate>
/
<geoCoordinate id="EE1B73EBEF63A2228E90FEC7FE81FE8A" direction="west" orientation="longitude" pageId="20" pageNumber="321" precision="15" value="-52.888336">W 52°53'18&quot;</geoCoordinate>
,
<collectorName id="26DA70FAEF63A2228A06FEE3FE00FE8A" box="[276,392,294,319]" pageId="20" pageNumber="321">LF Sgarbi</collectorName>
,
<collectionCode id="ED3E8DE9EF63A2228A81FEE3FE58FE8A" box="[403,464,295,319]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15304" name="Algonquin Provincial Park" pageId="20" pageNumber="321">APM</collectionCode>
<collectorName id="26DA70FAEF63A2228AC5FEE3FDCCFE8B" box="[471,580,295,319]" pageId="20" pageNumber="321">Santos, E</collectorName>
. Raimundi cols. (
<collectionCode id="ED3E8DE9EF63A2228815FEE3FCD0FE8A" box="[775,856,295,319]" pageId="20" pageNumber="321">CZNC</collectionCode>
).
<typeStatus id="5494AB8EEF63A2228879FEE3FC54FE8A" box="[875,988,295,319]" pageId="20" pageNumber="321" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D29DEA5EF63A22288F4FEE3FB98FE8A" box="[998,1040,294,319]" pageId="20" pageNumber="321" type="male">1 ♂</specimenCount>
i same data as holotype (IBN).
</materialsCitation>
</paragraph>
<paragraph id="8B90152CEF63A2228BD5FE8EFD11FE7A" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="321">
<materialsCitation id="3B471F71EF63A2228BD5FE8EFD11FE7A" ID-GBIF-Occurrence="1585352610" collectingDate="1978-12-27" collectingDateMax="2012-02-24" collectingDateMin="1978-12-27" collectionCode="APM, CZNC, FAMU" collectorName="LF Sgarbi &amp; Santos, E &amp; Brokopondo &amp; JG Peters" country="Brazil" elevation="22514" latitude="5.0" location="Brokopondo district" longLatPrecision="1305" longitude="-54.983334" municipality="Mineiros" pageId="20" pageNumber="321" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Goias">
<emphasis id="B95BC93EEF63A2228BD5FE8EFE33FED6" bold="true" box="[199,443,330,355]" pageId="20" pageNumber="321">Additional material.</emphasis>
<collectingCountry id="F33855BCEF63A2228AD1FE8FFDA5FED6" box="[451,557,331,355]" name="Brazil" pageId="20" pageNumber="321">BRAZIL</collectingCountry>
:
<specimenCount id="9D29DEA5EF63A222892AFE8FFDE9FED6" box="[568,609,330,355]" pageId="20" pageNumber="321" type="female">1 ♀</specimenCount>
i from
<collectingRegion id="49EBDBCEEF63A22289A0FE8EFD7EFED6" box="[690,758,330,355]" country="Brazil" name="Goias" pageId="20" pageNumber="321">Goiás</collectingRegion>
,
<collectingMunicipality id="6BF48F56EF63A2228811FE8EFCE3FED6" box="[771,875,330,355]" pageId="20" pageNumber="321">Mineiros</collectingMunicipality>
, riacho/ponte, L
<elevation id="0002F21FEF63A2228F37FE8FFB11FED7" box="[1061,1177,330,355]" metricMagnitude="4" metricUnit="m" metricValue="2.2514" pageId="20" pageNumber="321" unit="m" value="22514.0">
<quantity id="4CD7B8C9EF63A2228F37FE8FFB11FED7" box="[1061,1177,330,355]" metricMagnitude="4" metricUnit="m" metricValue="2.2514" pageId="20" pageNumber="321" unit="m" value="22514.0">22, 514 m</quantity>
</elevation>
,
<geoCoordinate id="EE1B73EBEF63A2228FB4FE8FFAB4FED6" box="[1190,1340,330,355]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="15" value="-17.210001">S 17° 12' 36&quot;</geoCoordinate>
/
<geoCoordinate id="EE1B73EBEF63A2228E46FE8FFF61FE32" direction="west" orientation="longitude" pageId="20" pageNumber="321" precision="15" value="-52.660835">W 52° 39' 39&quot;</geoCoordinate>
