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<document id="EA1E62C3D2264C461D4F85C9FC4DB376" ID-CLB-Dataset="4300" ID-DOI="10.18590/euscorpius.2005.vol2005.iss19.1" ID-GBIF-Dataset="3d25137e-959e-4b79-bc0c-2db2f500ffcd" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648178" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617134361280" checkinUser="carolina" docAuthor="Lourenço, Wilson R. &amp; Geniez, Philippe" docDate="2005" docId="03866767F341866AA273F9F9FBFDF882" docLanguage="en" docName="Euscorpius.19.1-6.pdf.imf" docOrigin="Euscorpius 19 (19)" docSource="https://mds.marshall.edu/euscorpius/vol2005/iss19/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Buthus bonito Lourenço &amp; Geniez 2005, sp. n." docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FFBF1F1FF343866CA14DFFBFFFEEFFE3" masterDocTitle="A new scorpion species of the genus Buthus Leach, 1815 (Scorpiones, Buthidae) from Morocco" masterLastPageNumber="6" masterPageNumber="1" pageNumber="1" updateTime="1698935394561" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5052111B2451A829A3E5FA70CA8F9E78">A new scorpion species of the genus Buthus Leach, 1815 (Scorpiones, Buthidae) from Morocco</mods:title>
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<mods:namePart id="3ADB1E18FA6E9C854C69C490E16FDF99">Lourenço, Wilson R.</mods:namePart>
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<mods:namePart id="9DA29C65A02AB2338A1530497AF09018">Geniez, Philippe</mods:namePart>
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<mods:title id="1B33A6913214FBD82F7B86F895714738">Euscorpius</mods:title>
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<mods:date id="B4444305A9CD248949CDEE0A75401FA3">2005</mods:date>
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<mods:number id="6F119466445D56E90A42105274A0A158">2005-12-31</mods:number>
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<treatment id="03866767F341866AA273F9F9FBFDF882" ID-DOI="http://doi.org/10.5281/zenodo.5507119" ID-GBIF-Taxon="187973124" ID-Zenodo-Dep="5507119" LSID="urn:lsid:zoobank.org:act:F9C186CE-05F6-4710-98FA-C03ED648E3A5" httpUri="http://treatment.plazi.org/id/03866767F341866AA273F9F9FBFDF882" lastPageId="6" lastPageNumber="5" pageId="2" pageNumber="1">
<subSubSection id="C33585FAF341866EA273F9F9FB25F981" box="[830,1227,1606,1634]" pageId="2" pageNumber="1" type="nomenclature">
<paragraph id="8B90D671F341866EA273F9F9FB25F981" blockId="2.[830,1227,1606,1634]" box="[830,1227,1606,1634]" pageId="2" pageNumber="1">
<heading id="D0D8611DF341866EA273F9F9FB25F981" box="[830,1227,1606,1634]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1" reason="3">
<emphasis id="B95B0A63F341866EA273F9F9FB25F981" bold="true" box="[830,1227,1606,1634]" pageId="2" pageNumber="1">
<taxonomicName id="4C2FADF2F341866EA273F9F9FC04F982" ID-CoL="NYYT" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[830,1002,1606,1633]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F341866EA273F9F9FC04F982" bold="true" box="[830,1002,1606,1633]" italics="true" pageId="2" pageNumber="1">Buthus bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F341866EA2BFF9F7FBDAF981" box="[1010,1076,1608,1634]" pageId="2" pageNumber="1" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="1314CAF4F341866EA50EF9F7FB2CF981" box="[1091,1218,1608,1634]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–8" captionStart-2="Figure 9" captionStart-3="Figure 10" captionStartId-0="3.[1056,1142,822,845]" captionStartId-1="4.[192,278,1855,1878]" captionStartId-2="5.[192,268,902,925]" captionStartId-3="6.[192,268,1172,1195]" captionTargetBox-0="[157,1013,160,978]" captionTargetBox-1="[275,1376,163,1807]" captionTargetBox-2="[192,1440,76,879]" captionTargetBox-3="[189,1443,189,1131]" captionTargetId-0="figure-7@3.[149,1033,144,982]" captionTargetId-1="figure-10@4.[265,1402,154,1814]" captionTargetId-2="graphics-8@5.[264,1365,117,879]" captionTargetId-3="figure-10@6.[192,1440,192,1128]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figures 1–3: Buthus bonito sp. n. Male holotype. 1. Chelicera, dorsal aspect. 2. Metasomal segment V and telson, lateral aspect. 3. Granulation on the dentate margins of the pedipalp chela movable finger." captionText-1="Figures 4–8: Buthus bonito sp. n. Male holotype. Trichobothrial pattern. 4–5. Chela, dorso-external and ventral aspects. 6–7. Patella, dorsal and external aspects. 8. Femur, dorsal aspect." captionText-2="Figure 9: Map of Morocco, showing the site where the new species was collected (black star)." captionText-3="Figure 10: Juvenile specimen of Buthus bonito sp. n. just after molting process in its natural habitat. Note: this specimen was not collected." figureDoi-0="http://doi.org/10.5281/zenodo.4648180" figureDoi-1="http://doi.org/10.5281/zenodo.4648182" figureDoi-2="http://doi.org/10.5281/zenodo.4648184" figureDoi-3="http://doi.org/10.5281/zenodo.4648186" httpUri-0="https://zenodo.org/record/4648180/files/figure.png" httpUri-1="https://zenodo.org/record/4648182/files/figure.png" httpUri-2="https://zenodo.org/record/4648184/files/figure.png" httpUri-3="https://zenodo.org/record/4648186/files/figure.png" pageId="2" pageNumber="1">Figs. 1–10</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33585FAF341866EA273F938FAF7F919" pageId="2" pageNumber="1" type="materials_examined">
