223 lines
22 KiB
XML
223 lines
22 KiB
XML
<document id="B74CB3C8CF37B793D8D480405EA51C55" ID-DOI="10.11646/zootaxa.3999.3.7" ID-GBIF-Dataset="9d12960e-b90a-4df7-b7a7-25c05e587130" ID-ISSN="1175-5326" ID-Zenodo-Dep="242627" ID-ZooBank="DDB20916-F3A8-42CF-A1C7-BA0E99C1BEB6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461282335071" checkinUser="plazi" docAuthor="Peng, Zhong, Li, Qi-Li, Li, Li-Zhen & Gu, Fu-Kang" docDate="2015" docId="0386383E47054A71FF2A1578FC384DF8" docLanguage="en" docName="zt03999p429.pdf" docOrigin="Zootaxa 3999 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paederus (Gnathopaederus) szechuanus Chapin 1927" docType="treatment" docVersion="8" lastPageNumber="425" masterDocId="FFBF404647064A75FFBD1434FFE34F17" masterDocTitle="New data on the subgenus Gnathopaederus of the genus Paederus (Coleoptera: Staphylinidae: Paederinae) of mainland China" masterLastPageNumber="429" masterPageNumber="421" pageNumber="424" updateTime="1698614385559" updateUser="plazi">
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<mods:titleInfo id="FF5FD101470984F9FCEC3846768035BC">
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<mods:title id="2AE546CA03609A070FB0853B5CB70728">New data on the subgenus Gnathopaederus of the genus Paederus (Coleoptera: Staphylinidae: Paederinae) of mainland China</mods:title>
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<mods:name id="375A876DF295EB7A500B750223B8C450" type="personal">
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<mods:namePart id="95A1A7CC1D4111CA37247C1AB393464A">Peng, Zhong</mods:namePart>
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<mods:namePart id="E28AB3CEAA49A74CE9B419F81C1CA9B9">Li, Qi-Li</mods:namePart>
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<mods:namePart id="35E82F0BBB4E6E8AFFBEF7276C9752B8">Li, Li-Zhen</mods:namePart>
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<mods:namePart id="B42297094537CB80F533EF1FC078ED33">Gu, Fu-Kang</mods:namePart>
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<mods:title id="62EDBD741F34CCDC735F4781E03FC38C">Zootaxa</mods:title>
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<mods:date id="C0217A8E7592361E34A81F8C8D964BD6">2015</mods:date>
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<mods:number id="E7757C06A5F9C7AE87B480179E522372">3999</mods:number>
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<mods:identifier id="ABD3A480C647286F6810793FA445F58E" type="DOI">10.11646/zootaxa.3999.3.7</mods:identifier>
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<treatment id="0386383E47054A71FF2A1578FC384DF8" ID-DOI="http://doi.org/10.5281/zenodo.6097736" ID-GBIF-Taxon="119640858" ID-Zenodo-Dep="6097736" LSID="urn:lsid:plazi:treatment:0386383E47054A71FF2A1578FC384DF8" httpUri="http://treatment.plazi.org/id/0386383E47054A71FF2A1578FC384DF8" lastPageId="4" lastPageNumber="425" pageId="3" pageNumber="424">
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<subSubSection id="C335DAA347054A76FF2A1578FE8D4E90" pageId="3" pageNumber="424" type="nomenclature">
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<paragraph id="8B90892847054A76FF2A1578FCDD4E71" blockId="3.[151,830,332,391]" box="[151,830,332,358]" pageId="3" pageNumber="424">
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<heading id="D0D83E4447054A76FF2A1578FCDD4E71" bold="true" box="[151,830,332,358]" fontSize="11" level="1" pageId="3" pageNumber="424" reason="1">
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<taxonomicName id="4C2FF2AB47054A76FF2A1578FCDD4E71" ID-CoL="4BX6G" authority="Chapin, 1927" authorityName="Chapin" authorityYear="1927" box="[151,830,332,358]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="szechuanus" subGenus="Gnathopaederus">
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<emphasis id="B95B553A47054A76FF2A1578FCDD4E71" bold="true" box="[151,830,332,358]" pageId="3" pageNumber="424">
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<emphasis id="B95B553A47054A76FF2A1578FEEB4E71" bold="true" box="[151,264,332,358]" italics="true" pageId="3" pageNumber="424">Paederus</emphasis>