,
<collectingDate id="EFD5CA04EF63A2228BE6FEAAFEE0FE32" box="[244,360,366,391]" pageId="20" pageNumber="321" value="2012-02-24">
<date id="FF9133ECEF63A2228BE6FEAAFEE0FE32" box="[244,360,366,391]" pageId="20" pageNumber="321" value="2012-02-24">24.ii.2012</date>
</collectingDate>
,
<collectingMethod id="526E6D3BEF63A2228A66FEAAFDFFFE33" box="[372,631,366,391]" pageId="20" pageNumber="321">pennsylvania light trap</collectingMethod>
,
<collectorName id="26DA70FAEF63A2228991FEABFD7FFE32" box="[643,759,366,391]" pageId="20" pageNumber="321">LF Sgarbi</collectorName>
,
<collectionCode id="ED3E8DE9EF63A2228810FEABFCB7FE32" box="[770,831,367,391]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15304" name="Algonquin Provincial Park" pageId="20" pageNumber="321">APM</collectionCode>
<collectorName id="26DA70FAEF63A2228854FEABFC3BFE33" box="[838,947,367,391]" pageId="20" pageNumber="321">Santos, E</collectorName>
. Raimundi cols. (
<collectionCode id="ED3E8DE9EF63A2228F65FEABFB40FE32" box="[1143,1224,367,391]" pageId="20" pageNumber="321">CZNC</collectionCode>
). SURINAM:
<specimenCount id="9D29DEA5EF63A2228E78FEABFA1CFE32" box="[1386,1428,366,391]" pageId="20" pageNumber="321" type="male">3 ♂</specimenCount>
i (
<collectionCode id="ED3E8DE9EF63A2228B8CFE57FF7AFE1E" box="[158,242,403,427]" pageId="20" pageNumber="321">FAMU</collectionCode>
, except
<specimenCount id="9D29DEA5EF63A2228A5CFE57FEF0FE1E" box="[334,376,402,427]" pageId="20" pageNumber="321" type="male">1 ♂</specimenCount>
i IBN759CM) from
<location id="8EF043F7EF63A2228949FE56FCB7FE1E" LSID="urn:lsid:plazi:treatment:0386A43AEF63A2228B85FF53FCB5F80C:8EF043F7EF63A2228949FE56FCB7FE1E" box="[603,831,402,427]" country="Brazil" latitude="5.0" longLatPrecision="1305" longitude="-54.983334" municipality="Mineiros" name="Brokopondo district" pageId="20" pageNumber="321" stateProvince="Goias">Brokopondo district</location>
, creek on N edge
<collectorName id="26DA70FAEF63A2228F1BFE56FB1DFE1E" box="[1033,1173,402,427]" pageId="20" pageNumber="321">Brokopondo</collectorName>
(approx.
<geoCoordinate id="EE1B73EBEF63A2228E10FE56FACEFE1E" box="[1282,1350,402,427]" direction="north" orientation="latitude" pageId="20" pageNumber="321" precision="925" value="5.0">N5° 0'</geoCoordinate>
/
<geoCoordinate id="EE1B73EBEF63A2228E44FE57FF33FE7A" direction="west" orientation="longitude" pageId="20" pageNumber="321" precision="925" value="-54.983334">W 54° 59'</geoCoordinate>
),
<quantity id="4CD7B8C9EF63A2228BDDFE73FE82FE7B" box="[207,266,439,463]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="321" unit="m" value="90.0">
<elevation id="0002F21FEF63A2228BDDFE73FE82FE7B" box="[207,266,439,463]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="321" unit="m" value="90.0">90 m</elevation>
</quantity>
,
<collectingDate id="EFD5CA04EF63A2228A04FE72FE10FE7A" box="[278,408,438,463]" pageId="20" pageNumber="321" value="1978-12-27">
<date id="FF9133ECEF63A2228A04FE72FE10FE7A" box="[278,408,438,463]" pageId="20" pageNumber="321" value="1978-12-27">27.xii.1978</date>
</collectingDate>
, WL &amp;
<collectorName id="26DA70FAEF63A2228AE1FE73FDD5FE7B" box="[499,605,439,463]" pageId="20" pageNumber="321">JG Peters</collectorName>
cols.