<paragraph id="8B90D671F341866EA273F938FAF7F919" blockId="2.[830,1440,1671,1786]" pageId="2" pageNumber="1">
<emphasis id="B95B0A63F341866EA273F938FC07F97D" bold="true" box="[830,1001,1671,1694]" pageId="2" pageNumber="1">Type material.</emphasis>
<materialsCitation id="3B47DC2CF341866EA2BEF938FBF6F95E" ID-GBIF-Occurrence="3352090301" collectorName="Khnifiss" country="Morocco" location="Tarfaya" pageId="2" pageNumber="1" specimenCount="1">
<collectingCountry id="F33896E1F341866EA2BEF938FBB8F97D" box="[1011,1110,1671,1694]" name="Morocco" pageId="2" pageNumber="1">Morocco</collectingCountry>
, northeast of
<location id="8EF080AAF341866EA5BDF938FAA8F97D" LSID="urn:lsid:plazi:treatment:03866767F341866AA273F9F9FBFDF882:8EF080AAF341866EA5BDF938FAA8F97D" box="[1264,1350,1671,1694]" country="Morocco" name="Tarfaya" pageId="2" pageNumber="1">Tarfaya</location>
, near to the
<collectorName id="26DAB3A7F341866EA227F919FC2BF95E" box="[874,965,1702,1725]" pageId="2" pageNumber="1">Khnifiss</collectorName>
lagoon
</materialsCitation>
,
<date id="FF91F0B1F341866EA567F919FB72F95E" box="[1066,1180,1702,1725]" pageId="2" pageNumber="1" value="2004-05-31">
31/
<collectionCode id="ED3E4EB4F341866EA501F919FBB1F95E" box="[1100,1119,1702,1725]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="1" type="Museum">V</collectionCode>
/2004
</date>
(M. Aymerich leg.);
<specimenCount id="9D291DF8F341866EA4DEF919FC9CF938" pageId="2" pageNumber="1" type="male">1 male</specimenCount>
<typeStatus id="549468D3F341866EA232F97BFC32F938" box="[895,988,1732,1755]" pageId="2" pageNumber="1" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D291DF8F341866EA2BCF97BFBBDF938" box="[1009,1107,1732,1755]" pageId="2" pageNumber="1" type="female">2 female</specimenCount>
<typeStatus id="549468D3F341866EA52CF97BFB29F938" box="[1121,1223,1732,1755]" pageId="2" pageNumber="1" type="paratype">paratypes</typeStatus>
<superScript id="7C5A7B39F341866EA58AF901FB3EF92E" attach="left" box="[1223,1232,1726,1741]" fontSize="6" pageId="2" pageNumber="1">*</superScript>
.
<materialsCitation id="3B47DC2CF341866EA5A9F97BFAF8F919" ID-GBIF-Occurrence="3352090302" collectorName="Paris" country="Morocco" location="Deposited" municipality="Museum" pageId="2" pageNumber="1" specimenCount="1">
<location id="8EF080AAF341866EA5A9F97BFABEF938" LSID="urn:lsid:plazi:treatment:03866767F341866AA273F9F9FBFDF882:8EF080AAF341866EA5A9F97BFABEF938" box="[1252,1360,1732,1755]" country="Morocco" municipality="Museum" name="Deposited" pageId="2" pageNumber="1">Deposited</location>
in the
<collectingMunicipality id="6BF44C0BF341866EA273F95CFC72F919" box="[830,924,1763,1786]" pageId="2" pageNumber="1">Muséum</collectingMunicipality>
national d’Histoire naturelle,
<collectorName id="26DAB3A7F341866EA590F95CFAF8F919" box="[1245,1302,1763,1786]" pageId="2" pageNumber="1">Paris</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<paragraph id="8B90D671F341866EA273F88EFCA9F8A3" blockId="2.[830,1440,1841,1920]" box="[830,839,1841,1856]" pageId="2" pageNumber="1">
<superScript id="7C5A7B39F341866EA273F88EFCA9F8A3" attach="left" box="[830,839,1841,1856]" fontSize="6" pageId="2" pageNumber="1">*</superScript>
</paragraph>
<paragraph id="8B90D671F341866EA216F886FC65F863" blockId="2.[830,1440,1841,1920]" pageId="2" pageNumber="1">female specimens are still alive. They will be subsequently deposited in the Muséum’s collections after biological observations.</paragraph>
<caption id="DF5086F9F340866FA56DFC89FB1CFC2A" ID-DOI="http://doi.org/10.5281/zenodo.4648180" ID-Zenodo-Dep="4648180" httpUri="https://zenodo.org/record/4648180/files/figure.png" pageId="3" pageNumber="2" startId="3.[1056,1142,822,845]" targetBox="[157,1013,160,978]" targetPageId="3">
<paragraph id="8B90D671F340866FA56DFC89FB1CFC2A" blockId="3.[1056,1442,822,969]" pageId="3" pageNumber="2">
<emphasis id="B95B0A63F340866FA56DFC89FB59FCAE" bold="true" box="[1056,1207,822,845]" pageId="3" pageNumber="2">Figures 1–3:</emphasis>
<taxonomicName id="4C2FADF2F340866FA58BFC88FABAFCAF" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[1222,1364,823,844]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F340866FA58BFC88FABAFCAF" box="[1222,1364,823,844]" italics="true" pageId="3" pageNumber="2">Buthus bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F340866FA429FC87FA4FFCAE" box="[1380,1441,824,845]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
Male holotype.
<emphasis id="B95B0A63F340866FA583FCEEFB0FFC85" bold="true" box="[1230,1249,849,870]" pageId="3" pageNumber="2">1.</emphasis>
Chelicera, dorsal aspect.
<emphasis id="B95B0A63F340866FA523FCD5FB6FFC9C" bold="true" box="[1134,1153,874,895]" pageId="3" pageNumber="2">2.</emphasis>
Metasomal segment V and telson, lateral aspect.
<emphasis id="B95B0A63F340866FA5BBFC3CFAE6FC7B" bold="true" box="[1270,1288,899,920]" pageId="3" pageNumber="2">3.</emphasis>
Granulation on the dentate margins of the pedipalp chela movable finger.