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(
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<emphasis id="B95B553A47054A76FEA51578FE034E71" bold="true" box="[280,480,332,358]" italics="true" pageId="3" pageNumber="424">Gnathopaederus</emphasis>
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)
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<emphasis id="B95B553A47054A76FE4F1578FD9D4E71" bold="true" box="[498,638,332,358]" italics="true" pageId="3" pageNumber="424">szechuanus</emphasis>
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(Chapin, 1927)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
<paragraph id="8B90892847054A76FF2A155AFE8D4E90" blockId="3.[151,830,332,391]" box="[151,366,366,391]" pageId="3" pageNumber="424">(Figs 1B–C, 1H, 3)</paragraph>
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||
</subSubSection>
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||
<caption id="DF50D9A047054A76FF2A130CFDEA48BD" httpUri="https://zenodo.org/record/242629/files/figure.png" pageId="3" pageNumber="424" targetBox="[166,1435,455,1817]" targetPageId="3">
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<paragraph id="8B90892847054A76FF2A130CFDEA48BD" blockId="3.[151,1437,1848,1962]" pageId="3" pageNumber="424">
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<emphasis id="B95B553A47054A76FF2A130CFEF6485A" bold="true" box="[151,277,1848,1870]" pageId="3" pageNumber="424">FIGURE 3.</emphasis>
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<taxonomicName id="4C2FF2AB47054A76FEA0130CFE1B485A" box="[285,504,1848,1869]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="szechuanus">
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<emphasis id="B95B553A47054A76FEA0130CFE1B485A" box="[285,504,1848,1869]" italics="true" pageId="3" pageNumber="424">Paederus szechuanus</emphasis>
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</taxonomicName>
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.
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<emphasis id="B95B553A47054A76FDBA130CFDFA485A" bold="true" box="[519,537,1848,1869]" pageId="3" pageNumber="424">A</emphasis>
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male left mandible
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<emphasis id="B95B553A47054A76FD50130CFD1D485A" bold="true" box="[749,766,1848,1869]" pageId="3" pageNumber="424">B</emphasis>
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male right mandible
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<emphasis id="B95B553A47054A76FC62130CFC124859" bold="true" box="[991,1009,1848,1870]" pageId="3" pageNumber="424">C</emphasis>
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female right mandible
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<emphasis id="B95B553A47054A76FB5B130CFB1B485A" bold="true" box="[1254,1272,1848,1869]" pageId="3" pageNumber="424">D</emphasis>
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||
male labrum
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<emphasis id="B95B553A47054A76FA36130CFA7F485A" bold="true" box="[1419,1436,1848,1869]" pageId="3" pageNumber="424">E</emphasis>
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female labrum
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<emphasis id="B95B553A47054A76FE921363FEDD487B" bold="true" box="[303,318,1879,1900]" pageId="3" pageNumber="424">F</emphasis>
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female tergite VIII
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<emphasis id="B95B553A47054A76FDB51363FDFF487A" bold="true" box="[520,540,1879,1901]" pageId="3" pageNumber="424">G</emphasis>
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female sternite VIII
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<emphasis id="B95B553A47054A76FD521363FCE0487B" bold="true" box="[751,771,1879,1900]" pageId="3" pageNumber="424">H</emphasis>
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female sternite IX
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<emphasis id="B95B553A47054A76FC781363FC2C487B" bold="true" box="[965,975,1879,1900]" pageId="3" pageNumber="424">I</emphasis>