</materialsCitation>
</paragraph>
<paragraph id="8B90152CEF63A2228BD5FE1EFCB2FBEF" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="321">
Male imago. Length (mm): body, 6.06.5; fore wing, 6.07.0; hind wing, 2.9; cerci, 17.020.0; fore leg, 3.6. General coloration yellowish, widely shaded with gray (
<figureCitation id="131409A9EF63A222880CFE3AFC0BFDA2" box="[798,899,510,535]" captionStart="FIGURES 167 170" captionStartId="44.[151,264,1960,1983]" captionTargetBox="[257,1322,202,1910]" captionTargetId="figure@44.[246,1329,193,1922]" captionTargetPageId="44" captionText="FIGURES 167 170. Male imago, general view, dorsal: 167, C. dasilvai sp. nov.; 168, C. zunigae sp. nov.; 169, C. pereirae sp. nov.; 170, C. janae sp. nov." httpUri="https://zenodo.org/record/838480/files/figure.png" pageId="20" pageNumber="321">Fig. 169</figureCitation>
). Head completely shaded with black dorsally, brownish gray on gena, antenna (except for hyaline flagellum) and ventrally. Thorax (
<figureCitation id="131409A9EF63A2228FB8FDE6FA9BFD8E" box="[1194,1299,546,571]" captionStart="FIGURES 167 170" captionStartId="44.[151,264,1960,1983]" captionTargetBox="[257,1322,202,1910]" captionTargetId="figure@44.[246,1329,193,1922]" captionTargetPageId="44" captionText="FIGURES 167 170. Male imago, general view, dorsal: 167, C. dasilvai sp. nov.; 168, C. zunigae sp. nov.; 169, C. pereirae sp. nov.; 170, C. janae sp. nov." httpUri="https://zenodo.org/record/838480/files/figure.png" pageId="20" pageNumber="321">Fig. 169</figureCitation>
). Pronotum translucent, completely shaded with gray, darker on anterior portion, medial line, and margins; prosternum yellowish white, shaded with gray; propleura anteriorly grayish, posteriorly black. Mesonotum yellowish, shaded with gray on medial line, parapsidal sutures, and between
<collectionCode id="ED3E8DE9EF63A222882FFD4BFCE7FD12" box="[829,879,655,679]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15543" name="Parasitic Seed Plants" pageId="20" pageNumber="321">PSP</collectionCode>
; pleura and sterna yellowish white, shaded with gray; sutures above coxa blackish; furcasternum shaded with gray. Metanotum yellowish white, shaded with gray medially. Legs. Fore leg completely shaded gray, slightly paler at base of tarsomeres; middle and hind legs whitish, shaded completely with gray. Wings. Membrane hyaline, except C and Sc areas shaded with brownish gray, veins hyaline, except C, Sc, and R1 brownish. Hind wing hyaline, except C and Sc areas very slightly shaded with gray. Abdomen (
<figureCitation id="131409A9EF63A2228A04FC81FEFFFCEB" box="[278,375,837,862]" captionStart="FIGURES 167 170" captionStartId="44.[151,264,1960,1983]" captionTargetBox="[257,1322,202,1910]" captionTargetId="figure@44.[246,1329,193,1922]" captionTargetPageId="44" captionText="FIGURES 167 170. Male imago, general view, dorsal: 167, C. dasilvai sp. nov.; 168, C. zunigae sp. nov.; 169, C. pereirae sp. nov.; 170, C. janae sp. nov." httpUri="https://zenodo.org/record/838480/files/figure.png" pageId="20" pageNumber="321">Fig. 169</figureCitation>
) whitish translucent, widely and strongly shaded with gray on terga, except on pleural folds. Abdominal sterna translucent white, shaded with gray, slightly darker on medial line and inner margin of gill sclerites. Genitalia (
<figureCitation id="131409A9EF63A2228A64FC49FD89FC13" box="[374,513,909,934]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Figs. 7174</figureCitation>
): yellowish white; sternum IX with truncate hind margin (
<figureCitation id="131409A9EF63A2228F9BFC49FB54FC13" box="[1161,1244,909,934]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Fig. 71</figureCitation>
), medial line and hind margin black; pedestal of each side touching basally, with apically acute parastylus (not visible in
<figureCitation id="131409A9EF63A2228E29FC75FA07FC7F" box="[1339,1423,945,970]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Fig. 72</figureCitation>
), round inner corner strongly projected; forceps short and translucent, shaded with gray; base of main lobe of penes sclerotized, strongly expanded laterally and proximally, with dorsal surface finely granulated (arrow in
<figureCitation id="131409A9EF63A2228E28FC3DFA07FBA7" box="[1338,1423,1017,1042]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Fig. 73</figureCitation>
), apical 1/3 narrowing abruptly, rounded and curved ventrally (
<figureCitation id="131409A9EF63A222884FFBD9FC3CFB83" box="[861,948,1053,1078]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Fig. 74</figureCitation>
); secondary membranous lobe translucent, slender and cylindrical. Caudal filament translucent whitish.