</paragraph>
</caption>
<subSubSection id="C33585FAF340866FA18DFC4AFF1BFB46" pageId="3" pageNumber="2" type="etymology">
<paragraph id="8B90D671F340866FA18DFC4AFF1BFB46" blockId="3.[192,802,1013,1189]" pageId="3" pageNumber="2">
<emphasis id="B95B0A63F340866FA18DFC4AFEACFBEF" bold="true" box="[192,322,1013,1036]" pageId="3" pageNumber="2">Etymology.</emphasis>
The name ‘bonito’ (beautiful in Spanish), makes reference to the beautiful color of the new species. Curiously, Spanish still is one of the European languages used in the extreme southwest of
<collectingCountry id="F33896E1F340866FA3D6FBEEFD15FB8B" box="[667,763,1105,1128]" name="Morocco" pageId="3" pageNumber="2">Morocco</collectingCountry>
, an area corresponding to a part of the former
<collectingCountry id="F33896E1F340866FA3EDFBCFFF00FB46" name="Western Sahara" pageId="3" pageNumber="2">Spanish Sahara</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33585FAF340866FA18DFB73FDA5F992" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8B90D671F340866FA18DFB73FEC0FB00" blockId="3.[192,803,1228,1649]" box="[192,302,1228,1251]" pageId="3" pageNumber="2">
<emphasis id="B95B0A63F340866FA18DFB73FEC0FB00" bold="true" box="[192,302,1228,1251]" pageId="3" pageNumber="2">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B90D671F340866FA18DFB55FDA5F992" blockId="3.[192,803,1228,1649]" pageId="3" pageNumber="2">
Scorpions of medium size, reaching a total length of
<quantity id="4CD77B94F340866FA247FB55FEDCFAC3" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="3" pageNumber="2" unit="mm" value="52.5" valueMax="55.0" valueMin="50.0">50 to 55 mm</quantity>
. General coloration very pale yellow, with only some slightly darker zones over the carapace and median carina of tergites, chelicera finger denticles, oblique rows of granules on pedipalp fingers and extremity of aculeus. These parts are blackish in juveniles that exhibit a more contrasted pattern than that of
<taxonomicName id="4C2FADF2F340866FA395FA1DFEFEFA3B" authorityName="Pocock. The" authorityYear="1889" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F340866FA395FA1DFEFEFA3B" italics="true" pageId="3" pageNumber="2">Buthus atlantis</emphasis>
</taxonomicName>
. Venter, pedipalps and legs very pale yellow. Carinae and granulations moderate to weak. Fixed and movable fingers with 10–11 rows of granules in both sexes. Pectines with 34–33 teeth in male and 29–29 and 34–
<quantity id="4CD77B94F340866FA1A5F983FECCF9B0" box="[232,290,1596,1619]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="3" pageNumber="2" unit="in" value="34.0">34 in</quantity>
females. Male pectines very long and largely overlapping in their proximal region.
</paragraph>
</subSubSection>
<subSubSection id="C33585FAF340866AA18DF927FBFDF882" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2" type="description">
<paragraph id="8B90D671F340866FA18DF927FED5F92D" blockId="3.[192,802,1688,1895]" pageId="3" pageNumber="2">
<emphasis id="B95B0A63F340866FA18DF927FED5F92D" bold="true" pageId="3" pageNumber="2">
Description based on male
<typeStatus id="549468D3F340866FA34AF927FD87F94C" box="[519,617,1688,1711]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
(measurements in
<tableCitation id="C6ADE3CAF340866FA190F908FEDCF92D" box="[221,306,1719,1742]" captionStart="Table 1" captionStartId="5.[192,257,1857,1880]" captionTargetBox="[241,626,1054,1782]" captionTargetId="graphics-119@5.[187,780,987,1813]" captionText="Table 1: Morphometric values (in mm) of the male holotype of Buthus bonito sp. n." httpUri="http://table.plazi.org/id/DF5086F9F3468669A18DF8FEFE74F893" pageId="3" pageNumber="2" tableUuid="DF5086F9F3468669A18DF8FEFE74F893">Table 1</tableCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B90D671F340866FA18DF96BFBC6FAE2" blockId="3.[192,802,1688,1895]" lastBlockId="3.[830,1441,1013,1895]" pageId="3" pageNumber="2">
<emphasis id="B95B0A63F340866FA18DF96BFED3F90F" bold="true" box="[192,317,1748,1772]" italics="true" pageId="3" pageNumber="2">Coloration.</emphasis>
Basically very pale yellow, with only some slightly darker zones over the carapace and median carina of tergites, chelicera finger denticles, oblique rows of granules on pedipalp fingers and extremity of aculeus. Prosoma: carapace yellowish, with carinae slightly dark brown; eyes surrounded by black pigment. Mesosoma: tergites yellowish; median carina brownish. Metasoma: all segments and vesicle yellowish; aculeus yellowish at its base and blackish at its extremity. Venter pale yellow. Chelicerae yellowish without any variegated spots; fingers yellowish with reddish to blackish teeth. Pedipalps: yellowish; fingers with the oblique rows of granules dark reddish to blackish. Legs yellowish without any spots.
</paragraph>
<paragraph id="8B90D671F340866AA273FAB7FE80FA9E" blockId="3.[830,1441,1013,1895]" lastBlockId="6.[192,802,1257,1405]" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2">
<emphasis id="B95B0A63F340866FA273FAB7FC22FAC3" bold="true" box="[830,972,1288,1312]" italics="true" pageId="3" pageNumber="2">Morphology.</emphasis>
Carapace moderately granular; anterior margin with a very weak concavity, almost straight. Carinae moderate; anterior median, central median and posterior median carinae moderately granular; ‘lyre’ configuration can, however, be well observed. All furrows moderate. Median ocular tubercle at the centre of carapace. Eyes separated by two and half ocular diameters. Four pairs of lateral eyes: the first three of moderate size, the last one only vestigial. Sternum very small and triangular, wider than long. Mesosoma: tergites with a thin but intense granulation. Three longitudinal carinae; lateral carinae absent in tergites I and II; vestigial on III and reduced on IV to VI. Tergite VII pentacarinate. Venter: genital operculum divided longitudinally, and formed by two semi oval plates. Pectines: pectinal tooth count 34–