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male tergite VIII
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<emphasis id="B95B553A47054A76FB391363FB72487B" bold="true" box="[1156,1169,1879,1900]" pageId="3" pageNumber="424">J</emphasis>
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||
male sternite VIII
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||
<emphasis id="B95B553A47054A76FAEC1363FA86487B" bold="true" box="[1361,1381,1879,1900]" pageId="3" pageNumber="424">K</emphasis>
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male tergite IX
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<emphasis id="B95B553A47054A76FEBD1342FED1489C" bold="true" box="[256,306,1909,1931]" pageId="3" pageNumber="424">L, O</emphasis>
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aedeagus in ventral view
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<emphasis id="B95B553A47054A76FD801342FDB6489C" bold="true" box="[573,597,1910,1931]" pageId="3" pageNumber="424">M</emphasis>
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aedeagus in lateral view
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<emphasis id="B95B553A47054A76FCE41342FC88489C" bold="true" box="[857,875,1910,1931]" pageId="3" pageNumber="424">N</emphasis>
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||
aedeagus in dorsal view
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||
<emphasis id="B95B553A47054A76FBD31342FB9E489C" bold="true" box="[1134,1149,1910,1931]" pageId="3" pageNumber="424">P</emphasis>
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||
the front part of parameres. Scale bars:
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||
<emphasis id="B95B553A47054A76FEB113A0FEDD48BD" bold="true" box="[268,318,1940,1962]" pageId="3" pageNumber="424">A–O</emphasis>
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||
0.5 mm,
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||
<emphasis id="B95B553A47054A76FE2313A1FE4E48BD" bold="true" box="[414,429,1941,1962]" pageId="3" pageNumber="424">P</emphasis>
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||
0.1 mm.
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||
</paragraph>
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||
</caption>
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||
<subSubSection id="C335DAA347024A71FF2A14A3FDC24EC7" pageId="4" pageNumber="425" type="materials_examined">
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<emphasis id="B95B553A47024A71FF2A14A3FE874FA7" bold="true" box="[151,356,151,176]" pageId="4" pageNumber="425">Material studied.</emphasis>
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<collectingCountry id="F338C9B847024A71FED614A3FE514FA7" box="[363,434,151,176]" name="China" pageId="4" pageNumber="425">China</collectingCountry>
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: Sichuan: 2 ♂♂, 5 ♀♀, Dujiangyan, Qingcheng Shan,
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,
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,
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<date id="FF91AFE847024A71FF2A1488FEC14FC3" box="[151,290,188,212]" pageId="4" pageNumber="425" value="2012-07-30">30.VII.2012</date>
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, Zhong Peng leg. (
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<collectionCode id="ED3E11ED47024A71FDBF1488FDB04FC3" box="[514,595,188,212]" pageId="4" pageNumber="425">SNUC</collectionCode>
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); 2 ♂♂, 4 ♀♀, Dujiangyan, Qingcheng Shan,
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<geoCoordinate id="EE1BEFEF47024A71FBC91488FB1A4FC2" box="[1140,1273,188,213]" direction="north" orientation="latitude" pageId="4" pageNumber="425" precision="15" value="30.949444">30°56'58''N</geoCoordinate>
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,
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,
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<quantity id="4CD724CD47024A71FF2A14EBFF0C4FE0" box="[151,239,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="4" pageNumber="425" unit="m" value="1650.0">1650 m</quantity>