</paragraph>
<paragraph id="8B90152CEF63A2228BD5FBA1FB57FAE3" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="321">
Female imago. Length (mm): body, 9.2; fore wing, 9.0; hind wing and cerci broken. Similar to male imago, except pigmentation more distinct and few other differences, as follows. Thorax. Pronotum shaded with gray, except pale transverse line between anterior and posterior portions, and pale longitudinal sublateral band; medial line concolorous with rest of notum. Mesonotum additionally shaded gray on anterior triangular mark (after mesonotal protuberance). Middle and hind legs shaded more slightly with gray. Wing membranes yellowish. Abdomen ventrally shaded only submedially and laterally on sterna VIIIX; sternum VIII with small anteromedian subquadrate socket (
<figureCitation id="131409A9EF63A2228A6EFAF9FE47FAE3" box="[380,463,1341,1366]" captionStart="FIGURES 85 93" captionStartId="24.[151,264,1909,1932]" captionTargetBox="[168,1418,342,1885]" captionTargetId="figure@24.[163,1418,337,1885]" captionTargetPageId="24" captionText="FIGURES 85 93. Campsurus povilla sp. nov., male imago: 85, genitalia, v. v.; 86, detail of pedestals, v. v.; 87, detail of penes, v. v. Nymph: 88, left mandibular tusk, detail of inner tubercles; 89, left mandible, occlusal view (the basal part and molae were displaced during slide preparation; 90, same, detail of apex; 91, right mandible, occlusal view. Female abdominal sternum VIII with detail of socket: 92, C. povilla sp. nov.; 93, C. zunigae sp. nov." httpUri="https://zenodo.org/record/838456/files/figure.png" pageId="20" pageNumber="321">Fig. 93</figureCitation>
). Caudal filaments translucent whitish (all except basal segment lost).
</paragraph>
<paragraph id="8B90152CEF63A2228BD5FAA6FE26FA2B" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="321">Egg. Length, 250 µm; width, 200 µm. Yellowish with whitish polar cap. Polar cap relatively large (width 0.7× maximum width of egg).</paragraph>
</subSubSection>
<subSubSection id="C33546A7EF63A2228BD5FA6DFE52FA53" pageId="20" pageNumber="321" type="etymology">
<paragraph id="8B90152CEF63A2228BD5FA6DFE52FA53" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="321">
<emphasis id="B95BC93EEF63A2228BD5FA6DFEC6FA77" bold="true" box="[199,334,1449,1474]" pageId="20" pageNumber="321">Etymology.</emphasis>
We are pleased to name this species after Sueli Pereira, a mayfly specialist who kindly provided material treated in this work.
</paragraph>
</subSubSection>
<subSubSection id="C33546A7EF63A2228BD5FA35FD7AF9BF" box="[199,754,1521,1546]" pageId="20" pageNumber="321" type="distribution">
<paragraph id="8B90152CEF63A2228BD5FA35FD7AF9BF" blockId="20.[151,1437,258,1978]" box="[199,754,1521,1546]" pageId="20" pageNumber="321">
<emphasis id="B95BC93EEF63A2228BD5FA35FED1F9BF" bold="true" box="[199,345,1521,1546]" pageId="20" pageNumber="321">Distribution</emphasis>
(
<figureCitation id="131409A9EF63A2228A7AFA35FE43F9BF" box="[360,459,1521,1546]" captionStart="FIGURES 179" captionStartId="48.[151,264,1621,1644]" captionTargetBox="[164,1412,193,1595]" captionTargetId="figure@48.[164,1412,193,1595]" captionTargetPageId="48" captionText="FIGURES 179. Distribution maps." httpUri="https://zenodo.org/record/838488/files/figure.png" pageId="20" pageNumber="321">Fig. 179</figureCitation>
).