<quantity id="4CD77B94F340866FA2FCF96AFC03F90F" box="[945,1005,1749,1772]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="3" pageNumber="2" unit="in" value="33.0">33 in</quantity>
male
<typeStatus id="549468D3F340866FA570F96AFB73F90F" box="[1085,1181,1749,1772]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
, and 29–29, 34–
<quantity id="4CD77B94F340866FA42EF96AFA4EF90F" box="[1379,1440,1749,1772]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="3" pageNumber="2" unit="in" value="34.0">34 in</quantity>
female
<typeStatus id="549468D3F340866FA2DFF94BFC12F8E8" box="[914,1020,1780,1803]" pageId="3" pageNumber="2" type="paratype">paratypes</typeStatus>
; middle basal lamella of the pectines not dilated in both sexes. Sternites smooth, with elongated spiracles; four carinae on sternite VII; other sternites without carinae and with two weak furrows. Metasoma: segments I to III with 10 moderately crenulated carinae, ventral more strongly marked on segments II and III; segment IV with 8 moderately crenulated carinae; segment V with five carinae; the ventrolateral carinae crenulate with 2/3 lobate denticles posteriorly; ventral median carina not divided posteriorly; anal arc composed of 9–10 ventral teeth, and two lateral lobes. All segments with a smooth dorsal depression; intercarinal smooth granular, except for the lateral and ventral aspects of segment V which presents a very thin granulation. Telson, vesicle globular with some very thin granulations on the lateral and ventral surfaces; aculeus strongly curved, and shorter than the vesicle; subaculear tooth absent. Cheliceral dentition as defined by Vachon (1963) for the family
<taxonomicName id="4C2FADF2F3468669A564FA30FB63FA45" box="[1065,1165,1423,1446]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
; external distal and internal distal denticles of approximately the same length; basal denticles of movable finger small but not fused; ventral aspect of both fingers and manus covered with long dense setae. Pedipalps: femur pentacarinate; patella with eight carinae; chela smooth with only vestigial carinae on ventral surface; all faces weakly granular to smooth. Fixed and movable fingers with 10–11 oblique rows of granules. Internal and external accessory granules present but moderate to weak; three accessory granules on the distal end of movable finger next to the terminal denticle. Legs: tarsus (telotarsus) with two longitudinal rows of 6–7 long setae ventrally; basitarsus with moderate bristle comb; it is, however, uncertain if the new species could be an exclusively psammophilic element (see habitat comments); tibial spur strong on legs III and IV; prolateral spurs moderate to strong on legs I to IV. Trichobothriotaxy: trichobothrial pattern of
<typeStatus id="549468D3F345866AA3A6FAB7FCCCFAFC" box="[747,802,1288,1311]" pageId="6" pageNumber="5">Type</typeStatus>
A, orthobothriotaxic as defined by Vachon (1974). Dorsal trichobothria of femur arranged in β- configuration (Vachon, 1975).
</paragraph>
<caption id="DF5086F9F3478668A18DF880FCEFF88C" ID-DOI="http://doi.org/10.5281/zenodo.4648182" ID-Zenodo-Dep="4648182" httpUri="https://zenodo.org/record/4648182/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,278,1855,1878]" targetBox="[275,1376,163,1807]" targetPageId="4">
<paragraph id="8B90D671F3478668A18DF880FCEFF88C" blockId="4.[192,1440,1855,1903]" pageId="4" pageNumber="3">
<emphasis id="B95B0A63F3478668A18DF880FEA1F8B5" bold="true" box="[192,335,1855,1878]" pageId="4" pageNumber="3">Figures 4–8:</emphasis>
<taxonomicName id="4C2FADF2F3478668A01AF8FFFE31F8B6" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[343,479,1856,1877]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F3478668A01AF8FFFE31F8B6" box="[343,479,1856,1877]" italics="true" pageId="4" pageNumber="3">Buthus bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F3478668A0A5F8FEFDF3F8B5" box="[488,541,1857,1878]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
Male holotype. Trichobothrial pattern.
<emphasis id="B95B0A63F3478668A2E8F8FEFC21F8B5" bold="true" box="[933,975,1857,1878]" pageId="4" pageNumber="3">4–5.</emphasis>
Chela, dorso-external and ventral aspects.
<emphasis id="B95B0A63F3478668A435F8FEFA4EF8B5" bold="true" box="[1400,1440,1857,1878]" pageId="4" pageNumber="3">6–7.</emphasis>
Patella, dorsal and external aspects.
<emphasis id="B95B0A63F3478668A351F8E5FDC0F88C" bold="true" box="[540,558,1882,1903]" pageId="4" pageNumber="3">8.</emphasis>
Femur, dorsal aspect.
</paragraph>
</caption>
<caption id="DF5086F9F3468669A18DFC39FBB7FC7E" ID-DOI="http://doi.org/10.5281/zenodo.4648184" ID-Zenodo-Dep="4648184" box="[192,1113,902,925]" httpUri="https://zenodo.org/record/4648184/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,268,902,925]" targetBox="[192,1440,76,879]" targetPageId="5">
<paragraph id="8B90D671F3468669A18DFC39FBB7FC7E" blockId="5.[192,1113,902,925]" box="[192,1113,902,925]" pageId="5" pageNumber="4">
<emphasis id="B95B0A63F3468669A18DFC39FEC6FC7E" bold="true" box="[192,296,902,925]" pageId="5" pageNumber="4">Figure 9:</emphasis>
Map of Morocco, showing the site where the new species was collected (black star).