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,
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<date id="FF91AFE847024A71FF4114D4FE644FEF" box="[252,391,224,248]" pageId="4" pageNumber="425" value="2012-07-29">29.VII.2012</date>
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, Dai, Peng & Yin leg. (
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<collectionCode id="ED3E11ED47024A71FD2E14D4FD074FEF" box="[659,740,224,248]" pageId="4" pageNumber="425">SNUC</collectionCode>
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); 3 ♂♂, Baoxing Hsien, Fengtongzhai Natural Reserve,
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<quantity id="4CD724CD47024A71FAD914D4FF4C4E0B" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="425" unit="m" value="1900.0">1900 m</quantity>
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,
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<date id="FF91AFE847024A71FF7D1530FEDA4E0B" box="[192,313,260,284]" pageId="4" pageNumber="425" value="2014-10-02">02.X.2014</date>
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, Hao Xu & Jian-Yue Qiu leg. (
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<collectionCode id="ED3E11ED47024A71FD0B1530FCE44E0B" box="[694,775,260,284]" pageId="4" pageNumber="425">SNUC</collectionCode>
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); 2 ♀♀, Baoxing Hsien, Fengtongzhai Natural Reserve,
|
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<geoCoordinate id="EE1BEFEF47024A71FF2A151CFF164E57" box="[151,245,296,320]" direction="north" orientation="latitude" pageId="4" pageNumber="425" precision="925" value="30.533333">30°32'N</geoCoordinate>
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,
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<geoCoordinate id="EE1BEFEF47024A71FEBA1513FE924E57" box="[263,369,295,320]" direction="east" orientation="longitude" pageId="4" pageNumber="425" precision="925" value="102.933334">102°56'E</geoCoordinate>
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,
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<quantity id="4CD724CD47024A71FE3F1513FE3C4E28" box="[386,479,295,320]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="425" unit="m" value="1500.0">1500 m</quantity>
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,
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<date id="FF91AFE847024A71FE4C1513FD904E57" box="[497,627,295,320]" pageId="4" pageNumber="425" value="2012-06-25">25.VI.2012</date>
|
||
, Huang leg. (
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<collectionCode id="ED3E11ED47024A71FCA7151CFC884E57" box="[794,875,296,320]" pageId="4" pageNumber="425">SNUC</collectionCode>
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); 2 ♂♂, 2 ♀♀, Tianquan Hsien, Liangluxiang,
|
||
<geoCoordinate id="EE1BEFEF47024A71FF2A1578FEFF4E72" box="[151,284,332,357]" direction="north" orientation="latitude" pageId="4" pageNumber="425" precision="15" value="29.93861">29°56'19''N</geoCoordinate>
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,
|
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<geoCoordinate id="EE1BEFEF47024A71FE951578FE5B4E73" box="[296,440,332,356]" direction="east" orientation="longitude" pageId="4" pageNumber="425" precision="15" value="102.391106">102°23'28''E</geoCoordinate>
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,
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<quantity id="4CD724CD47024A71FE791578FDF84E73" box="[452,539,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="425" unit="m" value="1500.0">1500 m</quantity>
|
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,
|
||
<date id="FF91AFE847024A71FD9A1578FD504E73" box="[551,691,332,356]" pageId="4" pageNumber="425" value="2012-07-09">09.VII.2012</date>
|
||
, Dai, Peng & Yin leg. (
|
||
<collectionCode id="ED3E11ED47024A71FC061578FBEF4E73" box="[955,1036,332,356]" pageId="4" pageNumber="425">SNUC</collectionCode>
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); 5 ♂♂, 3 ♀♀, Leshan, Emei Shan,
|
||