<collectingCountry id="F33855BCEF63A2228ACCFA35FDAAF9BF" box="[478,546,1521,1546]" name="Brazil" pageId="20" pageNumber="321">Brazil</collectingCountry>
(
<collectingRegion id="49EBDBCEEF63A2228923FA35FDF0F9BF" box="[561,632,1521,1546]" country="Brazil" name="Goias" pageId="20" pageNumber="321">Goiás</collectingRegion>
),
<collectingCountry id="F33855BCEF63A222899EFA35FD66F9BF" box="[652,750,1521,1546]" name="Suriname" pageId="20" pageNumber="321">Surinam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33546A7EF63A2228BD5F9D1FB69F8B3" pageId="20" pageNumber="321" type="diagnosis">
<paragraph id="8B90152CEF63A2228BD5F9D1FB69F8B3" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="321">
<emphasis id="B95BC93EEF63A2228BD5F9D1FEC9F99B" bold="true" box="[199,321,1557,1582]" pageId="20" pageNumber="321">Diagnosis.</emphasis>
<taxonomicName id="4C2F6EAFEF63A2228A5AF9D2FDA5F998" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[328,557,1558,1581]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pereirae" status="sp. nov.">
<emphasis id="B95BC93EEF63A2228A5AF9D2FDA5F998" box="[328,557,1558,1581]" italics="true" pageId="20" pageNumber="321">Campsurus pereirae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A2228927F9D1FD07F99B" box="[565,655,1557,1582]" pageId="20" pageNumber="321" rank="species">
<emphasis id="B95BC93EEF63A2228927F9D1FD07F99B" bold="true" box="[565,655,1557,1582]" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
is known from adults of both sexes, and it can be distinguished from other species of
<taxonomicName id="4C2F6EAFEF63A2228A5EF9FEFE43F9E4" authorityName="Eaton" authorityYear="1868" box="[332,459,1594,1617]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B95BC93EEF63A2228A5EF9FEFE43F9E4" box="[332,459,1594,1617]" italics="true" pageId="20" pageNumber="321">Campsurus</emphasis>
</taxonomicName>
by: 1) posterior margin of male abdominal sternum IX truncate (
<figureCitation id="131409A9EF63A2228F8CF9FDFB78F9E7" box="[1182,1264,1593,1618]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Fig. 71</figureCitation>
); 2) pedestal of each side touching basally, with apically acute parastylus (
<figureCitation id="131409A9EF63A222883EF999FC08F9C3" box="[812,896,1629,1654]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Fig. 72</figureCitation>
), round inner corner strongly projected; 3) base of main lobe of penes sclerotized, strongly expanded laterally and proximally with its dorsal surface finely granulated (
<figureCitation id="131409A9EF63A2228A0FF961FEF8F90B" box="[285,368,1701,1726]" captionStart="FIGURES 68 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="20" pageNumber="321">Fig. 73</figureCitation>
), apical 1/3 narrowing abruptly, rounded and slightly curved ventrally; secondary membranous lobe cylindrical and slender; 4) small size (length of male fore wings 6.57.0 mm); 5) female socket single, subquadrate (
<figureCitation id="131409A9EF63A2228A3FF929FE09F8B3" box="[301,385,1773,1798]" captionStart="FIGURES 85 93" captionStartId="24.[151,264,1909,1932]" captionTargetBox="[168,1418,342,1885]" captionTargetId="figure@24.[163,1418,337,1885]" captionTargetPageId="24" captionText="FIGURES 85 93. Campsurus povilla sp. nov., male imago: 85, genitalia, v. v.; 86, detail of pedestals, v. v.; 87, detail of penes, v. v. Nymph: 88, left mandibular tusk, detail of inner tubercles; 89, left mandible, occlusal view (the basal part and molae were displaced during slide preparation; 90, same, detail of apex; 91, right mandible, occlusal view. Female abdominal sternum VIII with detail of socket: 92, C. povilla sp. nov.; 93, C. zunigae sp. nov." httpUri="https://zenodo.org/record/838456/files/figure.png" pageId="20" pageNumber="321">Fig. 93</figureCitation>
); 6) egg with large polar cap, ratio maximum width of egg/width of PC =1.4.
</paragraph>
</subSubSection>
<subSubSection id="C33546A7EF63A2228BD5F8D5FCB5F80C" pageId="20" pageNumber="321" type="discussion">
<paragraph id="8B90152CEF63A2228BD5F8D5FCB5F80C" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="321">
<emphasis id="B95BC93EEF63A2228BD5F8D5FEC4F89F" bold="true" box="[199,332,1809,1834]" pageId="20" pageNumber="321">Discussion.</emphasis>
The extremely expanded penes and the fine granules present on them are unique to
<taxonomicName id="4C2F6EAFEF63A2228FE7F8D6FAFCF89C" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1269,1396,1810,1833]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pereirae" status="sp. nov.">
<emphasis id="B95BC93EEF63A2228FE7F8D6FAFCF89C" box="[1269,1396,1810,1833]" italics="true" pageId="20" pageNumber="321">C. pereirae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A2228E68F8D5FF40F8FB" pageId="20" pageNumber="321" rank="species">
<emphasis id="B95BC93EEF63A2228E68F8D5FF40F8FB" bold="true" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
, distinguishing this species from the closely related
<taxonomicName id="4C2F6EAFEF63A2228838F8F2FC59F8F8" authorityName="Traver ." authorityYear="1947" box="[810,977,1846,1869]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="cuyuniensis">
<emphasis id="B95BC93EEF63A2228838F8F2FC59F8F8" box="[810,977,1846,1869]" italics="true" pageId="20" pageNumber="321">C. cuyuniensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F6EAFEF63A22288F0F8F2FBF5F8F8" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[994,1149,1846,1869]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="nessimiani" status="sp. nov.">
<emphasis id="B95BC93EEF63A22288F0F8F2FBF5F8F8" box="[994,1149,1846,1869]" italics="true" pageId="20" pageNumber="321">C. nessimiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A2228F95F8F1FB6BF8FB" box="[1159,1251,1845,1870]" pageId="20" pageNumber="321" rank="species">
<emphasis id="B95BC93EEF63A2228F95F8F1FB6BF8FB" bold="true" box="[1159,1251,1845,1870]" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C2F6EAFEF63A2228FE6F8F2FAF9F8FB" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1268,1393,1845,1870]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="dasilvai" status="sp. nov.">
<emphasis id="B95BC93EEF63A2228FE6F8F2FAF9F8FB" box="[1268,1393,1845,1870]" italics="true" pageId="20" pageNumber="321">C. dasilvai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A2228E68F8F1FF40F8C7" pageId="20" pageNumber="321" rank="species">
<emphasis id="B95BC93EEF63A2228E68F8F1FF40F8C7" bold="true" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C2F6EAFEF63A2228BC5F89EFED9F8C4" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[215,337,1882,1905]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="zunigae" status="sp. nov.">
<emphasis id="B95BC93EEF63A2228BC5F89EFED9F8C4" box="[215,337,1882,1905]" italics="true" pageId="20" pageNumber="321">C. zunigae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A2228A4BF89DFE3CF8C7" box="[345,436,1881,1906]" pageId="20" pageNumber="321" rank="species">
<emphasis id="B95BC93EEF63A2228A4BF89DFE3CF8C7" bold="true" box="[345,436,1881,1906]" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C2F6EAFEF63A2228AFFF89EFDC6F8C4" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[493,590,1882,1905]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="janae" status="sp. nov.">
<emphasis id="B95BC93EEF63A2228AFFF89EFDC6F8C4" box="[493,590,1882,1905]" italics="true" pageId="20" pageNumber="321">C. janae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A2228944F89DFD39F8C7" box="[598,689,1881,1906]" pageId="20" pageNumber="321" rank="species">
<emphasis id="B95BC93EEF63A2228944F89DFD39F8C7" bold="true" box="[598,689,1881,1906]" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="4C2F6EAFEF63A22289A8F89EFC08F8C4" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[698,896,1882,1905]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="janae" status="sp. nov.">
<emphasis id="B95BC93EEF63A22289A8F89EFC08F8C4" box="[698,896,1882,1905]" italics="true" pageId="20" pageNumber="321">Campsurus janae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A222889BF89DFC6CF8C7" box="[905,996,1881,1906]" pageId="20" pageNumber="321" rank="species">
<emphasis id="B95BC93EEF63A222889BF89DFC6CF8C7" bold="true" box="[905,996,1881,1906]" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
also presents penes with an extremely wide base, but the penes are smooth, and the pedestals are much larger and shorter than in
<taxonomicName id="4C2F6EAFEF63A2228FA4F8BAFABFF820" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1206,1335,1918,1941]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pereirae" status="sp. nov.">
<emphasis id="B95BC93EEF63A2228FA4F8BAFABFF820" box="[1206,1335,1918,1941]" italics="true" pageId="20" pageNumber="321">C. pereirae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF63A2228E52F8B9FA14F823" box="[1344,1436,1917,1942]" pageId="20" pageNumber="321" rank="species">
<emphasis id="B95BC93EEF63A2228E52F8B9FA14F823" bold="true" box="[1344,1436,1917,1942]" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
Again, the finely granulated penes are unique to
<taxonomicName id="4C2F6EAFEF63A22289AAF866FCBFF80C" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[696,823,1954,1977]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pereirae">
<emphasis id="B95BC93EEF63A22289AAF866FCBFF80C" box="[696,823,1954,1977]" italics="true" pageId="20" pageNumber="321">C. pereirae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>