</paragraph>
</caption>
<paragraph id="8B90D671F3468669A1BCFBA1FD9FF915" pageId="5" pageNumber="4">
<table id="F92F24D1F3467993A1BCFBA1FD9CF915" box="[241,626,1054,1782]" gridcols="2" gridrows="24" pageId="5" pageNumber="4">
<tr id="351FD433F3467993A1BCFBA1FD9CFBD6" box="[241,626,1054,1077]" gridrow="0" pageId="5" pageNumber="4">
<th id="76CEBD4FF3467993A1BCFBA1FE09FBD6" box="[241,487,1054,1077]" gridcol="0" gridrow="0" pageId="5" pageNumber="4">Total length</th>
<th id="76CEBD4FF3467993A30EFBA1FD9CFBD6" box="[579,626,1054,1077]" gridcol="1" gridrow="0" pageId="5" pageNumber="4">54.6</th>
</tr>
<tr id="351FD433F3467993A1BCFB83FD9CFBB0" box="[241,626,1084,1107]" gridrow="1" pageId="5" pageNumber="4" rowspan-1="1">
<td id="76CEBD4FF3467993A1BCFB83FE09FBB0" box="[241,487,1084,1107]" gridcol="0" gridrow="1" pageId="5" pageNumber="4">Carapace:</td>
</tr>
<tr id="351FD433F3467993A1BCFBE4FD9CFB91" box="[241,626,1115,1138]" gridrow="2" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFBE4FE09FB91" box="[241,487,1115,1138]" gridcol="0" gridrow="2" pageId="5" pageNumber="4">- length</td>
<td id="76CEBD4FF3467993A30EFBE4FD9CFB91" box="[579,626,1115,1138]" gridcol="1" gridrow="2" pageId="5" pageNumber="4">6.3</td>
</tr>
<tr id="351FD433F3467993A1BCFBC5FD9CFB72" box="[241,626,1146,1169]" gridrow="3" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFBC5FE09FB72" box="[241,487,1146,1169]" gridcol="0" gridrow="3" pageId="5" pageNumber="4">- anterior width</td>
<td id="76CEBD4FF3467993A30EFBC5FD9CFB72" box="[579,626,1146,1169]" gridcol="1" gridrow="3" pageId="5" pageNumber="4">4.5</td>
</tr>
<tr id="351FD433F3467993A1BCFB27FD9CFB4C" box="[241,626,1176,1199]" gridrow="4" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFB27FE09FB4C" box="[241,487,1176,1199]" gridcol="0" gridrow="4" pageId="5" pageNumber="4">- posterior width</td>
<td id="76CEBD4FF3467993A30EFB27FD9CFB4C" box="[579,626,1176,1199]" gridcol="1" gridrow="4" pageId="5" pageNumber="4">7.2</td>
</tr>
<tr id="351FD433F3467993A1BCFB08FD9CFB2D" box="[241,626,1207,1230]" gridrow="5" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFB08FD9CFB2D" box="[241,626,1207,1230]" colspan="2" colspanRight="1" gridcol="0" gridrow="5" pageId="5" pageNumber="4">Metasomal segment I:</td>
</tr>
<tr id="351FD433F3467993A1BCFB69FD9CFB0E" box="[241,626,1238,1261]" gridrow="6" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFB69FE09FB0E" box="[241,487,1238,1261]" gridcol="0" gridrow="6" pageId="5" pageNumber="4">- length</td>
<td id="76CEBD4FF3467993A30EFB69FD9CFB0E" box="[579,626,1238,1261]" gridcol="1" gridrow="6" pageId="5" pageNumber="4">4.5</td>
</tr>
<tr id="351FD433F3467993A1BCFB4BFD9CFAE8" box="[241,626,1268,1291]" gridrow="7" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFB4BFE09FAE8" box="[241,487,1268,1291]" gridcol="0" gridrow="7" pageId="5" pageNumber="4">- width</td>
<td id="76CEBD4FF3467993A30EFB4BFD9CFAE8" box="[579,626,1268,1291]" gridcol="1" gridrow="7" pageId="5" pageNumber="4">4.2</td>
</tr>
<tr id="351FD433F3467993A1BCFAACFD9CFAC9" box="[241,626,1299,1322]" gridrow="8" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFAACFD9CFAC9" box="[241,626,1299,1322]" colspan="2" colspanRight="1" gridcol="0" gridrow="8" pageId="5" pageNumber="4">Metasomal segment V:</td>
</tr>
<tr id="351FD433F3467993A1BCFA8DFD9CFAAA" box="[241,626,1330,1353]" gridrow="9" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFA8DFE09FAAA" box="[241,487,1330,1353]" gridcol="0" gridrow="9" pageId="5" pageNumber="4">- length</td>
<td id="76CEBD4FF3467993A30EFA8DFD9CFAAA" box="[579,626,1330,1353]" gridcol="1" gridrow="9" pageId="5" pageNumber="4">7.7</td>
</tr>
<tr id="351FD433F3467993A1BCFAEFFD9CFA84" box="[241,626,1360,1383]" gridrow="10" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFAEFFE09FA84" box="[241,487,1360,1383]" gridcol="0" gridrow="10" pageId="5" pageNumber="4">- width</td>
<td id="76CEBD4FF3467993A30EFAEFFD9CFA84" box="[579,626,1360,1383]" gridcol="1" gridrow="10" pageId="5" pageNumber="4">3.1</td>
</tr>
<tr id="351FD433F3467993A1BCFAD0FD9CFA65" box="[241,626,1391,1414]" gridrow="11" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFAD0FE09FA65" box="[241,487,1391,1414]" gridcol="0" gridrow="11" pageId="5" pageNumber="4">- depth</td>
<td id="76CEBD4FF3467993A30EFAD0FD9CFA65" box="[579,626,1391,1414]" gridcol="1" gridrow="11" pageId="5" pageNumber="4">2.6</td>
</tr>
<tr id="351FD433F3467993A1BCFA31FD9CFA46" box="[241,626,1422,1445]" gridrow="12" pageId="5" pageNumber="4" rowspan-1="1">
<td id="76CEBD4FF3467993A1BCFA31FE09FA46" box="[241,487,1422,1445]" gridcol="0" gridrow="12" pageId="5" pageNumber="4">Vesicle:</td>
</tr>
<tr id="351FD433F3467993A1BCFA13FD9CFA20" box="[241,626,1452,1475]" gridrow="13" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFA13FE09FA20" box="[241,487,1452,1475]" gridcol="0" gridrow="13" pageId="5" pageNumber="4">- width</td>
<td id="76CEBD4FF3467993A30EFA13FD9CFA20" box="[579,626,1452,1475]" gridcol="1" gridrow="13" pageId="5" pageNumber="4">2.8</td>
</tr>
<tr id="351FD433F3467993A1BCFA74FD9CFA01" box="[241,626,1483,1506]" gridrow="14" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCFA74FE09FA01" box="[241,487,1483,1506]" gridcol="0" gridrow="14" pageId="5" pageNumber="4">- depth</td>
<td id="76CEBD4FF3467993A30EFA74FD9CFA01" box="[579,626,1483,1506]" gridcol="1" gridrow="14" pageId="5" pageNumber="4">2.8</td>
</tr>