<geoCoordinate id="EE1BEFEF47024A71FF2A155BFEF84E9F" box="[151,283,367,392]" direction="north" orientation="latitude" pageId="4" pageNumber="425" precision="15" value="29.564444">29°33'52''N</geoCoordinate>
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,
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<geoCoordinate id="EE1BEFEF47024A71FE951544FE5B4E9F" box="[296,440,368,392]" direction="east" orientation="longitude" pageId="4" pageNumber="425" precision="15" value="103.37723">103°22'38''E</geoCoordinate>
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,
|
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<quantity id="4CD724CD47024A71FE781544FDFF4E90" box="[453,540,368,392]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="425" unit="m" value="1100.0">1100 m</quantity>
|
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,
|
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<date id="FF91AFE847024A71FD951544FD504E9F" box="[552,691,368,392]" pageId="4" pageNumber="425" value="2012-07-27">27.VII.2012</date>
|
||
, Zhong Peng leg. (
|
||
<collectionCode id="ED3E11ED47024A71FC371544FC384E9F" box="[906,987,368,392]" pageId="4" pageNumber="425">SNUC</collectionCode>
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); 1 ♀, Leshan, Emei Shan,
|
||
<geoCoordinate id="EE1BEFEF47024A71FAAF1544FA7B4E9F" box="[1298,1432,368,392]" direction="north" orientation="latitude" pageId="4" pageNumber="425" precision="15" value="29.552221">29°33'08''N</geoCoordinate>
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,
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<geoCoordinate id="EE1BEFEF47024A71FF2A15A0FEC54EBB" box="[151,294,404,428]" direction="east" orientation="longitude" pageId="4" pageNumber="425" precision="15" value="103.37195">103°22'19''E</geoCoordinate>
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||
,
|
||
<quantity id="4CD724CD47024A71FE8915A0FE6F4EBB" box="[308,396,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="425" unit="m" value="1300.0">1300 m</quantity>
|
||
,
|
||
<date id="FF91AFE847024A71FE2415A0FDC74EBA" box="[409,548,404,429]" pageId="4" pageNumber="425" value="2013-07-05">05.VII.2013</date>
|
||
, Xiao Bian leg. (
|
||
<collectionCode id="ED3E11ED47024A71FD5B15A0FCD44EBB" box="[742,823,404,428]" pageId="4" pageNumber="425">SNUC</collectionCode>
|
||
);
|
||
<specimenCount id="9D2942A147024A71FCF115A0FC9A4EBB" box="[844,889,404,428]" pageId="4" pageNumber="425" type="generic">1 ♂</specimenCount>
|
||
, 1 ♀, Xiling Xueshan,
|
||
<geoCoordinate id="EE1BEFEF47024A71FB3F15A0FB024EBB" box="[1154,1249,404,428]" direction="north" orientation="latitude" pageId="4" pageNumber="425" precision="925" value="30.616667">30°37'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1BEFEF47024A71FB5315A0FAB44EBB" box="[1262,1367,404,428]" direction="east" orientation="longitude" pageId="4" pageNumber="425" precision="925" value="103.13333">103°08'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD724CD47024A71FAD915A0FF4C4ED8" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="425" unit="m" value="2100.0">2100 m</quantity>
|
||
,
|
||
<date id="FF91AFE847024A71FF06158CFED24EC7" box="[187,305,440,464]" pageId="4" pageNumber="425" value="2013-02-04">04.II.2013</date>
|
||
, Zhang leg. (
|
||
<collectionCode id="ED3E11ED47024A71FE79158CFDF64EC7" box="[452,533,440,464]" pageId="4" pageNumber="425">SNUC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C335DAA347024A71FF7A15E8FC384DF8" pageId="4" pageNumber="425" type="discussion">
|
||
<paragraph id="8B90892847024A71FF7A15E8FC384DF8" blockId="4.[151,1437,151,752]" pageId="4" pageNumber="425">
|
||
<emphasis id="B95B553A47024A71FF7A15E8FEA74EE2" bold="true" box="[199,324,476,501]" pageId="4" pageNumber="425">Comment.</emphasis>
|
||
A comparison of the redescription and illustrations of
|
||
<taxonomicName id="4C2FF2AB47024A71FC1315E9FBAA4EE3" box="[942,1097,476,500]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="szechuanus">
|
||