<tr id="351FD433F3467993A1BCFA55FD9CF9E2" box="[241,626,1514,1537]" gridrow="15" pageId="5" pageNumber="4" rowspan-1="1">
<td id="76CEBD4FF3467993A1BCFA55FE09F9E2" box="[241,487,1514,1537]" gridcol="0" gridrow="15" pageId="5" pageNumber="4">Pedipalp:</td>
</tr>
<tr id="351FD433F3467993A1BCF9B7FD9CF9FC" box="[241,626,1544,1567]" gridrow="16" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCF9B7FE09F9FC" box="[241,487,1544,1567]" gridcol="0" gridrow="16" pageId="5" pageNumber="4">- Femur length</td>
<td id="76CEBD4FF3467993A30EF9B7FD9CF9FC" box="[579,626,1544,1567]" gridcol="1" gridrow="16" pageId="5" pageNumber="4">5.3</td>
</tr>
<tr id="351FD433F3467993A1BCF998FD9CF9DD" box="[241,626,1575,1598]" gridrow="17" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCF998FE09F9DD" box="[241,487,1575,1598]" gridcol="0" gridrow="17" pageId="5" pageNumber="4">- Femur width</td>
<td id="76CEBD4FF3467993A30EF998FD9CF9DD" box="[579,626,1575,1598]" gridcol="1" gridrow="17" pageId="5" pageNumber="4">1.7</td>
</tr>
<tr id="351FD433F3467993A1BCF9F9FD9CF9BE" box="[241,626,1606,1629]" gridrow="18" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCF9F9FE09F9BE" box="[241,487,1606,1629]" gridcol="0" gridrow="18" pageId="5" pageNumber="4">- Patella length</td>
<td id="76CEBD4FF3467993A30EF9F9FD9CF9BE" box="[579,626,1606,1629]" gridcol="1" gridrow="18" pageId="5" pageNumber="4">6.0</td>
</tr>
<tr id="351FD433F3467993A1BCF9DBFD9CF998" box="[241,626,1636,1659]" gridrow="19" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCF9DBFE09F998" box="[241,487,1636,1659]" gridcol="0" gridrow="19" pageId="5" pageNumber="4">- Patella width</td>
<td id="76CEBD4FF3467993A30EF9DBFD9CF998" box="[579,626,1636,1659]" gridcol="1" gridrow="19" pageId="5" pageNumber="4">2.3</td>
</tr>
<tr id="351FD433F3467993A1BCF93CFD9CF979" box="[241,626,1667,1690]" gridrow="20" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCF93CFE09F979" box="[241,487,1667,1690]" gridcol="0" gridrow="20" pageId="5" pageNumber="4">- Chela length</td>
<td id="76CEBD4FF3467993A30EF93CFD9CF979" box="[579,626,1667,1690]" gridcol="1" gridrow="20" pageId="5" pageNumber="4">10.5</td>
</tr>
<tr id="351FD433F3467993A1BCF91DFD9CF95A" box="[241,626,1698,1721]" gridrow="21" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCF91DFE09F95A" box="[241,487,1698,1721]" gridcol="0" gridrow="21" pageId="5" pageNumber="4">- Chela width</td>
<td id="76CEBD4FF3467993A30EF91DFD9CF95A" box="[579,626,1698,1721]" gridcol="1" gridrow="21" pageId="5" pageNumber="4">2.4</td>
</tr>
<tr id="351FD433F3467993A1BCF97FFD9CF934" box="[241,626,1728,1751]" gridrow="22" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCF97FFE09F934" box="[241,487,1728,1751]" gridcol="0" gridrow="22" pageId="5" pageNumber="4">- Chela depth</td>
<td id="76CEBD4FF3467993A30EF97FFD9CF934" box="[579,626,1728,1751]" gridcol="1" gridrow="22" pageId="5" pageNumber="4">2.4</td>
</tr>
<tr id="351FD433F3467993A1BCF960FD9CF915" box="[241,626,1759,1782]" gridrow="23" pageId="5" pageNumber="4">
<td id="76CEBD4FF3467993A1BCF960FE09F915" box="[241,487,1759,1782]" gridcol="0" gridrow="23" pageId="5" pageNumber="4">Movable finger: length</td>
<td id="76CEBD4FF3467993A30EF960FD9CF915" box="[579,626,1759,1782]" gridcol="1" gridrow="23" pageId="5" pageNumber="4">6.8</td>
</tr>
</table>
</paragraph>
<caption id="DF5086F9F3468669A18DF8FEFE74F893" ID-Table-UUID="DF5086F9F3468669A18DF8FEFE74F893" httpUri="http://table.plazi.org/id/DF5086F9F3468669A18DF8FEFE74F893" pageId="5" pageNumber="4" startId="5.[192,257,1857,1880]" targetBox="[241,626,1054,1782]" targetIsTable="true" targetPageId="5">
<paragraph id="8B90D671F3468669A18DF8FEFE74F893" blockId="5.[192,801,1857,1904]" pageId="5" pageNumber="4">
<emphasis id="B95B0A63F3468669A18DF8FEFECEF8BB" bold="true" box="[192,288,1857,1880]" pageId="5" pageNumber="4">Table 1:</emphasis>
Morphometric values (in mm) of the male holotype of
<taxonomicName id="4C2FADF2F3468669A197F8E4FE8EF893" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[218,352,1883,1904]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F3468669A197F8E4FE8EF893" box="[218,352,1883,1904]" italics="true" pageId="5" pageNumber="4">Buthus bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F3468669A02BF8E4FE74F893" box="[358,410,1883,1904]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF5086F9F345866AA18DFB2BFEADFB20" ID-DOI="http://doi.org/10.5281/zenodo.4648186" ID-Zenodo-Dep="4648186" httpUri="https://zenodo.org/record/4648186/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,268,1172,1195]" targetBox="[189,1443,189,1131]" targetPageId="6">
<paragraph id="8B90D671F345866AA18DFB2BFEADFB20" blockId="6.[192,1440,1172,1219]" pageId="6" pageNumber="5">
<emphasis id="B95B0A63F345866AA18DFB2BFED9FB48" bold="true" box="[192,311,1172,1195]" pageId="6" pageNumber="5">Figure 10:</emphasis>
Juvenile specimen of
<taxonomicName id="4C2FADF2F345866AA35AFB2AFD71FB49" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[535,671,1173,1194]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F345866AA35AFB2AFD71FB49" box="[535,671,1173,1194]" italics="true" pageId="6" pageNumber="5">Buthus bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F345866AA3E8FB2AFD34FB49" box="[677,730,1173,1194]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
just after molting process in its natural habitat. Note: this specimen was not collected.
</paragraph>
</caption>
<paragraph id="8B90D671F345866AA18DFA20FEB4FA55" blockId="6.[192,804,1439,1891]" box="[192,346,1439,1462]" pageId="6" pageNumber="5">
<emphasis id="B95B0A63F345866AA18DFA20FEB4FA55" bold="true" box="[192,346,1439,1462]" pageId="6" pageNumber="5">Relationships</emphasis>
</paragraph>
<paragraph id="8B90D671F345866AA18DFA03FC39FA7A" blockId="6.[192,804,1439,1891]" lastBlockId="6.[830,1440,1256,1433]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA18DFA03FEB2FA37" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[192,348,1468,1492]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F345866AA18DFA03FEB2FA37" box="[192,348,1468,1492]" italics="true" pageId="6" pageNumber="5">Buthus bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F345866AA024FA02FE44FA37" box="[361,426,1469,1492]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
does not belong to the “
<taxonomicName id="4C2FADF2F345866AA395FA03FEC8FA10" baseAuthorityName="Amoreux" baseAuthorityYear="1889" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="occitanus">
<emphasis id="B95B0A63F345866AA395FA03FEC8FA10" italics="true" pageId="6" pageNumber="5">Buthus occitanus</emphasis>
</taxonomicName>
complex of species”, but is rather associated with
<taxonomicName id="4C2FADF2F345866AA1B3FA46FDACF9F2" authority="Pocock. The" authorityName="Pocock. The" authorityYear="1889" box="[254,578,1529,1553]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F345866AA1B3FA46FE48F9F2" box="[254,422,1529,1553]" italics="true" pageId="6" pageNumber="5">Buthus atlantis</emphasis>
Pocock. The
</taxonomicName>
definition of these groups is based on carinae development.
<taxonomicName id="4C2FADF2F345866AA3D5F9A6FCCFF9D3" baseAuthorityName="Amoreux" baseAuthorityYear="1889" box="[664,801,1561,1584]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="occitanus">
<emphasis id="B95B0A63F345866AA3D5F9A6FCCFF9D3" box="[664,801,1561,1584]" italics="true" pageId="6" pageNumber="5">B. occitanus</emphasis>
</taxonomicName>
related species show very strongly developed carinae, whereas, those associated with
<taxonomicName id="4C2FADF2F345866AA35DF9E9FD90F98E" authorityName="Pocock. The" authorityYear="1889" box="[528,638,1621,1645]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F345866AA35DF9E9FD90F98E" box="[528,638,1621,1645]" italics="true" pageId="6" pageNumber="5">B. atlantis</emphasis>
</taxonomicName>
have moderate to weak carinae. The new species can, however, be distinguished from
<taxonomicName id="4C2FADF2F345866AA03FF92CFDFDF948" authorityName="Pocock. The" authorityYear="1889" box="[370,531,1683,1707]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F345866AA03FF92CFDFDF948" box="[370,531,1683,1707]" italics="true" pageId="6" pageNumber="5">Buthus atlantis</emphasis>
</taxonomicName>
by the following characters: (i) in
<taxonomicName id="4C2FADF2F345866AA07AF90DFE75F92A" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[311,411,1713,1737]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F345866AA07AF90DFE75F92A" box="[311,411,1713,1737]" italics="true" pageId="6" pageNumber="5">B. bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F345866AA0E9F90DFE0EF92A" box="[420,480,1714,1737]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
lateral carinae of tergites are very much reduced or absent; (ii) aculeus in
<taxonomicName id="4C2FADF2F345866AA3F1F96EFCCFF90B" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[700,801,1744,1768]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F345866AA3F1F96EFCCFF90B" box="[700,801,1744,1768]" italics="true" pageId="6" pageNumber="5">B. bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F345866AA18DF94FFF11F8E4" box="[192,255,1776,1799]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
is shorter than the vesicle and strongly curved, whereas in
<taxonomicName id="4C2FADF2F345866AA00FF8B1FE5DF8C6" authorityName="Pocock. The" authorityYear="1889" box="[322,435,1805,1829]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F345866AA00FF8B1FE5DF8C6" box="[322,435,1805,1829]" italics="true" pageId="6" pageNumber="5">B. atlantis</emphasis>
</taxonomicName>
it is longer than the vesicle and weakly curved (see Lourenço, 2003, Figs. 20 and 24); (iii) anal arc in the new species shows two lobes, whereas in
<taxonomicName id="4C2FADF2F345866AA28BFB56FBD5FAE3" authorityName="Pocock. The" authorityYear="1889" box="[966,1083,1256,1280]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F345866AA28BFB56FBD5FAE3" box="[966,1083,1256,1280]" italics="true" pageId="6" pageNumber="5">B. atlantis</emphasis>
</taxonomicName>
three lobes are frequently observed; (iv) adults of
<taxonomicName id="4C2FADF2F345866AA567FAB8FB25FAFC" authorityName="Pocock. The" authorityYear="1889" box="[1066,1227,1287,1311]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F345866AA567FAB8FB25FAFC" box="[1066,1227,1287,1311]" italics="true" pageId="6" pageNumber="5">Buthus atlantis</emphasis>
</taxonomicName>
are much bigger in size with
<quantity id="4CD77B94F345866AA2E7FA99FBD7FADE" box="[938,1081,1318,1341]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="6" pageNumber="5" unit="mm" value="85.0" valueMax="90.0" valueMin="80.0">80 to 90 mm</quantity>
in total length; (v) juveniles exhibit a contrasted blackish and yellowish pattern while those of
<taxonomicName id="4C2FADF2F345866AA2D2FADBFBE1FA98" authorityName="Pocock. The" authorityYear="1889" box="[927,1039,1379,1403]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F345866AA2D2FADBFBE1FA98" box="[927,1039,1379,1403]" italics="true" pageId="6" pageNumber="5">B. atlantis</emphasis>
</taxonomicName>
present a relatively uniform yellowish coloration.
</paragraph>
<paragraph id="8B90D671F345866AA273FA04FC4DFA12" blockId="6.[830,1434,1466,1889]" pageId="6" pageNumber="5">
<emphasis id="B95B0A63F345866AA273FA04FC4DFA12" bold="true" pageId="6" pageNumber="5">
Check-list of the known
<taxonomicName id="4C2FADF2F345866AA519FA05FB4AFA31" authorityName="Leach" authorityYear="1815" box="[1108,1188,1466,1490]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B95B0A63F345866AA519FA05FB4AFA31" bold="true" box="[1108,1188,1466,1490]" italics="true" pageId="6" pageNumber="5">Buthus</emphasis>
</taxonomicName>
species distributed in
<collectingCountry id="F33896E1F345866AA273FA65FC4DFA12" box="[830,931,1498,1521]" name="Morocco" pageId="6" pageNumber="5">Morocco</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="8B90D671F345866AA273FA47FB5BF9EC" blockId="6.[830,1434,1466,1889]" box="[830,1205,1528,1552]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273FA47FB5BF9EC" authority="(Amoreux, 1889)" baseAuthorityName="Amoreux" baseAuthorityYear="1889" box="[830,1205,1528,1552]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="occitanus">
<emphasis id="B95B0A63F345866AA273FA47FC1BF9EC" box="[830,1013,1528,1552]" italics="true" pageId="6" pageNumber="5">Buthus occitanus</emphasis>
(Amoreux, 1889)
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F9A9FB7FF9CD" blockId="6.[830,1434,1466,1889]" box="[830,1169,1558,1582]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F9A9FB7FF9CD" authority="(C. L. Koch, 1839)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1839" box="[830,1169,1558,1582]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="paris">
<emphasis id="B95B0A63F345866AA273F9A9FC28F9CD" box="[830,966,1558,1582]" italics="true" pageId="6" pageNumber="5">Buthus paris</emphasis>
(C.L. Koch, 1839)
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F98AFB7EF9AE" blockId="6.[830,1434,1466,1889]" box="[830,1168,1589,1613]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F98AFB7EF9AE" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" box="[830,1168,1589,1613]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="mardochei">
<emphasis id="B95B0A63F345866AA273F98AFBEFF9AE" box="[830,1025,1589,1613]" italics="true" pageId="6" pageNumber="5">Buthus mardochei</emphasis>
Simon, 1878
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F9EBFB99F988" blockId="6.[830,1434,1466,1889]" box="[830,1143,1620,1644]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F9EBFB99F988" authority="Pocock, 1889" authorityName="Pocock. The" authorityYear="1889" box="[830,1143,1620,1644]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B95B0A63F345866AA273F9EBFC31F98F" box="[830,991,1620,1644]" italics="true" pageId="6" pageNumber="5">Buthus atlantis</emphasis>
Pocock, 1889
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F9CDFB73F969" blockId="6.[830,1434,1466,1889]" box="[830,1181,1650,1674]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F9CDFB73F969" authority="Birula, 1903" authorityName="Birula" authorityYear="1903" box="[830,1181,1650,1674]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="B95B0A63F345866AA273F9CDFBFFF969" box="[830,1041,1650,1674]" italics="true" pageId="6" pageNumber="5">Buthus maroccanus</emphasis>
Birula, 1903
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F92EFB49F94A" blockId="6.[830,1434,1466,1889]" box="[830,1191,1681,1705]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F92EFB49F94A" authority="Vachon, 1949" authorityName="Vachon" authorityYear="1949" box="[830,1191,1681,1705]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="malhommei">
<emphasis id="B95B0A63F345866AA273F92EFBE5F94A" box="[830,1035,1681,1705]" italics="true" pageId="6" pageNumber="5">Buthus malhommei</emphasis>
Vachon, 1949
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F90FFB34F924" blockId="6.[830,1434,1466,1889]" box="[830,1242,1712,1736]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F90FFB34F924" authority="Lourenco, 2003" authorityName="Lourenco" authorityYear="2003" box="[830,1242,1712,1736]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="mariefranceae">
<emphasis id="B95B0A63F345866AA273F90FFBC5F92B" box="[830,1067,1712,1736]" italics="true" pageId="6" pageNumber="5">Buthus mariefranceae</emphasis>
Lourenço, 2003
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F971FB62F905" blockId="6.[830,1434,1466,1889]" box="[830,1164,1742,1766]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F971FB62F905" authority="Lourenco, 2003" authorityName="Lourenco" authorityYear="2003" box="[830,1164,1742,1766]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="rochati">
<emphasis id="B95B0A63F345866AA273F971FC32F905" box="[830,988,1742,1766]" italics="true" pageId="6" pageNumber="5">Buthus rochati</emphasis>
Lourenço, 2003
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F952FB4EF8E6" blockId="6.[830,1434,1466,1889]" box="[830,1184,1773,1797]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F952FB4EF8E6" authority="Lourenco, 2003" authorityName="Lourenco" authorityYear="2003" box="[830,1184,1773,1797]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="lienhardi">
<emphasis id="B95B0A63F345866AA273F952FC1FF8E6" box="[830,1009,1773,1797]" italics="true" pageId="6" pageNumber="5">Buthus lienhardi</emphasis>
Lourenço, 2003
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F8B3FB72F8C0" blockId="6.[830,1434,1466,1889]" box="[830,1180,1804,1828]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F8B3FB72F8C0" authority="Lourenco, 2003" authorityName="Lourenco" authorityYear="2003" box="[830,1180,1804,1828]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="albengai">
<emphasis id="B95B0A63F345866AA273F8B3FC02F8C7" box="[830,1004,1804,1828]" italics="true" pageId="6" pageNumber="5">Buthus albengai</emphasis>
Lourenço, 2003
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F895FB08F8A1" blockId="6.[830,1434,1466,1889]" box="[830,1254,1834,1858]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F895FB08F8A1" authority="Lourenco &amp; Slimani, 2004" authorityName="Lourenco &amp; Slimani" authorityYear="2004" box="[830,1254,1834,1858]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="draa">
<emphasis id="B95B0A63F345866AA273F895FC2FF8A1" box="[830,961,1834,1858]" italics="true" pageId="6" pageNumber="5">Buthus draa</emphasis>
Lourenço &amp; Slimani, 2004
</taxonomicName>
</paragraph>
<paragraph id="8B90D671F345866AA273F8F6FBFDF882" blockId="6.[830,1434,1466,1889]" box="[830,1043,1865,1889]" pageId="6" pageNumber="5">
<taxonomicName id="4C2FADF2F345866AA273F8F6FC3DF882" authority="Lourenço &amp; Geniez, 2005" authorityName="Lourenço &amp; Geniez" authorityYear="2005" box="[830,979,1865,1889]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="bonito" status="sp. nov.">
<emphasis id="B95B0A63F345866AA273F8F6FC3DF882" box="[830,979,1865,1889]" italics="true" pageId="6" pageNumber="5">Buthus bonito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268B718F345866AA297F8F5FBFDF882" box="[986,1043,1866,1889]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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