<emphasis id="B95B553A47024A71FC1315E9FBAA4EE3" box="[942,1097,476,500]" italics="true" pageId="4" pageNumber="425">P. szechuanus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFBEF4D947024A71FBD315E8FADA4EE3" author="Li" box="[1134,1337,476,501]" pageId="4" pageNumber="425" refString="Li, X. - Y. & Zhou, H. - Z. (2009) Phylogenetic analysis and taxonomy of the subgenus Gnathopaederus Chapin and related groups of Paederus s. l. (Coleoptera: Staphylinidae: Paederinae). Invertebrate Systematics, 23, 422 - 444. http: // dx. doi. org / 10.1071 / IS 09015" type="journal article" year="2009">Li & Zhou (2009)</bibRefCitation>
|
||
with the above material revealed some differences in the sexual characters. According to
|
||
<bibRefCitation id="EFBEF4D947024A71FB8515CBFAE84D0F" author="Li" box="[1080,1291,511,536]" pageId="4" pageNumber="425" refString="Li, X. - Y. & Zhou, H. - Z. (2009) Phylogenetic analysis and taxonomy of the subgenus Gnathopaederus Chapin and related groups of Paederus s. l. (Coleoptera: Staphylinidae: Paederinae). Invertebrate Systematics, 23, 422 - 444. http: // dx. doi. org / 10.1071 / IS 09015" type="journal article" year="2009">Li & Zhou (2009)</bibRefCitation>
|
||
, the anterior margin of the male labrum is distinctly sinuate (less sinuate,
|
||
<figureCitation id="131495AD47024A71FCDF1610FC464D2B" box="[866,933,548,573]" captionStart="FIGURE 3" captionStartId="3.[151,250,1848,1870]" captionTargetBox="[166,1435,455,1817]" captionTargetId="figure@3.[151,1435,446,1826]" captionTargetPageId="3" captionText="FIGURE 3. Paederus szechuanus. A male left mandible B male right mandible C female right mandible D male labrum E female labrum F female tergite VIII G female sternite VIII H female sternite IX I male tergite VIII J male sternite VIII K male tergite IX L, O aedeagus in ventral view M aedeagus in lateral view N aedeagus in dorsal view P the front part of parameres. Scale bars: A – O 0.5 mm, P 0.1 mm." httpUri="https://zenodo.org/record/242629/files/figure.png" pageId="4" pageNumber="425">Fig. 3</figureCitation>
|
||
D), the apical portion of the dorsal plate is curved to the right in dorsal view (left in the material listed above,
|
||
<figureCitation id="131495AD47024A71FCC21673FC5C4D77" box="[895,959,583,608]" captionStart="FIGURE 3" captionStartId="3.[151,250,1848,1870]" captionTargetBox="[166,1435,455,1817]" captionTargetId="figure@3.[151,1435,446,1826]" captionTargetPageId="3" captionText="FIGURE 3. Paederus szechuanus. A male left mandible B male right mandible C female right mandible D male labrum E female labrum F female tergite VIII G female sternite VIII H female sternite IX I male tergite VIII J male sternite VIII K male tergite IX L, O aedeagus in ventral view M aedeagus in lateral view N aedeagus in dorsal view P the front part of parameres. Scale bars: A – O 0.5 mm, P 0.1 mm." httpUri="https://zenodo.org/record/242629/files/figure.png" pageId="4" pageNumber="425">Fig. 3</figureCitation>
|
||
N) and the internal sac of the aedeagus has S-shaped sclerotized spines (a long curved spine and a short spine in the material listed above,
|
||
<figureCitation id="131495AD47024A71FB031658FAE14D93" box="[1214,1282,620,645]" captionStart="FIGURE 3" captionStartId="3.[151,250,1848,1870]" captionTargetBox="[166,1435,455,1817]" captionTargetId="figure@3.[151,1435,446,1826]" captionTargetPageId="3" captionText="FIGURE 3. Paederus szechuanus. A male left mandible B male right mandible C female right mandible D male labrum E female labrum F female tergite VIII G female sternite VIII H female sternite IX I male tergite VIII J male sternite VIII K male tergite IX L, O aedeagus in ventral view M aedeagus in lateral view N aedeagus in dorsal view P the front part of parameres. Scale bars: A – O 0.5 mm, P 0.1 mm." httpUri="https://zenodo.org/record/242629/files/figure.png" pageId="4" pageNumber="425">Figs 3</figureCitation>
|
||
L–O). This species is widespread in the eastern Qionglai mountains and subject to considerable intraspecific variation of characters such as size, coloration and punctation, as well as the shape of the head, pronotum, elytra, parameres and internal structures of the aedeagus, but the shape of the labrum is constant.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |