treatments-xml/data/03/83/C8/0383C80DB631A91E588BFB03FBA9E2CD.xml

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<mods:title id="A8E35A2E71E7C789D144FE9808F8D74B">Taxonomy of European Damaeidae (Acari: Oribatida) VII. Redescription of Neobelba pseudopapillipes Bulanova-Zachvatkina, 1967 with comments on its generic status</mods:title>
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<mods:namePart id="9A4BF8F35889104497454503886F30D7">Miko, Ladislav</mods:namePart>
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<mods:namePart id="7F35A37C55186839562DBE416C3D4982">Kolesnikov, Vasiliy B.</mods:namePart>
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<subSubSection id="C3302A90B631A915588BFB03FBCBE3A8" box="[151,1053,1165,1191]" pageId="2" pageNumber="376" type="description">
<paragraph id="8B95791BB631A915588BFB03FBCBE3A8" blockId="2.[151,1053,1165,1191]" box="[151,1053,1165,1191]" pageId="2" pageNumber="376">
<heading id="D0DDCE77B631A915588BFB03FBCBE3A8" bold="true" box="[151,1053,1165,1191]" fontSize="11" level="1" pageId="2" pageNumber="376" reason="1">
<emphasis id="B95EA509B631A915588BFB03FBCBE3A8" bold="true" box="[151,1053,1165,1191]" pageId="2" pageNumber="376">
Redescription of 
<taxonomicName id="4C2A0298B631A9155970FB03FBCBE3A8" ID-CoL="996e3ae7-3fe3-4e6a-8847-dce0c8dcca98" authority="Bulanova-Zachvatkina, 1967" authorityName="Bulanova-Zachvatkina" authorityYear="1967" box="[364,1053,1165,1191]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B631A9155970FB03FD7BE3A8" bold="true" box="[364,685,1165,1191]" italics="true" pageId="2" pageNumber="376">Neobelba pseudopapillipes</emphasis>
<bibRefCitation id="EFBB04EAB631A9155AA9FB03FBCBE3A8" author="Bulanova-Zachvatkina" box="[693,1053,1165,1191]" pageId="2" pageNumber="376" refString="Bulanova-Zachvatkina, E. M. (1967) Pancirnyje kleschtschi - Oribatidy. Vysshaya Shkola, Moscow, 254 pp. [in Russian]" type="book" year="1967">Bulanova-Zachvatkina, 1967</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3302A90B631A915588BFB5AFAE7E2E1" pageId="2" pageNumber="376" type="diagnosis">
<paragraph id="8B95791BB631A915588BFB5AFAE7E2E1" blockId="2.[151,1437,1236,2022]" pageId="2" pageNumber="376">
<emphasis id="B95EA509B631A915588BFB5AFDA9E3E2" bold="true" box="[151,639,1236,1261]" pageId="2" pageNumber="376">Diagnosis (adult, immatures unknown).</emphasis>
Damaeid mite of medium size with increased number of setae on trochanters and femora: setal formula of trochanters 1-1-4-3, setal formula of femora 
<date id="FF945FDBB631A9155C71FB76FB32E21F" box="[1133,1252,1272,1296]" pageId="2" pageNumber="376" value="2010-10-09" valueMax="2010-10-09" valueMin="2010-10-09">10-10-9-9.</date>
Solenidia of all legs, with exception of 
<emphasis id="B95EA509B631A91559BEFA93FE63E23B" box="[418,437,1309,1332]" italics="true" pageId="2" pageNumber="376">φ</emphasis>
1 on tibia I and 
<emphasis id="B95EA509B631A9155A79FA93FDAEE23B" box="[613,632,1309,1332]" italics="true" pageId="2" pageNumber="376">φ</emphasis>
on tibia IV, relatively short. All solenidia of genu I 
<emphasis id="B95EA509B631A9155CDDFA93FB19E23B" box="[1217,1231,1309,1332]" italics="true" pageId="2" pageNumber="376">–</emphasis>
III and tibia II 
<emphasis id="B95EA509B631A9155D6CFA93FAA8E23B" box="[1392,1406,1309,1332]" italics="true" pageId="2" pageNumber="376">–</emphasis>
IV with associated companion seta 
<emphasis id="B95EA509B631A9155A17FACDFDCFE254" box="[523,537,1347,1371]" italics="true" pageId="2" pageNumber="376">d</emphasis>
. Prodorsum without propodolateral apophysis 
<emphasis id="B95EA509B631A9155C28FACAFB94E254" box="[1076,1090,1348,1371]" italics="true" pageId="2" pageNumber="376">P</emphasis>
, with one pair of tubercles in dorsosejugal area (
<emphasis id="B95EA509B631A9155968FAE6FE65E28B" box="[372,435,1384,1412]" italics="true" pageId="2" pageNumber="376">B a–B p</emphasis>
). Both sensillus and interlamellar seta with elongated and attenuated, flagellate tips. Distinct round area (
<emphasis id="B95EA509B631A915599BFA00FE67E2A9" box="[391,433,1422,1446]" italics="true" pageId="2" pageNumber="376">apd</emphasis>
) resembling area porosa present anterior to insertion of interlamellar seta. Notogastral setae, at least in the posterior part of notogaster, clearly bent and characteristically thickened near base, when seen in lateral view. Ventral side without apparent tubercles. Discidium narrow, bent posterolaterad, claw-like.
</paragraph>
</subSubSection>
<subSubSection id="C3302A90B631A91558DBFA77FF0AE0E9" pageId="2" pageNumber="376" type="materials_examined">
<paragraph id="8B95791BB631A91558DBFA77FEEAE171" blockId="2.[151,1437,1236,2022]" pageId="2" pageNumber="376">
<emphasis id="B95EA509B631A91558DBFA77FC59E11D" bold="true" box="[199,911,1529,1554]" pageId="2" pageNumber="376">
Material examined. 
<typeStatus id="5491C7B9B631A91559A3FA77FDF7E11D" box="[447,545,1529,1554]" pageId="2" pageNumber="376" type="neotype">Neotype</typeStatus>
and 14 paraneotypes: 
<collectingCountry id="F33D398BB631A9155B5CFA77FC59E11D" box="[832,911,1529,1554]" name="Russia" pageId="2" pageNumber="376">Russia</collectingCountry>
</emphasis>
, Voronezh region, Ramonskiy district, forest near village of Aydarovo, 
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, 
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, about 
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a.s.l.: mixed broad-leaved forest with dominance of 
<emphasis id="B95EA509B631A91558A9F9CCFE9EE156" box="[181,328,1602,1625]" italics="true" pageId="2" pageNumber="376">
<taxonomicName id="4C2A0298B631A91558A9F9CCFE92E156" box="[181,324,1602,1625]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="376" phylum="Tracheophyta" rank="species" species="spp">Quercus spp</taxonomicName>
.
</emphasis>
, 
<taxonomicName id="4C2A0298B631A915594AF9CCFDD9E155" box="[342,527,1602,1626]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="376" phylum="Tracheophyta" rank="species" species="tremula">
<emphasis id="B95EA509B631A915594AF9CCFDD9E155" box="[342,527,1602,1626]" italics="true" pageId="2" pageNumber="376">Populus tremula</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2A0298B631A9155A01F9CCFD11E155" box="[541,711,1602,1626]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="376" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="B95EA509B631A9155A01F9CCFD11E155" box="[541,711,1602,1626]" italics="true" pageId="2" pageNumber="376">Betula pendula</emphasis>
</taxonomicName>
and 
<emphasis id="B95EA509B631A9155AE2F9CCFC79E155" box="[766,943,1602,1626]" italics="true" pageId="2" pageNumber="376">
<taxonomicName id="4C2A0298B631A9155AE2F9CCFC7DE155" box="[766,939,1602,1626]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="376" phylum="Tracheophyta" rank="species" species="silvestris">Pinus silvestris</taxonomicName>
,
</emphasis>
in the forest floor; collected 
<date id="FF945FDBB631A9155CE8F9CFFAB0E155" box="[1268,1382,1601,1626]" pageId="2" pageNumber="376" value="2012-06-16">16.6.2012</date>
, leg. V. Kolesnikov.
</paragraph>
<paragraph id="8B95791BB631A91558DBF907FA46E001" blockId="2.[151,1437,1236,2022]" pageId="2" pageNumber="376">
The 
<typeStatus id="5491C7B9B631A915591CF904FEE6E1AD" box="[256,304,1674,1698]" pageId="2" pageNumber="376">type</typeStatus>
series will be deposited in the following collections: The 
<typeStatus id="5491C7B9B631A9155BE1F904FB80E1AD" box="[1021,1110,1674,1698]" pageId="2" pageNumber="376" type="neotype">neotype</typeStatus>
and 2 paraneotypes in the Acarological Collections of Senckenberg Museum für Naturkunde Görlitz (
<collectingCountry id="F33D398BB631A9155C2BF920FB73E1C9" box="[1079,1189,1710,1734]" name="Germany" pageId="2" pageNumber="376">Germany</collectingCountry>
), 2 paraneotypes in Arachnological Collections in zoological department of the National Museum in Prague (
<collectingCountry id="F33D398BB631A9155CA5F95FFAA0E1E5" box="[1209,1398,1745,1770]" name="Czech Republic" pageId="2" pageNumber="376">Czech Republic</collectingCountry>
), 3 paraneotypes in private collection of Ladislav Miko and remaining 7 paraneotypes in collection of V. Kolesnikov.
</paragraph>
<paragraph id="8B95791BB631A91558DBF897FF0AE0E9" blockId="2.[151,1437,1236,2022]" pageId="2" pageNumber="376">
<emphasis id="B95EA509B631A91558DBF897FB92E03E" bold="true" box="[199,1092,1817,1842]" pageId="2" pageNumber="376">
Further material examined (not included in the 
<typeStatus id="5491C7B9B631A9155B0FF897FCA6E03D" box="[787,880,1817,1842]" pageId="2" pageNumber="376" type="neotype">neotype</typeStatus>
series). 
<collectingCountry id="F33D398BB631A9155BCBF897FBE8E03D" box="[983,1086,1817,1842]" name="Slovakia" pageId="2" pageNumber="376">Slovakia</collectingCountry>
,
</emphasis>
East Slovak region, Košice – Kavečany, in the area of zoological gardens, 
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, 
<geoCoordinate id="EE1E1FDCB631A9155B26F8B3FC14E059" box="[826,962,1853,1878]" direction="east" orientation="longitude" pageId="2" pageNumber="376" precision="15" value="21.198332">21°11’54”E</geoCoordinate>
), 
<quantity id="4CD2D4FEB631A9155BC5F8B0FBF5E059" box="[985,1059,1854,1878]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="376" unit="m" value="430.0">430 m</quantity>
a.s.l., mixed broad-leaved forest with dominance of 
<taxonomicName id="4C2A0298B631A915596FF8ECFDE2E075" box="[371,564,1890,1914]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="376" phylum="Tracheophyta" rank="species" species="betulus">
<emphasis id="B95EA509B631A915596FF8ECFDE2E075" box="[371,564,1890,1914]" italics="true" pageId="2" pageNumber="376">Carpinus betulus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A0298B631A9155A70F8ECFD1CE076" box="[620,714,1890,1913]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="376" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EA509B631A9155A70F8ECFD1CE076" box="[620,714,1890,1913]" italics="true" pageId="2" pageNumber="376">Quercus</emphasis>
</taxonomicName>
<taxonomicName id="4C2A0298B631A9155ACEF8ECFD2FE076" box="[722,761,1890,1913]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="376" phylum="Tracheophyta" rank="species" species="spp">spp</taxonomicName>
., with deforested grassy patches, in the litter. 4 individuals, collected 
<date id="FF945FDBB631A9155915F808FEB8E091" box="[265,366,1926,1950]" pageId="2" pageNumber="376" value="1989-08-02">2.8.1989</date>
, leg Ľubomír Kováč. 2 individuals deposited in Arachnological Collections in zoological department of the National Museum in Prague (
<collectingCountry id="F33D398BB631A9155AA2F827FCACE0CD" box="[702,890,1961,1986]" name="Czech Republic" pageId="2" pageNumber="376">Czech Republic</collectingCountry>
), 2 individuals in private collection of Ladislav Miko.
</paragraph>
</subSubSection>
<subSubSection id="C3302A90B630A91F58DBFF19FE27E3C6" lastPageId="8" lastPageNumber="382" pageId="3" pageNumber="377" type="description">
<paragraph id="8B95791BB630A91458DBFF19FD52E7F7" blockId="3.[151,1436,151,248]" pageId="3" pageNumber="377">
<emphasis id="B95EA509B630A91458DBFF19FE84E7BF" bold="true" box="[199,338,151,176]" pageId="3" pageNumber="377">Description</emphasis>
(adult). 
<figureCitation id="1311659EB630A91459AAFF19FDCBE7BF" box="[438,541,151,176]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[151,250,1918,1940]" captionStartId-1="5.[151,250,1129,1151]" captionStartId-2="6.[151,250,1945,1967]" captionStartId-3="7.[151,250,1941,1963]" captionTargetBox-0="[168,1421,1064,1896]" captionTargetBox-1="[179,1423,207,1093]" captionTargetBox-2="[186,1401,204,1905]" captionTargetBox-3="[224,1374,213,1885]" captionTargetId-0="figure@4.[151,1436,1053,1901]" captionTargetId-1="figure@5.[151,1436,193,1108]" captionTargetId-2="figure@6.[151,1436,193,1924]" captionTargetId-3="figure@7.[195,1385,193,1907]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." captionText-1="FIGURE 2. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — lateral view, details of prodorsum and gnatosoma. A—lateral view of the body, B—prodorsum, C—chelicere, D—palp, E—notogastral setae of individuals collected in Slovakia. Bar representing 100 µm (A, E) or 50 µm (B – D). Details of chelicere (setae cha and chb) and palp (distal part of palp tarsus) further enlarged to allow observation of details. Abbreviations: apd—area prodorsal, dis—discidium, gl—notogastral gland opening. Other markings represent names of setae, tubercles or lyrifissures." captionText-2="FIGURE 3. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — legs. A—leg I, B—femur and genu II, C—femur and genu III, D—leg IV, E—enlarged detail of solenidion and coupled seta d of tibia IV. Bar representing 100 µm (A – D)." captionText-3="FIGURE 4. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — legs II and III (drawings of V. B. Kolesnikov). A—leg II (segments individually depicted), B—leg III (segments individually depicted). Bar representing 50 µm." httpUri-0="https://zenodo.org/record/229454/files/figure.png" httpUri-1="https://zenodo.org/record/229455/files/figure.png" httpUri-2="https://zenodo.org/record/229456/files/figure.png" httpUri-3="https://zenodo.org/record/229457/files/figure.png" pageId="3" pageNumber="377">
Figs 1 
<emphasis id="B95EA509B630A91459E2FF17FDDAE7BF" box="[510,524,153,176]" italics="true" pageId="3" pageNumber="377">–</emphasis>
4
</figureCitation>
. 
<emphasis id="B95EA509B630A9145A36FF16FD78E7A0" box="[554,686,152,175]" italics="true" pageId="3" pageNumber="377">Dimensions</emphasis>
Total body length 480 
<emphasis id="B95EA509B630A9145BAAFF17FC12E7BF" box="[950,964,153,176]" italics="true" pageId="3" pageNumber="377">–</emphasis>
530 (510), ventral length 430 
<emphasis id="B95EA509B630A9145D0AFF17FAF2E7BF" box="[1302,1316,153,176]" italics="true" pageId="3" pageNumber="377">–</emphasis>
498 (452); maximal width of notogaster 260 
<emphasis id="B95EA509B630A9145A0EFF33FDF6E7DB" box="[530,544,189,212]" italics="true" pageId="3" pageNumber="377">–</emphasis>
300 (278), maximal width of prodorsum 185 
<emphasis id="B95EA509B630A9145C03FF33FBFBE7DB" box="[1055,1069,189,212]" italics="true" pageId="3" pageNumber="377">–</emphasis>
230 (208); n=10. Length of legs and other detailed measurements in 
<tableCitation id="C6A84CA0B630A9145A36FF51FDA8E7F7" box="[554,638,223,248]" captionStart="TABLE 1" captionStartId="3.[151,239,296,319]" captionText="TABLE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — measurements." pageId="3" pageNumber="377">Table 1</tableCitation>
.
</paragraph>
<caption id="DF552993B630A914588BFEA6FB13E630" box="[151,1221,295,319]" pageId="3" pageNumber="377">
<paragraph id="8B95791BB630A914588BFEA6FB13E630" blockId="3.[151,1221,295,319]" box="[151,1221,295,319]" pageId="3" pageNumber="377">
<emphasis id="B95EA509B630A914588BFEA6FEDCE630" bold="true" box="[151,266,296,319]" pageId="3" pageNumber="377">TABLE 1.</emphasis>
<taxonomicName id="4C2A0298B630A914590CFEA9FBD1E630" authority="Bulanova-Zachvatkina, 1967" authorityName="Bulanova-Zachvatkina" authorityYear="1967" box="[272,1031,295,319]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="pseudopapillipes" subGenus="Neobelba">
<emphasis id="B95EA509B630A914590CFEA9FD6DE630" box="[272,699,295,319]" italics="true" pageId="3" pageNumber="377">Metabelba (Neobelba) pseudopapillipes</emphasis>
(Bulanova-Zachvatkina, 1967)
</taxonomicName>
—measurements.
</paragraph>
</caption>
<paragraph id="8B95791BB630A914588BFEE6FE32E671" blockId="3.[151,484,360,382]" box="[151,484,360,382]" pageId="3" pageNumber="377">Lengths of setae (all in µm), n=5</paragraph>
<paragraph id="8B95791BB630A914588BFE28FB82E6B3" blockId="3.[151,1108,422,444]" box="[151,1108,422,444]" pageId="3" pageNumber="377">Prodorsum Notogaster Ventral part of body</paragraph>
<paragraph id="8B95791BB630A914588BFE68FDFBE446" blockId="3.[151,1111,485,743]" lastBlockId="3.[151,557,819,841]" pageId="3" pageNumber="377">
<emphasis id="B95EA509B630A914588BFE68FF7BE6F4" box="[151,173,486,507]" italics="true" pageId="3" pageNumber="377">ro</emphasis>
67 (62 
<emphasis id="B95EA509B630A914592AFE68FE95E6F4" box="[310,323,486,507]" italics="true" pageId="3" pageNumber="377">–</emphasis>
70) 
<emphasis id="B95EA509B630A9145A13FE68FDF7E6F1" box="[527,545,486,510]" italics="true" pageId="3" pageNumber="377">c1</emphasis>
96 (90 
<emphasis id="B95EA509B630A9145AB2FE68FD6DE6F4" box="[686,699,486,507]" italics="true" pageId="3" pageNumber="377">–</emphasis>
99) 
<emphasis id="B95EA509B630A9145B95FE6BFC75E6F4" box="[905,931,485,507]" italics="true" pageId="3" pageNumber="377">1a</emphasis>
31 (30 
<emphasis id="B95EA509B630A9145C35FE68FBE0E6F4" box="[1065,1078,486,507]" italics="true" pageId="3" pageNumber="377">–</emphasis>
32) 
<emphasis id="B95EA509B630A914588BFD88FF7FE513" box="[151,169,518,540]" italics="true" pageId="3" pageNumber="377">le</emphasis>
75 (70 
<emphasis id="B95EA509B630A914592AFD89FE95E513" box="[310,323,519,540]" italics="true" pageId="3" pageNumber="377">–</emphasis>
78) 
<emphasis id="B95EA509B630A9145A13FD88FDCBE52F" box="[527,541,518,544]" italics="true" pageId="3" pageNumber="377">l a</emphasis>
86 (81 
<emphasis id="B95EA509B630A9145AB2FD89FD6DE513" box="[686,699,519,540]" italics="true" pageId="3" pageNumber="377">–</emphasis>
90) 
<emphasis id="B95EA509B630A9145B95FD88FC75E513" box="[905,931,518,540]" italics="true" pageId="3" pageNumber="377">2a</emphasis>
32 (30 
<emphasis id="B95EA509B630A9145C35FD89FBE0E513" box="[1065,1078,519,540]" italics="true" pageId="3" pageNumber="377">–</emphasis>
34) 
<emphasis id="B95EA509B630A914588BFDA7FF7DE531" box="[151,171,553,574]" italics="true" pageId="3" pageNumber="377">in</emphasis>
124 (115 
<emphasis id="B95EA509B630A9145953FDA7FE8DE531" box="[335,347,553,574]" italics="true" pageId="3" pageNumber="377">–</emphasis>
130) 
<emphasis id="B95EA509B630A9145A13FDA9FDCBE54D" box="[527,541,551,578]" italics="true" pageId="3" pageNumber="377">l p</emphasis>
92 (87 
<emphasis id="B95EA509B630A9145AB2FDA7FD6DE531" box="[686,699,553,574]" italics="true" pageId="3" pageNumber="377">–</emphasis>
97) 
<emphasis id="B95EA509B630A9145B95FDA9FC77E532" box="[905,929,551,573]" italics="true" pageId="3" pageNumber="377">3c</emphasis>
37 (34 
<emphasis id="B95EA509B630A9145C35FDA7FBE0E531" box="[1065,1078,553,574]" italics="true" pageId="3" pageNumber="377">–</emphasis>
39) 
<emphasis id="B95EA509B630A914588BFDC4FF7BE550" box="[151,173,586,607]" italics="true" pageId="3" pageNumber="377">ex</emphasis>
37 (34 
<emphasis id="B95EA509B630A914592AFDC4FE95E550" box="[310,323,586,607]" italics="true" pageId="3" pageNumber="377">–</emphasis>
39) 
<emphasis id="B95EA509B630A9145A13FDC7FDF5E56D" box="[527,547,585,610]" italics="true" pageId="3" pageNumber="377">h3</emphasis>
95 (92 
<emphasis id="B95EA509B630A9145AB2FDC4FD6DE550" box="[686,699,586,607]" italics="true" pageId="3" pageNumber="377">–</emphasis>
98) 
<emphasis id="B95EA509B630A9145B95FDC7FC74E550" box="[905,930,585,607]" italics="true" pageId="3" pageNumber="377">4b</emphasis>
38 (37 
<emphasis id="B95EA509B630A9145C35FDC4FBE0E550" box="[1065,1078,586,607]" italics="true" pageId="3" pageNumber="377">–</emphasis>
39) 
<emphasis id="B95EA509B630A914588BFDE5FF7DE58F" box="[151,171,619,640]" italics="true" pageId="3" pageNumber="377">ss</emphasis>
165 (155 
<emphasis id="B95EA509B630A9145953FDE5FE8AE58F" box="[335,348,619,640]" italics="true" pageId="3" pageNumber="377">–</emphasis>
170) 
<emphasis id="B95EA509B630A9145A13FDE4FDF5E58C" box="[527,547,618,643]" italics="true" pageId="3" pageNumber="377">h1</emphasis>
64 (63 
<emphasis id="B95EA509B630A9145AB2FDE5FD6DE58F" box="[686,699,619,640]" italics="true" pageId="3" pageNumber="377">–</emphasis>
66) 
<emphasis id="B95EA509B630A9145B95FDE5FC4BE58C" box="[905,925,619,643]" italics="true" pageId="3" pageNumber="377">g1</emphasis>
28 (27 
<emphasis id="B95EA509B630A9145C35FDE5FBE0E58F" box="[1065,1078,619,640]" italics="true" pageId="3" pageNumber="377">–</emphasis>
31) 
<emphasis id="B95EA509B630A9145A13FD02FDF5E5AA" box="[527,547,652,677]" italics="true" pageId="3" pageNumber="377">p1</emphasis>
60 (58 
<emphasis id="B95EA509B630A9145AB2FD02FD6DE5AE" box="[686,699,652,673]" italics="true" pageId="3" pageNumber="377">–</emphasis>
62) 
<emphasis id="B95EA509B630A9145B95FD02FC75E5AE" box="[905,931,652,673]" italics="true" pageId="3" pageNumber="377">ag</emphasis>
37 (34 
<emphasis id="B95EA509B630A9145C35FD02FBE0E5AE" box="[1065,1078,652,673]" italics="true" pageId="3" pageNumber="377">–</emphasis>
38) 
<emphasis id="B95EA509B630A9145A13FD23FDF5E5C9" box="[527,547,685,710]" italics="true" pageId="3" pageNumber="377">p3</emphasis>
34 (33 
<emphasis id="B95EA509B630A9145AB2FD23FD6DE5CD" box="[686,699,685,706]" italics="true" pageId="3" pageNumber="377">–</emphasis>
36) 
<emphasis id="B95EA509B630A9145B95FD23FC7CE5C9" box="[905,938,685,710]" italics="true" pageId="3" pageNumber="377">an1</emphasis>
34 (30 
<emphasis id="B95EA509B630A9145C35FD23FBE0E5CD" box="[1065,1078,685,706]" italics="true" pageId="3" pageNumber="377">–</emphasis>
38) 
<emphasis id="B95EA509B630A9145B95FD43FC7CE5E8" box="[905,938,717,743]" italics="true" pageId="3" pageNumber="377">ad1</emphasis>
40 (38 
<emphasis id="B95EA509B630A9145C35FD41FBE0E5EB" box="[1065,1078,719,740]" italics="true" pageId="3" pageNumber="377">–</emphasis>
41) Lengths of legs and leg segments (n=5).
</paragraph>
<paragraph id="8B95791BB630A9145983FCEBFB1DE3CA" pageId="3" pageNumber="377">
<table id="F92A8BBBB63056E85883FCEBFAE3E3CA" box="[159,1333,869,1221]" gridcols="4" gridrows="9" pageId="3" pageNumber="377">
<tr id="351A7B59B63056E85883FCEBFAE3E475" box="[159,1333,869,890]" gridrow="0" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FCEBFDE2E475" box="[159,564,869,890]" gridcol="0" gridrow="0" pageId="3" pageNumber="377">Leg I</th>
<th id="76CB1225B63056E85A83FCEBFCE3E475" box="[671,821,869,890]" gridcol="1" gridrow="0" pageId="3" pageNumber="377">Leg II</th>
<th id="76CB1225B63056E85B83FCEBFBE2E475" box="[927,1076,869,890]" gridcol="2" gridrow="0" pageId="3" pageNumber="377">Leg III</th>
<th id="76CB1225B63056E85C83FCEBFAE3E475" box="[1183,1333,869,890]" gridcol="3" gridrow="0" pageId="3" pageNumber="377">Leg IV</th>
</tr>
<tr id="351A7B59B63056E85883FC03FAE3E4AC" box="[159,1333,909,931]" gridrow="1" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FC03FDE2E4AC" box="[159,564,909,931]" gridcol="0" gridrow="1" pageId="3" pageNumber="377">trochanter -</th>
<td id="76CB1225B63056E85A83FC03FCE3E4AC" box="[671,821,909,931]" gridcol="1" gridrow="1" pageId="3" pageNumber="377">-</td>
<td id="76CB1225B63056E85B83FC03FBE2E4AC" box="[927,1076,909,931]" gridcol="2" gridrow="1" pageId="3" pageNumber="377">
62(60 
<emphasis id="B95EA509B630A9145BFCFC00FC3BE4AC" box="[992,1005,910,931]" italics="true" pageId="3" pageNumber="377">–</emphasis>
63)
</td>
<td id="76CB1225B63056E85C83FC03FAE3E4AC" box="[1183,1333,909,931]" gridcol="3" gridrow="1" pageId="3" pageNumber="377">
110 (109 
<emphasis id="B95EA509B630A9145CE5FC00FAD3E4AC" box="[1273,1285,910,931]" italics="true" pageId="3" pageNumber="377">–</emphasis>
111)
</td>
</tr>
<tr id="351A7B59B63056E85883FC38FAE3E4C2" box="[159,1333,950,973]" gridrow="2" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FC38FDE2E4C2" box="[159,564,950,973]" gridcol="0" gridrow="2" pageId="3" pageNumber="377">
Femur 116 (114 
<emphasis id="B95EA509B630A91459E4FC36FDD2E4C2" box="[504,516,952,973]" italics="true" pageId="3" pageNumber="377">–</emphasis>
117)
</th>
<td id="76CB1225B63056E85A83FC38FCE3E4C2" box="[671,821,950,973]" gridcol="1" gridrow="2" pageId="3" pageNumber="377">
126 (122 
<emphasis id="B95EA509B630A9145AE5FC36FCD0E4C2" box="[761,774,952,973]" italics="true" pageId="3" pageNumber="377">–</emphasis>
131)
</td>
<td id="76CB1225B63056E85B83FC38FBE2E4C2" box="[927,1076,950,973]" gridcol="2" gridrow="2" pageId="3" pageNumber="377">
93 (89 
<emphasis id="B95EA509B630A9145BFCFC36FC3BE4C2" box="[992,1005,952,973]" italics="true" pageId="3" pageNumber="377">–</emphasis>
95)
</td>
<td id="76CB1225B63056E85C83FC38FAE3E4C2" box="[1183,1333,950,973]" gridcol="3" gridrow="2" pageId="3" pageNumber="377">
104 (102 
<emphasis id="B95EA509B630A9145CE6FC36FAD0E4C2" box="[1274,1286,952,973]" italics="true" pageId="3" pageNumber="377">–</emphasis>
106)
</td>
</tr>
<tr id="351A7B59B63056E85883FC6EFAE3E4F9" box="[159,1333,992,1014]" gridrow="3" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FC6EFDE2E4F9" box="[159,564,992,1014]" gridcol="0" gridrow="3" pageId="3" pageNumber="377">
genu 46 (45 
<emphasis id="B95EA509B630A91459FCFC6FFE3BE4F9" box="[480,493,993,1014]" italics="true" pageId="3" pageNumber="377">–</emphasis>
49)
</th>
<td id="76CB1225B63056E85A83FC6EFCE3E4F9" box="[671,821,992,1014]" gridcol="1" gridrow="3" pageId="3" pageNumber="377">
46 (45 
<emphasis id="B95EA509B630A9145AFCFC6FFD3BE4F9" box="[736,749,993,1014]" italics="true" pageId="3" pageNumber="377">–</emphasis>
47)
</td>
<td id="76CB1225B63056E85B83FC6EFBE2E4F9" box="[927,1076,992,1014]" gridcol="2" gridrow="3" pageId="3" pageNumber="377">
32 (30 
<emphasis id="B95EA509B630A9145BFCFC6FFC3BE4F9" box="[992,1005,993,1014]" italics="true" pageId="3" pageNumber="377">–</emphasis>
32)
</td>
<td id="76CB1225B63056E85C83FC6EFAE3E4F9" box="[1183,1333,992,1014]" gridcol="3" gridrow="3" pageId="3" pageNumber="377">
45 (44 
<emphasis id="B95EA509B630A9145CFCFC6FFB3BE4F9" box="[1248,1261,993,1014]" italics="true" pageId="3" pageNumber="377">–</emphasis>
48)
</td>
</tr>
<tr id="351A7B59B63056E85883FB87FAE3E310" box="[159,1333,1033,1055]" gridrow="4" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FB87FDE2E310" box="[159,564,1033,1055]" gridcol="0" gridrow="4" pageId="3" pageNumber="377">
Tibia 64 (62 
<emphasis id="B95EA509B630A91459FCFB84FE3BE310" box="[480,493,1034,1055]" italics="true" pageId="3" pageNumber="377">–</emphasis>
66)
</th>
<td id="76CB1225B63056E85A83FB87FCE3E310" box="[671,821,1033,1055]" gridcol="1" gridrow="4" pageId="3" pageNumber="377">
70 (68 
<emphasis id="B95EA509B630A9145AFCFB84FD3BE310" box="[736,749,1034,1055]" italics="true" pageId="3" pageNumber="377">–</emphasis>
73)
</td>
<td id="76CB1225B63056E85B83FB87FBE2E310" box="[927,1076,1033,1055]" gridcol="2" gridrow="4" pageId="3" pageNumber="377">
66 (65 
<emphasis id="B95EA509B630A9145BFCFB84FC3BE310" box="[992,1005,1034,1055]" italics="true" pageId="3" pageNumber="377">–</emphasis>
67)
</td>
<td id="76CB1225B63056E85C83FB87FAE3E310" box="[1183,1333,1033,1055]" gridcol="3" gridrow="4" pageId="3" pageNumber="377">
81 (79 
<emphasis id="B95EA509B630A9145CFCFB84FB3BE310" box="[1248,1261,1034,1055]" italics="true" pageId="3" pageNumber="377">–</emphasis>
84)
</td>
</tr>
<tr id="351A7B59B63056E85883FBBDFAE3E346" box="[159,1333,1075,1097]" gridrow="5" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FBBDFDE2E346" box="[159,564,1075,1097]" gridcol="0" gridrow="5" pageId="3" pageNumber="377">
tarsus 148 (146 
<emphasis id="B95EA509B630A91459E5FBBAFDD0E346" box="[505,518,1076,1097]" italics="true" pageId="3" pageNumber="377">–</emphasis>
150)
</th>
<td id="76CB1225B63056E85A83FBBDFCE3E346" box="[671,821,1075,1097]" gridcol="1" gridrow="5" pageId="3" pageNumber="377">
151 (149 
<emphasis id="B95EA509B630A9145AE5FBBAFCD0E346" box="[761,774,1076,1097]" italics="true" pageId="3" pageNumber="377">–</emphasis>
152)
</td>
<td id="76CB1225B63056E85B83FBBDFBE2E346" box="[927,1076,1075,1097]" gridcol="2" gridrow="5" pageId="3" pageNumber="377">
162 (159 
<emphasis id="B95EA509B630A9145BE5FBBAFBD0E346" box="[1017,1030,1076,1097]" italics="true" pageId="3" pageNumber="377">–</emphasis>
164)
</td>
<td id="76CB1225B63056E85C83FBBDFAE3E346" box="[1183,1333,1075,1097]" gridcol="3" gridrow="5" pageId="3" pageNumber="377">
161 (158 
<emphasis id="B95EA509B630A9145CE6FBBAFAD0E346" box="[1274,1286,1076,1097]" italics="true" pageId="3" pageNumber="377">–</emphasis>
163)
</td>
</tr>
<tr id="351A7B59B63056E85883FBD2FAE3E37D" box="[159,1333,1116,1138]" gridrow="6" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FBD2FDE2E37D" box="[159,564,1116,1138]" gridcol="0" gridrow="6" pageId="3" pageNumber="377">
total 374 (370 
<emphasis id="B95EA509B630A91459E5FBD3FDD0E37D" box="[505,518,1117,1138]" italics="true" pageId="3" pageNumber="377">–</emphasis>
376)
</th>
<td id="76CB1225B63056E85A83FBD2FCE3E37D" box="[671,821,1116,1138]" gridcol="1" gridrow="6" pageId="3" pageNumber="377">
393 (392 
<emphasis id="B95EA509B630A9145AE5FBD3FCD0E37D" box="[761,774,1117,1138]" italics="true" pageId="3" pageNumber="377">–</emphasis>
400)
</td>
<td id="76CB1225B63056E85B83FBD2FBE2E37D" box="[927,1076,1116,1138]" gridcol="2" gridrow="6" pageId="3" pageNumber="377">
413 (406 
<emphasis id="B95EA509B630A9145BE5FBD3FBD0E37D" box="[1017,1030,1117,1138]" italics="true" pageId="3" pageNumber="377">–</emphasis>
417)
</td>
<td id="76CB1225B63056E85C83FBD2FAE3E37D" box="[1183,1333,1116,1138]" gridcol="3" gridrow="6" pageId="3" pageNumber="377">
501 (492 
<emphasis id="B95EA509B630A9145CE6FBD3FAD0E37D" box="[1274,1286,1117,1138]" italics="true" pageId="3" pageNumber="377">–</emphasis>
510)
</td>
</tr>
<tr id="351A7B59B63056E85883FB0BFAE3E394" box="[159,1333,1157,1179]" gridrow="7" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FB0BFDE2E394" box="[159,564,1157,1179]" gridcol="0" gridrow="7" pageId="3" pageNumber="377">leg/ventral length 0,83</th>
<td id="76CB1225B63056E85A83FB0BFCE3E394" box="[671,821,1157,1179]" gridcol="1" gridrow="7" pageId="3" pageNumber="377">0,87</td>
<td id="76CB1225B63056E85B83FB0BFBE2E394" box="[927,1076,1157,1179]" gridcol="2" gridrow="7" pageId="3" pageNumber="377">0,91</td>
<td id="76CB1225B63056E85C83FB0BFAE3E394" box="[1183,1333,1157,1179]" gridcol="3" gridrow="7" pageId="3" pageNumber="377">1,11</td>
</tr>
<tr id="351A7B59B63056E85883FB21FAE3E3CA" box="[159,1333,1199,1221]" gridrow="8" pageId="3" pageNumber="377">
<th id="76CB1225B63056E85883FB21FDE2E3CA" box="[159,564,1199,1221]" gridcol="0" gridrow="8" pageId="3" pageNumber="377">leg/total body length 0,73</th>
<td id="76CB1225B63056E85A83FB21FCE3E3CA" box="[671,821,1199,1221]" gridcol="1" gridrow="8" pageId="3" pageNumber="377">0,77</td>
<td id="76CB1225B63056E85B83FB21FBE2E3CA" box="[927,1076,1199,1221]" gridcol="2" gridrow="8" pageId="3" pageNumber="377">0,81</td>
<td id="76CB1225B63056E85C83FB21FAE3E3CA" box="[1183,1333,1199,1221]" gridcol="3" gridrow="8" pageId="3" pageNumber="377">0,98</td>
</tr>
</table>
</paragraph>
<paragraph id="8B95791BB630A91458DBFA84FB94E138" blockId="3.[151,1437,1289,2026]" pageId="3" pageNumber="377">
<emphasis id="B95EA509B630A91458DBFA84FE9BE22E" box="[199,333,1290,1313]" italics="true" pageId="3" pageNumber="377">Integument.</emphasis>
Color medium reddish brown, individuals from 
<collectingCountry id="F33D398BB630A9145B6EFA87FC02E22D" box="[882,980,1289,1314]" name="Slovakia" pageId="3" pageNumber="377">Slovakia</collectingCountry>
lighter, yellowish brown. Almost whole body surface covered by thick layer of cerotegument, mostly amorphous and/or granular (epimeral area), with filamentous excrescences present as well, being most prominent in sejugal area. Rostral and lamellar seta covered by cerotegument, except of tip of 
<emphasis id="B95EA509B630A9145A31FAFCFD93E286" box="[557,581,1394,1417]" italics="true" pageId="3" pageNumber="377">ro</emphasis>
(
<figureCitation id="1311659EB630A9145A45FAFFFD4BE285" box="[601,669,1393,1418]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="3" pageNumber="377">Fig. 1</figureCitation>
D). Club-like bunch of filamentous and amorphous cerotegument present on distal part of sensillus, about between half and three quarters of its length, just before distal attenuated part (
<figureCitation id="1311659EB630A91458CEFA38FEC7E2C0" box="[210,273,1462,1487]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="3" pageNumber="377">Fig. 1</figureCitation>
C). Notogastral setae without cerotegument cover. Legs covered by thick layer of cerotegument, except distal part of all tarsi (only showed on 
<figureCitation id="1311659EB630A9145A6BFA57FD15E2FD" box="[631,707,1497,1522]" captionStart="FIGURE 3" captionStartId="6.[151,250,1945,1967]" captionTargetBox="[186,1401,204,1905]" captionTargetId="figure@6.[151,1436,193,1924]" captionTargetPageId="6" captionText="FIGURE 3. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — legs. A—leg I, B—femur and genu II, C—femur and genu III, D—leg IV, E—enlarged detail of solenidion and coupled seta d of tibia IV. Bar representing 100 µm (A – D)." httpUri="https://zenodo.org/record/229456/files/figure.png" pageId="3" pageNumber="377">Figs 3</figureCitation>
and 
<figureCitation id="1311659EB630A9145B19FA54FCC3E2FD" box="[773,789,1498,1522]" captionStart="FIGURE 4" captionStartId="7.[151,250,1941,1963]" captionTargetBox="[224,1374,213,1885]" captionTargetId="figure@7.[195,1385,193,1907]" captionTargetPageId="7" captionText="FIGURE 4. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — legs II and III (drawings of V. B. Kolesnikov). A—leg II (segments individually depicted), B—leg III (segments individually depicted). Bar representing 50 µm." httpUri="https://zenodo.org/record/229457/files/figure.png" pageId="3" pageNumber="377">4</figureCitation>
, in other Figs omitted). Underlying cuticle smooth or microtuberculate. Adults bearing gastronotic exuviae of immature stages, often with small particles of adhered foreign material, exuviae however loosely attached and often lost during extraction.
</paragraph>
<paragraph id="8B95791BB630A91358DBF9CFFAC0E582" blockId="3.[151,1437,1289,2026]" lastBlockId="4.[151,1437,151,1019]" lastPageId="4" lastPageNumber="378" pageId="3" pageNumber="377">
<emphasis id="B95EA509B630A91458DBF9CFFE92E156" box="[199,324,1601,1625]" italics="true" pageId="3" pageNumber="377">Prodorsum</emphasis>
. 
<figureCitation id="1311659EB630A914594BF9CFFE4BE155" box="[343,413,1601,1626]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="3" pageNumber="377">Figs 1</figureCitation>
A, 2A, 2B. Prodorsum conical, broadly triangular in dorsal view, sharply narrowed in sejugal area. Rostrum rounded, centrally with weakened cuticle appearing dorsally as lighter triangular area. Insertions of rostral setae (
<emphasis id="B95EA509B630A91459CCF904FE31E1AE" box="[464,487,1674,1697]" italics="true" pageId="3" pageNumber="377">ro</emphasis>
) on small tubercles, which are positioned at end of short ridges or lines running parallel to prodorsum edge. Insertions of lamellar setae (
<emphasis id="B95EA509B630A9145B0BF923FCFDE1CA" box="[791,811,1709,1733]" italics="true" pageId="3" pageNumber="377">le</emphasis>
) on posterior edge of arch, created by more sclerotised ribs running along lateral edge of prodorsum, and further between 
<emphasis id="B95EA509B630A9145B60F95FFC46E1E6" box="[892,912,1745,1769]" italics="true" pageId="3" pageNumber="377">le</emphasis>
insertions, in this area less visible and forming 2-3 less distinct lines. Apophysis 
<emphasis id="B95EA509B630A9145A0BF978FDFEE002" box="[535,552,1782,1805]" italics="true" pageId="3" pageNumber="377">P</emphasis>
absent, edge of prodorsum lateral to bothridia almost rounded, in some cases most lateral part of prodorsum with short, straight edges, subparallel to body axis. Single enantiophyse present in dorsosejugal area (considered to be centrodorsal enantiophyse 
<emphasis id="B95EA509B630A9145B6EF8B0FC50E05A" box="[882,902,1854,1877]" italics="true" pageId="3" pageNumber="377">D</emphasis>
, for questions of homology see discussion in Remarks), well developed, with anterior tubercle (
<emphasis id="B95EA509B630A9145AD6F8ECFD3AE076" box="[714,748,1890,1913]" italics="true" pageId="3" pageNumber="377">Da</emphasis>
) smaller, broadly triangular and sitting on sclerotised oblique ridge running from posterior end of bothridial area. Posterior centrodorsal tubercle (
<emphasis id="B95EA509B630A9145C55F808FBBDE092" box="[1097,1131,1926,1949]" italics="true" pageId="3" pageNumber="377">Dp</emphasis>
) larger, about twice longer than 
<emphasis id="B95EA509B630A91458D2F824FF26E0CE" box="[206,240,1962,1985]" italics="true" pageId="3" pageNumber="377">Da</emphasis>
, conical, its base hidden under anterior edge of notogaster. Enantiophyses 
<emphasis id="B95EA509B630A9145C2DF824FB94E0CE" box="[1073,1090,1962,1985]" italics="true" pageId="3" pageNumber="377">B</emphasis>
(postbothridial) and 
<emphasis id="B95EA509B630A9145D31F824FAEAE0CE" box="[1325,1340,1962,1985]" italics="true" pageId="3" pageNumber="377">L</emphasis>
(lateral) absent, but slight elevations and thickenings of cuticle above the apophyse 
<emphasis id="B95EA509B630A9145BE3F840FBC3E0E5" box="[1023,1045,1998,2026]" italics="true" pageId="3" pageNumber="377">S a</emphasis>
and posterior to bothridia may in dorsal view resemble flat tubercles 
<emphasis id="B95EA509B637A9135A38FF17FDEAE7BB" box="[548,572,153,180]" italics="true" pageId="4" pageNumber="378">L a</emphasis>
.and 
<emphasis id="B95EA509B637A9135A6EFF17FD44E7BF" box="[626,658,153,180]" italics="true" pageId="4" pageNumber="378">B a.</emphasis>
Parastigmatic apophyses spiniform, oblique, with tips overlapping in dorsal view. 
<emphasis id="B95EA509B637A913592DFF31FE91E7D4" box="[305,327,191,219]" italics="true" pageId="4" pageNumber="378">S a</emphasis>
with broader base, and, when seen ventrally, much longer than (about twice as long as) 
<emphasis id="B95EA509B637A9135D63FF31FA43E7D4" box="[1407,1429,191,219]" italics="true" pageId="4" pageNumber="378">S p</emphasis>
. Apophyse 
<emphasis id="B95EA509B637A9135908FF68FEFDE60D" box="[276,299,230,258]" italics="true" pageId="4" pageNumber="378">S p</emphasis>
more narrow, with subparallel edges, better observable in ventral view, sharp-tiped. Bothridium positioned on elevated and more sclerotised area, short but distinct ridge runs from its base anteromediad. Interlamellar seta (
<emphasis id="B95EA509B637A9135971FEBFFE55E647" box="[365,387,305,328]" italics="true" pageId="4" pageNumber="378">in</emphasis>
) inserted on small, distinct tubercle. Small, pore-like opening present behind insertion of 
<emphasis id="B95EA509B637A9135D9FFEBFFA4AE647" box="[1411,1436,305,328]" italics="true" pageId="4" pageNumber="378">ex</emphasis>
(
<figureCitation id="1311659EB637A9135883FEDAFF0CE662" box="[159,218,340,365]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="4" pageNumber="378">Fig 1</figureCitation>
C, 2A), distinct pore visible also in lateral view behind acetabulum II in sejugal area (
<figureCitation id="1311659EB637A9135CD6FEDAFADCE662" box="[1226,1290,340,365]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1151]" captionTargetBox="[179,1423,207,1093]" captionTargetId="figure@5.[151,1436,193,1108]" captionTargetPageId="5" captionText="FIGURE 2. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — lateral view, details of prodorsum and gnatosoma. A—lateral view of the body, B—prodorsum, C—chelicere, D—palp, E—notogastral setae of individuals collected in Slovakia. Bar representing 100 µm (A, E) or 50 µm (B – D). Details of chelicere (setae cha and chb) and palp (distal part of palp tarsus) further enlarged to allow observation of details. Abbreviations: apd—area prodorsal, dis—discidium, gl—notogastral gland opening. Other markings represent names of setae, tubercles or lyrifissures." httpUri="https://zenodo.org/record/229455/files/figure.png" pageId="4" pageNumber="378">Fig 2</figureCitation>
A). Unusual, distinctly pronounced area, resembling porose areas of other oribatids, present anterior to 
<emphasis id="B95EA509B637A9135C83FEF7FB63E69F" box="[1183,1205,377,400]" italics="true" pageId="4" pageNumber="378">in</emphasis>
insertion (prodorsal area, 
<emphasis id="B95EA509B637A91358CFFE12FF2BE6BB" box="[211,253,412,436]" italics="true" pageId="4" pageNumber="378">apd</emphasis>
). Rostral (
<emphasis id="B95EA509B637A913596AFE13FE5BE6BB" box="[374,397,413,436]" italics="true" pageId="4" pageNumber="378">ro</emphasis>
) and lamellar (
<emphasis id="B95EA509B637A9135A25FE12FD9BE6BB" box="[569,589,412,436]" italics="true" pageId="4" pageNumber="378">le</emphasis>
) setae of subequal length, curved inwards, with spinuli (
<emphasis id="B95EA509B637A9135CD4FE12FB0AE6BB" box="[1224,1244,412,436]" italics="true" pageId="4" pageNumber="378">le</emphasis>
) or barbs (
<emphasis id="B95EA509B637A9135D44FE13FAB9E6BB" box="[1368,1391,413,436]" italics="true" pageId="4" pageNumber="378">ro</emphasis>
) on outer curvature, 
<emphasis id="B95EA509B637A913594EFE4EFEB0E6D7" box="[338,358,448,472]" italics="true" pageId="4" pageNumber="378">le</emphasis>
slightly stronger at their base. Interlamellar seta (
<emphasis id="B95EA509B637A9135B81FE4FFC65E6D7" box="[925,947,449,472]" italics="true" pageId="4" pageNumber="378">in</emphasis>
) longer than 
<emphasis id="B95EA509B637A9135C51FE4FFBB2E6D7" box="[1101,1124,449,472]" italics="true" pageId="4" pageNumber="378">ro</emphasis>
and 
<emphasis id="B95EA509B637A9135C89FE4EFB7FE6D7" box="[1173,1193,448,472]" italics="true" pageId="4" pageNumber="378">le</emphasis>
, with sparse barbs in middle part and with long, flagellate tip, the latter often broken away (
<figureCitation id="1311659EB637A9135BF4FE6AFBFBE6F2" box="[1000,1069,484,509]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="4" pageNumber="378">Fig. 1</figureCitation>
C, 2B). Exobothridial seta (
<emphasis id="B95EA509B637A9135D66FE6BFA45E6F3" box="[1402,1427,485,508]" italics="true" pageId="4" pageNumber="378">ex</emphasis>
) relatively long, directed anteriad, bent, gradually attenuated distally, setiform. Bothridial funnel as usual in 
<taxonomicName id="4C2A0298B637A913588BFDA2FECEE54A" box="[151,280,556,581]" class="Arachnida" family="Damaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="378" phylum="Arthropoda" rank="family">Damaeidae</taxonomicName>
, cup-like, relatively large, positioned relatively far laterally. Distinct fields of sigillae (muscle insertions) present on prodorsum, two anteriad—anteriomediad to bothridia and 
<emphasis id="B95EA509B637A9135C5CFDDFFB80E567" box="[1088,1110,593,616]" italics="true" pageId="4" pageNumber="378">in</emphasis>
, one in interbothridial area. Lateral edges of two anterior fields strengthened and more sclerotised, creating a kind of narrow ridge.
</paragraph>
<paragraph id="8B95791BB637A91358DBFD17FB85E4F5" blockId="4.[151,1437,151,1019]" pageId="4" pageNumber="378">
<emphasis id="B95EA509B637A91358DBFD17FE94E5BF" box="[199,322,665,688]" italics="true" pageId="4" pageNumber="378">Notogaster</emphasis>
. 
<figureCitation id="1311659EB637A9135946FD16FE73E5BE" box="[346,421,664,689]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="4" pageNumber="378">Figs 1</figureCitation>
A, 2A. Notogaster circular or sub-circular in dorsal view, in lateral view almost hemispherical, very slightly eccentric with maximum height slightly posterior to center. Notogastral setae strong, setiform to spiniform, relatively long, pointing more or less radially outwards of notogaster. In dorsal view they may appear straight or almost straight (
<figureCitation id="1311659EB637A9135A56FC8AFD5FE412" box="[586,649,772,797]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="4" pageNumber="378">Fig. 1</figureCitation>
D), but almost all clearly bent when seen in lateral view. Proximal part of some setae, seen in lateral view, with distinct and very characteristic thickening, with external edge curved proximally almost rectangularly (
<figureCitation id="1311659EB637A9135A04FCC2FD8FE46A" box="[536,601,844,869]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="4" pageNumber="378">Fig. 1</figureCitation>
D– 
<emphasis id="B95EA509B637A9135A9CFCC2FD40E466" box="[640,662,844,873]" italics="true" pageId="4" pageNumber="378">h1</emphasis>
, 
<figureCitation id="1311659EB637A9135ABBFCC2FD3BE46A" box="[679,749,844,869]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1151]" captionTargetBox="[179,1423,207,1093]" captionTargetId="figure@5.[151,1436,193,1108]" captionTargetPageId="5" captionText="FIGURE 2. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — lateral view, details of prodorsum and gnatosoma. A—lateral view of the body, B—prodorsum, C—chelicere, D—palp, E—notogastral setae of individuals collected in Slovakia. Bar representing 100 µm (A, E) or 50 µm (B – D). Details of chelicere (setae cha and chb) and palp (distal part of palp tarsus) further enlarged to allow observation of details. Abbreviations: apd—area prodorsal, dis—discidium, gl—notogastral gland opening. Other markings represent names of setae, tubercles or lyrifissures." httpUri="https://zenodo.org/record/229455/files/figure.png" pageId="4" pageNumber="378">Fig. 2</figureCitation>
A). In some individuals, some notogastral setae (usually of rows 
<emphasis id="B95EA509B637A91358C4FCFDFF36E484" box="[216,224,883,907]" italics="true" pageId="4" pageNumber="378">l</emphasis>
or 
<emphasis id="B95EA509B637A9135911FCFDFECDE484" box="[269,283,883,907]" italics="true" pageId="4" pageNumber="378">h</emphasis>
) may be bent outwards distally, creating slightly S-form shape in lateral view (
<figureCitation id="1311659EB637A9135CD8FCFDFADAE484" box="[1220,1292,883,908]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1151]" captionTargetBox="[179,1423,207,1093]" captionTargetId="figure@5.[151,1436,193,1108]" captionTargetPageId="5" captionText="FIGURE 2. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — lateral view, details of prodorsum and gnatosoma. A—lateral view of the body, B—prodorsum, C—chelicere, D—palp, E—notogastral setae of individuals collected in Slovakia. Bar representing 100 µm (A, E) or 50 µm (B – D). Details of chelicere (setae cha and chb) and palp (distal part of palp tarsus) further enlarged to allow observation of details. Abbreviations: apd—area prodorsal, dis—discidium, gl—notogastral gland opening. Other markings represent names of setae, tubercles or lyrifissures." httpUri="https://zenodo.org/record/229455/files/figure.png" pageId="4" pageNumber="378">Fig. 2</figureCitation>
E). Posterior notogastral setae (
<emphasis id="B95EA509B637A913597FFC16FE4EE4BB" box="[355,408,920,948]" italics="true" pageId="4" pageNumber="378">p1-p3</emphasis>
<emphasis id="B95EA509B637A9135984FC19FE77E4BF" bold="true" box="[408,417,919,944]" pageId="4" pageNumber="378">)</emphasis>
shorter, bent, without thickening at base, pointing laterad or posterolaterad. Circumgastric row of muscle sigillae distinct, well visible. Two pairs of minute light spots present on notogaster posteriad or posteromediad to insertions of setae 
<emphasis id="B95EA509B637A9135A2DFC6CFD91E4F5" box="[561,583,994,1018]" italics="true" pageId="4" pageNumber="378">la</emphasis>
and 
<emphasis id="B95EA509B637A9135A61FC6CFD45E4F5" box="[637,659,994,1018]" italics="true" pageId="4" pageNumber="378">lp</emphasis>
(indicated by black arrows on 
<figureCitation id="1311659EB637A9135BF2FC6CFBFAE4F5" box="[1006,1068,994,1019]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="4" pageNumber="378">Fig. 1</figureCitation>
A).
</paragraph>
<caption id="DF552993B637A913588BF8F0FCE4E0FF" httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="4" pageNumber="378" targetBox="[168,1421,1064,1896]" targetPageId="4">
<paragraph id="8B95791BB637A913588BF8F0FCE4E0FF" blockId="4.[151,1436,1918,2032]" pageId="4" pageNumber="378">
<emphasis id="B95EA509B637A913588BF8F0FEC1E09C" bold="true" box="[151,279,1918,1940]" pageId="4" pageNumber="378">FIGURE 1.</emphasis>
<taxonomicName id="4C2A0298B637A913593EF8F0FBD3E09B" authority="Bulanova-Zachvatkina, 1967" authorityName="Bulanova-Zachvatkina" authorityYear="1967" box="[290,1029,1918,1940]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="pseudopapillipes" subGenus="Neobelba">
<emphasis id="B95EA509B637A913593EF8F0FE58E09B" box="[290,398,1918,1940]" italics="true" pageId="4" pageNumber="378">Metabelba</emphasis>
(
<emphasis id="B95EA509B637A91359BEF8F0FDD5E09B" box="[418,515,1918,1940]" italics="true" pageId="4" pageNumber="378">Neobelba</emphasis>
) 
<emphasis id="B95EA509B637A9135A0AF8F0FD14E09B" box="[534,706,1918,1940]" italics="true" pageId="4" pageNumber="378">pseudopapillipes</emphasis>
(Bulanova-Zachvatkina, 1967)
</taxonomicName>
—habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C–D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster.
</paragraph>
</caption>
<caption id="DF552993B636A912588BFBE7FBA3E3F5" httpUri="https://zenodo.org/record/229455/files/figure.png" pageId="5" pageNumber="379" targetBox="[179,1423,207,1093]" targetPageId="5">
<paragraph id="8B95791BB636A912588BFBE7FBA3E3F5" blockId="5.[151,1436,1129,1274]" pageId="5" pageNumber="379">
<emphasis id="B95EA509B636A912588BFBE7FEC3E371" bold="true" box="[151,277,1129,1151]" pageId="5" pageNumber="379">FIGURE 2.</emphasis>
<taxonomicName id="4C2A0298B636A9125901FBE7FC20E370" authority="Bulanova-Zachvatkina, 1967" authorityName="Bulanova-Zachvatkina" authorityYear="1967" box="[285,1014,1129,1151]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="pseudopapillipes" subGenus="Neobelba">
<emphasis id="B95EA509B636A9125901FBE7FE5FE370" box="[285,393,1129,1151]" italics="true" pageId="5" pageNumber="379">Metabelba</emphasis>
(
<emphasis id="B95EA509B636A9125987FBE7FE2AE370" box="[411,508,1129,1151]" italics="true" pageId="5" pageNumber="379">Neobelba</emphasis>
) 
<emphasis id="B95EA509B636A9125A10FBE7FD6EE370" box="[524,696,1129,1151]" italics="true" pageId="5" pageNumber="379">pseudopapillipes</emphasis>
(Bulanova-Zachvatkina, 1967)
</taxonomicName>
—lateral view, details of prodorsum and gnatosoma. A—lateral view of the body, B—prodorsum, C—chelicere, D—palp, E—notogastral setae of individuals collected in Slovakia. Bar representing 100 µm (A, E) or 50 µm (B–D). Details of chelicere (setae 
<emphasis id="B95EA509B636A9125C34FB28FB9AE3B3" box="[1064,1100,1190,1212]" italics="true" pageId="5" pageNumber="379">cha</emphasis>
and 
<emphasis id="B95EA509B636A9125C9CFB28FB72E3B3" box="[1152,1188,1190,1212]" italics="true" pageId="5" pageNumber="379">chb</emphasis>
) and palp (distal part of palp tarsus) further enlarged to allow observation of details. Abbreviations: apd—area prodorsal, dis—discidium, gl—notogastral gland opening. Other markings represent names of setae, tubercles or lyrifissures.
</paragraph>
</caption>
<paragraph id="8B95791BB636A91258DBFAA8FE9EE2FC" blockId="5.[151,1436,1318,1997]" pageId="5" pageNumber="379">
<taxonomicName id="4C2A0298B636A91258DBFAA8FE80E231" box="[199,342,1318,1342]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="5" pageNumber="379" phylum="Nematoda" rank="genus">
<emphasis id="B95EA509B636A91258DBFAA8FE80E231" box="[199,342,1318,1342]" italics="true" pageId="5" pageNumber="379">Gnathosoma</emphasis>
</taxonomicName>
. Subcapitulum as usual for family, diarthric, with three pairs of setae. Setae fine, setiform, curved, in ventral view seta 
<emphasis id="B95EA509B636A9125964FAC5FE5AE26D" box="[376,396,1355,1378]" italics="true" pageId="5" pageNumber="379">m</emphasis>
appearing longest, but in lateral view almost equal to 
<emphasis id="B95EA509B636A9125BF0FAC4FC2CE26D" box="[1004,1018,1354,1378]" italics="true" pageId="5" pageNumber="379">h</emphasis>
, seta 
<emphasis id="B95EA509B636A9125C26FAC5FB9EE26D" box="[1082,1096,1355,1378]" italics="true" pageId="5" pageNumber="379">a</emphasis>
slightly shorter. Setal formula of palp 0-2-1-3-9(1), solenidion 
<emphasis id="B95EA509B636A9125A17FAE1FDC8E289" box="[523,542,1391,1414]" italics="true" pageId="5" pageNumber="379">ω</emphasis>
adhered to surface of palp tarsus and hard to observe (
<figureCitation id="1311659EB636A9125C81FAE0FB35E288" box="[1181,1251,1390,1415]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1151]" captionTargetBox="[179,1423,207,1093]" captionTargetId="figure@5.[151,1436,193,1108]" captionTargetPageId="5" captionText="FIGURE 2. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — lateral view, details of prodorsum and gnatosoma. A—lateral view of the body, B—prodorsum, C—chelicere, D—palp, E—notogastral setae of individuals collected in Slovakia. Bar representing 100 µm (A, E) or 50 µm (B – D). Details of chelicere (setae cha and chb) and palp (distal part of palp tarsus) further enlarged to allow observation of details. Abbreviations: apd—area prodorsal, dis—discidium, gl—notogastral gland opening. Other markings represent names of setae, tubercles or lyrifissures." httpUri="https://zenodo.org/record/229455/files/figure.png" pageId="5" pageNumber="379">Fig. 2</figureCitation>
D). Chelicera of shape and relative size as usual in 
<taxonomicName id="4C2A0298B636A9125A0AFA1CFD41E2A4" box="[534,663,1426,1451]" class="Arachnida" family="Damaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="379" phylum="Arthropoda" rank="family">Damaeidae</taxonomicName>
, seta 
<emphasis id="B95EA509B636A9125AC8FA1CFD2BE2A5" box="[724,765,1426,1450]" italics="true" pageId="5" pageNumber="379">cha</emphasis>
longer, bent outwards proximally and inwards distally, with short barbs on external curvature and attenuated tip, seta 
<emphasis id="B95EA509B636A9125B01FA38FC90E2C1" box="[797,838,1462,1486]" italics="true" pageId="5" pageNumber="379">chb</emphasis>
straight, distally bent and with fringe of diminishing barbs (
<figureCitation id="1311659EB636A91258FFFA54FEF0E2FC" box="[227,294,1498,1523]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1151]" captionTargetBox="[179,1423,207,1093]" captionTargetId="figure@5.[151,1436,193,1108]" captionTargetPageId="5" captionText="FIGURE 2. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — lateral view, details of prodorsum and gnatosoma. A—lateral view of the body, B—prodorsum, C—chelicere, D—palp, E—notogastral setae of individuals collected in Slovakia. Bar representing 100 µm (A, E) or 50 µm (B – D). Details of chelicere (setae cha and chb) and palp (distal part of palp tarsus) further enlarged to allow observation of details. Abbreviations: apd—area prodorsal, dis—discidium, gl—notogastral gland opening. Other markings represent names of setae, tubercles or lyrifissures." httpUri="https://zenodo.org/record/229455/files/figure.png" pageId="5" pageNumber="379">Fig. 2</figureCitation>
C).
</paragraph>
<paragraph id="8B95791BB636A91258DBFA70FC85E1C2" blockId="5.[151,1436,1318,1997]" pageId="5" pageNumber="379">
<emphasis id="B95EA509B636A91258DBFA70FE55E119" box="[199,387,1534,1558]" italics="true" pageId="5" pageNumber="379">Epimeral region</emphasis>
. 
<figureCitation id="1311659EB636A912598AFA70FE02E118" box="[406,468,1534,1559]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="5" pageNumber="379">Fig 1</figureCitation>
B. Mentotectum relatively narrow, low. Ventral tubercles absent, however, pair of triangular internal cuticular thickenings present at place of posterior ventrosejugal tubercles (
<emphasis id="B95EA509B636A9125CDEF9ADFB37E135" box="[1218,1249,1571,1594]" italics="true" pageId="5" pageNumber="379">Vp</emphasis>
), which may be misinterpreted as tubercles. Ventrosejugal groove relatively broad and quite deep, laterally distinctly bordered by posterior edge of elongated apophyse 
<emphasis id="B95EA509B636A9125A5FF9E5FD8FE188" box="[579,601,1643,1671]" italics="true" pageId="5" pageNumber="379">S a</emphasis>
. Epimeres III with standard set of 3 setae, epimeral setal formula 3-1-3-4. Epimeral setae subequal in length, fine, setiform and smooth, shorter than other setae of body. Epimeres with muscular sigillae, well visible, particularly closer to axial part.
</paragraph>
<paragraph id="8B95791BB636A91258DBF957FE60E0C2" blockId="5.[151,1436,1318,1997]" pageId="5" pageNumber="379">
<emphasis id="B95EA509B636A91258DBF957FE47E1FE" box="[199,401,1753,1777]" italics="true" pageId="5" pageNumber="379">Anogenital region</emphasis>
. 
<figureCitation id="1311659EB636A9125983F957FE08E1FD" box="[415,478,1753,1778]" captionStart="FIGURE 1" captionStartId="4.[151,250,1918,1940]" captionTargetBox="[168,1421,1064,1896]" captionTargetId="figure@4.[151,1436,1053,1901]" captionTargetPageId="4" captionText="FIGURE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — habitus and dorsal setae. A—dorsal view, B—ventral view, C—bothridial area and sensillus, D—dorsal setae. Bar representing 100 µm (A, B) or 50 µm (C – D). Abbreviations: bo—bothridium, dis—discidium, apd—prodorsal area. Other markings refer to the name of setae or tubercular structures. Black arrows indicate the light spots on the notogaster." httpUri="https://zenodo.org/record/229454/files/figure.png" pageId="5" pageNumber="379">Fig. 1</figureCitation>
B. Discidium (
<emphasis id="B95EA509B636A9125A99F957FD73E1FE" box="[645,677,1753,1777]" italics="true" pageId="5" pageNumber="379">dis</emphasis>
) narrow, elongated and bent backwards, claw-like. Genital opening larger than anal opening, circumgenital area framed by indistinct anterior and posterior ridge, surface of genital plates along setal insertions rugged. Preanal sclerite with two distinct lateral projections. Setal formula of anogenital region as in other 
<taxonomicName id="4C2A0298B636A91259F2F8CBFDA7E051" box="[494,625,1861,1886]" class="Arachnida" family="Damaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="379" phylum="Arthropoda" rank="family">Damaeidae</taxonomicName>
: genital 
<emphasis id="B95EA509B636A9125AC9F8C8FD3AE052" box="[725,748,1862,1885]" italics="true" pageId="5" pageNumber="379">g:</emphasis>
6; aggenital 
<emphasis id="B95EA509B636A9125B95F8C8FC73E052" box="[905,933,1862,1885]" italics="true" pageId="5" pageNumber="379">ag</emphasis>
: 1; anal 
<emphasis id="B95EA509B636A9125C0DF8C8FBFBE052" box="[1041,1069,1862,1885]" italics="true" pageId="5" pageNumber="379">an</emphasis>
: 2; adanal 
<emphasis id="B95EA509B636A9125CA8F8CBFB06E052" box="[1204,1232,1861,1885]" italics="true" pageId="5" pageNumber="379">ad</emphasis>
: 3. Adanal setae, particularly 
<emphasis id="B95EA509B636A912593EF8E7FE90E089" box="[290,326,1897,1926]" italics="true" pageId="5" pageNumber="379">ad3</emphasis>
, inserted far from anal opening and closer to lateral edge of ventral plate. All anogenital setae smooth, setiform, subequal in length, similar to epimeral setae. Lyrifissure 
<emphasis id="B95EA509B636A9125C1CF81EFBF2E0A7" box="[1024,1060,1936,1960]" italics="true" pageId="5" pageNumber="379">iad</emphasis>
apoanal—oblique and divergent from body axis posteriad.
</paragraph>
<caption id="DF552993B635A911588BF817FF36E0E3" httpUri="https://zenodo.org/record/229456/files/figure.png" pageId="6" pageNumber="380" targetBox="[186,1401,204,1905]" targetPageId="6">
<paragraph id="8B95791BB635A911588BF817FF36E0E3" blockId="6.[151,1436,1945,2028]" pageId="6" pageNumber="380">
<emphasis id="B95EA509B635A911588BF817FEC2E0A1" bold="true" box="[151,276,1945,1967]" pageId="6" pageNumber="380">FIGURE 3.</emphasis>
<taxonomicName id="4C2A0298B635A9115900F817FC20E0A0" authority="Bulanova-Zachvatkina, 1967" authorityName="Bulanova-Zachvatkina" authorityYear="1967" box="[284,1014,1945,1967]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="pseudopapillipes" subGenus="Neobelba">
<emphasis id="B95EA509B635A9115900F817FE5EE0A0" box="[284,392,1945,1967]" italics="true" pageId="6" pageNumber="380">Metabelba</emphasis>
(
<emphasis id="B95EA509B635A9115985F817FE2CE0A0" box="[409,506,1945,1967]" italics="true" pageId="6" pageNumber="380">Neobelba</emphasis>
) 
<emphasis id="B95EA509B635A9115A16F817FD60E0A0" box="[522,694,1945,1967]" italics="true" pageId="6" pageNumber="380">pseudopapillipes</emphasis>
(Bulanova-Zachvatkina, 1967)
</taxonomicName>
—legs. A—leg I, B—femur and genu II, C—femur and genu III, D—leg IV, E—enlarged detail of solenidion and coupled seta 
<emphasis id="B95EA509B635A9115C0CF836FBCAE0C1" box="[1040,1052,1976,1998]" italics="true" pageId="6" pageNumber="380">d</emphasis>
of tibia IV. Bar representing 100 µm (A–D).
</paragraph>
</caption>
<caption id="DF552993B634A910588BF81BFA4AE0C6" httpUri="https://zenodo.org/record/229457/files/figure.png" pageId="7" pageNumber="381" targetBox="[224,1374,213,1885]" targetPageId="7">
<paragraph id="8B95791BB634A910588BF81BFA4AE0C6" blockId="7.[151,1436,1940,1993]" pageId="7" pageNumber="381">
<emphasis id="B95EA509B634A910588BF81BFECEE0A5" bold="true" box="[151,280,1941,1963]" pageId="7" pageNumber="381">FIGURE 4.</emphasis>
<taxonomicName id="4C2A0298B634A9105938F81AFBDBE0A5" authority="Bulanova-Zachvatkina, 1967" authorityName="Bulanova-Zachvatkina" authorityYear="1967" box="[292,1037,1940,1962]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="pseudopapillipes" subGenus="Neobelba">
<emphasis id="B95EA509B634A9105938F81AFE46E0A5" box="[292,400,1940,1962]" italics="true" pageId="7" pageNumber="381">Metabelba</emphasis>
(
<emphasis id="B95EA509B634A91059B9F81AFDD0E0A5" box="[421,518,1940,1962]" italics="true" pageId="7" pageNumber="381">Neobelba</emphasis>
) 
<emphasis id="B95EA509B634A9105A07F81AFD11E0A5" box="[539,711,1940,1962]" italics="true" pageId="7" pageNumber="381">pseudopapillipes</emphasis>
(Bulanova-Zachvatkina, 1967)
</taxonomicName>
—legs II and III (drawings of V. B. Kolesnikov). A—leg II (segments individually depicted), B—leg III (segments individually depicted). Bar representing 50 µm.
</paragraph>
</caption>
<paragraph id="8B95791BB63BA91F58DBFF17FAB2E5C0" blockId="8.[151,1437,151,2034]" pageId="8" pageNumber="382">
<emphasis id="B95EA509B63BA91F58DBFF17FF2AE7BF" box="[199,252,153,176]" italics="true" pageId="8" pageNumber="382">Legs</emphasis>
. 
<figureCitation id="1311659EB63BA91F5910FF19FE83E7BF" box="[268,341,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,250,1945,1967]" captionTargetBox="[186,1401,204,1905]" captionTargetId="figure@6.[151,1436,193,1924]" captionTargetPageId="6" captionText="FIGURE 3. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — legs. A—leg I, B—femur and genu II, C—femur and genu III, D—leg IV, E—enlarged detail of solenidion and coupled seta d of tibia IV. Bar representing 100 µm (A – D)." httpUri="https://zenodo.org/record/229456/files/figure.png" pageId="8" pageNumber="382">Figs 3</figureCitation>
, 
<figureCitation id="1311659EB63BA91F5978FF16FEA3E7BF" box="[356,373,152,176]" captionStart="FIGURE 4" captionStartId="7.[151,250,1941,1963]" captionTargetBox="[224,1374,213,1885]" captionTargetId="figure@7.[195,1385,193,1907]" captionTargetPageId="7" captionText="FIGURE 4. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — legs II and III (drawings of V. B. Kolesnikov). A—leg II (segments individually depicted), B—leg III (segments individually depicted). Bar representing 50 µm." httpUri="https://zenodo.org/record/229457/files/figure.png" pageId="8" pageNumber="382">4</figureCitation>
. Legs monodactylous, relatively short (leg I shorter than body, leg IV about as long as body length, slightly longer than ventral length, other legs also shorter than body, and overall length of legs increasing from leg I to leg IV—see 
<tableCitation id="C6A84CA0B63BA91F59DBFF51FDCDE7F7" box="[455,539,223,248]" captionStart="TABLE 1" captionStartId="3.[151,239,296,319]" captionText="TABLE 1. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — measurements." pageId="8" pageNumber="382">Table 1</tableCitation>
). Leg segments with distinctly swollen distal parts (moniliform), except tarsi, where swollen part is proximal. Leg setae strong, setiform to spiniform, short to medium long, always shorter than their respective segment, mostly slightly bent, often with barbs or small spines at their outer curvatures. Dorsal setae of trochanters, femora, genua, and seta 
<emphasis id="B95EA509B63BA91F5A84FEC2FD61E66B" box="[664,695,332,356]" italics="true" pageId="8" pageNumber="382">ft”</emphasis>
of all tarsi very strong, robust, strongly and almost rectangularly bent proximally, mostly with long distinct spines at external curvature. Lateral setae of genua and tibiae usually similarly developed, but slightly smaller, other setae more narrow, or almost simply setiform, more straight and usually shorter. Leg setal formula except of seta 
<emphasis id="B95EA509B63BA91F5AAFFE36FD17E6DF" box="[691,705,440,464]" italics="true" pageId="8" pageNumber="382">d</emphasis>
on tibia IV identical to species of 
<taxonomicName id="4C2A0298B63BA91F5C23FE36FB61E6DF" box="[1087,1207,440,464]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63BA91F5C23FE36FB61E6DF" box="[1087,1207,440,464]" italics="true" pageId="8" pageNumber="382">Metabelba</emphasis>
</taxonomicName>
, as follows: leg I: 
<date id="FF945FDBB63BA91F5D99FE36FF18E6FB" pageId="8" pageNumber="382" value="2010-01-04">1- 10-4</date>
[1]-4(2)-20(2); leg II: 
<date id="FF945FDBB63BA91F59DFFE52FDC4E6FA" box="[451,530,476,501]" pageId="8" pageNumber="382" value="2010-01-04">1-10-4</date>
[1]-5[1]-17(2); leg III: 4-9-4[1]-5[1]-17(0); leg IV: 3-9-4-5[1]-14(0). Famulus (e) normal, setiform, emergent, relatively long. Solenidia of genu I, II and III each coupled with companion seta 
<emphasis id="B95EA509B63BA91F5D74FD8EFAA0E517" box="[1384,1398,512,536]" italics="true" pageId="8" pageNumber="382">d</emphasis>
, as well as solenidia of tibia II, III and IV. Length of solenidia on genua subequal to length of companion seta, tibial solenidia II–III slightly longer than respective companion setae, tibial solenidion IV about twice as long as companion seta or even slightly longer (
<figureCitation id="1311659EB63BA91F5A78FDE2FD71E58B" box="[612,679,620,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1945,1967]" captionTargetBox="[186,1401,204,1905]" captionTargetId="figure@6.[151,1436,193,1924]" captionTargetPageId="6" captionText="FIGURE 3. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — legs. A—leg I, B—femur and genu II, C—femur and genu III, D—leg IV, E—enlarged detail of solenidion and coupled seta d of tibia IV. Bar representing 100 µm (A – D)." httpUri="https://zenodo.org/record/229456/files/figure.png" pageId="8" pageNumber="382">Fig. 3</figureCitation>
D, E). Solenidia of tibia I free as typical for 
<taxonomicName id="4C2A0298B63BA91F5CB6FDE2FAFDE58A" box="[1194,1323,620,645]" class="Arachnida" family="Damaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="family">Damaeidae</taxonomicName>
, setiform, tactile, 
<emphasis id="B95EA509B63BA91F58F2FD1FFEDFE5A3" box="[238,265,657,684]" italics="true" pageId="8" pageNumber="382">φ1</emphasis>
very long, about 2,8 times longer than 
<emphasis id="B95EA509B63BA91F5AF9FD1FFCD6E5A3" box="[741,768,657,684]" italics="true" pageId="8" pageNumber="382">φ2</emphasis>
. Solenidia of tarsus I relatively short, setiform, pointed, solenidion 
<emphasis id="B95EA509B63BA91F590EFD39FEF8E5DD" box="[274,302,695,722]" italics="true" pageId="8" pageNumber="382">ω1</emphasis>
about twice as long as 
<emphasis id="B95EA509B63BA91F5A2FFD39FD98E5DD" box="[563,590,695,722]" italics="true" pageId="8" pageNumber="382">ω2</emphasis>
, distally bent. Solenidia of tarsus II straight, blunt, subequal in length.
</paragraph>
<paragraph id="8B95791BB63BA91F58DBFD52FE68E5FA" blockId="8.[151,1437,151,2034]" box="[199,446,732,757]" pageId="8" pageNumber="382">
<emphasis id="B95EA509B63BA91F58DBFD52FEECE5FA" bold="true" box="[199,314,732,757]" pageId="8" pageNumber="382">Ontogeny</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B95791BB63BA91F58DBFC8EFE27E3C6" blockId="8.[151,1437,151,2034]" pageId="8" pageNumber="382">
<emphasis id="B95EA509B63BA91F58DBFC8EFD76E416" bold="true" box="[199,672,768,793]" pageId="8" pageNumber="382">Geographical distribution and ecology.</emphasis>
<taxonomicName id="4C2A0298B63BA91F5AB5FC8FFC5DE417" box="[681,907,768,792]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63BA91F5AB5FC8FFC5DE417" box="[681,907,768,792]" italics="true" pageId="8" pageNumber="382">N. pseudopapillipes</emphasis>
</taxonomicName>
is recorded only very rarely. Except classical localities of 
<bibRefCitation id="EFBB04EAB63BA91F5931FCAAFD55E432" author="Bulanova-Zachvatkina" box="[301,643,804,829]" pageId="8" pageNumber="382" refString="Bulanova-Zachvatkina, E. M. (1967) Pancirnyje kleschtschi - Oribatidy. Vysshaya Shkola, Moscow, 254 pp. [in Russian]" type="book" year="1967">Bulanova-Zachvatkina (1967)</bibRefCitation>
in 
<collectingCountry id="F33D398BB63BA91F5AAEFCAAFCC1E432" box="[690,791,804,829]" name="Moldova" pageId="8" pageNumber="382">Moldova</collectingCountry>
and Central 
<collectingCountry id="F33D398BB63BA91F5BA4FCAAFBD3E432" box="[952,1029,804,829]" name="Russia" pageId="8" pageNumber="382">Russia</collectingCountry>
, and two localities presented here (
<collectingCountry id="F33D398BB63BA91F5882FCC6FED4E46E" box="[158,258,840,865]" name="Slovakia" pageId="8" pageNumber="382">Slovakia</collectingCountry>
and Russia—Voronezh region), we are aware of only two other potential records, both from Italy—one from Alps in province of Trentino (
<bibRefCitation id="EFBB04EAB63BA91F5A36FCE2FCD2E48A" author="Baratti" box="[554,772,876,901]" pageId="8" pageNumber="382" refString="Baratti, M., Migliorini, M. &amp; Bernini, F. (2000) Effetti dell'innevamento artificiale sugli Acari Oribatei (Acari, Oribatida) delle piste sciabili del Monte Bondone (Trentino, Italia). Studi Trentini di Scienze Naturali - Acta Biologica, 75, 147 - 159." type="journal article" year="2000">
Baratti 
<emphasis id="B95EA509B63BA91F5A9CFCE3FD6FE48B" box="[640,697,876,900]" italics="true" pageId="8" pageNumber="382">et al.</emphasis>
, 2000
</bibRefCitation>
), from ski slopes in elevation of about 
<quantity id="4CD2D4FEB63BA91F5CDDFCE2FA8FE48B" box="[1217,1369,876,901]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="8" pageNumber="382" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">1600-1700 m</quantity>
a.s.l., and another one from beech and oak forests in southern Tuscany, from elevation between 300 and 
<quantity id="4CD2D4FEB63BA91F5D08FC1FFA88E4A7" box="[1300,1374,913,937]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="382" unit="m" value="800.0">800 m</quantity>
a.s.l. (
<bibRefCitation id="EFBB04EAB63BA91F5882FC3AFE75E4C2" author="Migliorini" box="[158,419,948,973]" pageId="8" pageNumber="382" refString="Migliorini, M., Petrioli, A. &amp; Bernini, F. (2002) Comparative analysis of two edaphic zoocenoses (Oribatid mites and Carabid beetles) in five habitats of the &quot; Pietraporciana &quot; and &quot; Lucciolabella &quot; Nature Reserves (Orcia Valley, central Italy). Acta Oecologica, 23, 361 - 374. http: // dx. doi. org / 10.1016 / s 1146 - 609 x (02) 01163 - 3" type="journal article" year="2002">
Migliorini 
<emphasis id="B95EA509B63BA91F5900FC3BFE80E4C3" box="[284,342,948,972]" italics="true" pageId="8" pageNumber="382">et al.</emphasis>
, 2002
</bibRefCitation>
). However, in both cases the species found was reported as “ 
<taxonomicName id="4C2A0298B63BA91F5C7DFC3BFAB1E4C3" box="[1121,1383,948,973]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63BA91F5C7DFC3BFBADE4C3" box="[1121,1147,949,972]" italics="true" pageId="8" pageNumber="382">N.</emphasis>
cf. 
<emphasis id="B95EA509B63BA91F5CB5FC3AFAB1E4C3" box="[1193,1383,948,972]" italics="true" pageId="8" pageNumber="382">pseudopapillipes</emphasis>
</taxonomicName>
”, so there is no definitive clarity, if the species indeed belonged to 
<taxonomicName id="4C2A0298B63BA91F5B7CFC57FB96E4FF" box="[864,1088,984,1008]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63BA91F5B7CFC57FB96E4FF" box="[864,1088,984,1008]" italics="true" pageId="8" pageNumber="382">N. pseudopapillipes</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFBB04EAB63BA91F5C4CFC56FB31E4FE" author="Subias" box="[1104,1255,984,1009]" pageId="8" pageNumber="382" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1748 - 2002). Graellsia 60, 3 - 305." type="journal article" year="2004">Subías (2004</bibRefCitation>
, 
<bibRefCitation id="EFBB04EAB63BA91F5CE8FC57FAF8E4FE" author="Subias" box="[1268,1326,985,1009]" pageId="8" pageNumber="382" refString="Subias, L. (2013) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (Excepto fosiles). 540 pp (on-line version). Available from: http: // www. ucm. es / info / zoo / Artropodos / Catalogo. pdf (accessed 24 October 2014)" type="book" year="2013">2013</bibRefCitation>
) reported this species from “Europa Meridional”, but it remains unclear to us on which basis. Looking on confirmed localities of the species, it may be expected in lowland, alluvial and midland forests, dominated by 
<taxonomicName id="4C2A0298B63BA91F5D11FBAFFABDE337" box="[1293,1387,1057,1080]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="382" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EA509B63BA91F5D11FBAFFABDE337" box="[1293,1387,1057,1080]" italics="true" pageId="8" pageNumber="382">Quercus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A0298B63BA91F588BFBCBFF29E353" box="[151,255,1093,1116]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="382" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EA509B63BA91F588BFBCBFF29E353" box="[151,255,1093,1116]" italics="true" pageId="8" pageNumber="382">Carpinus</emphasis>
</taxonomicName>
, even if it cannot be excluded that the species occurs also in other 
<typeStatus id="5491C7B9B63BA91F5C06FBCBFB9CE352" box="[1050,1098,1093,1117]" pageId="8" pageNumber="382">type</typeStatus>
of litter of the forests in the (?southern) temperate zone and also in forest-steppic habitats in the south-eastern Europe. The species may be therefore expected, besides known localities in 
<collectingCountry id="F33D398BB63BA91F5AD1FB02FCCCE3AA" box="[717,794,1164,1189]" name="Russia" pageId="8" pageNumber="382">Russia</collectingCountry>
, 
<collectingCountry id="F33D398BB63BA91F5B37FB02FC46E3AA" box="[811,912,1164,1189]" name="Moldova" pageId="8" pageNumber="382">Moldova</collectingCountry>
and 
<collectingCountry id="F33D398BB63BA91F5BCEFB02FBE1E3AA" box="[978,1079,1164,1189]" name="Slovakia" pageId="8" pageNumber="382">Slovakia</collectingCountry>
, also in 
<collectingCountry id="F33D398BB63BA91F5CB8FB02FAD7E3AA" box="[1188,1281,1164,1189]" name="Ukraine" pageId="8" pageNumber="382">Ukraine</collectingCountry>
, 
<collectingCountry id="F33D398BB63BA91F5D0FFB02FA4FE3AA" box="[1299,1433,1164,1189]" name="Kazakhstan" pageId="8" pageNumber="382">Kazakhstan</collectingCountry>
, 
<collectingCountry id="F33D398BB63BA91F588BFB3EFF2AE3C6" box="[151,252,1200,1225]" name="Romania" pageId="8" pageNumber="382">Romania</collectingCountry>
and 
<collectingCountry id="F33D398BB63BA91F592EFB3FFE42E3C6" box="[306,404,1201,1225]" name="Hungary" pageId="8" pageNumber="382">Hungary</collectingCountry>
at least.
</paragraph>
</subSubSection>
<subSubSection id="C3302A90B63BA91E58DBFB5AFBA9E2CD" lastPageId="9" lastPageNumber="383" pageId="8" pageNumber="382" type="discussion">
<paragraph id="8B95791BB63BA91F58DBFB5AFCE0E2E1" blockId="8.[151,1437,151,2034]" pageId="8" pageNumber="382">
<emphasis id="B95EA509B63BA91F58DBFB5AFD69E3E2" bold="true" box="[199,703,1236,1261]" pageId="8" pageNumber="382">Remarks on species and its generic status.</emphasis>
<taxonomicName id="4C2A0298B63BA91F5ADAFB5BFC73E3E3" box="[710,933,1236,1260]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63BA91F5ADAFB5BFC73E3E3" box="[710,933,1236,1260]" italics="true" pageId="8" pageNumber="382">N. pseudopapillipes</emphasis>
</taxonomicName>
as re-described here fits very well to known characters of the species, as defined by 
<bibRefCitation id="EFBB04EAB63BA91F5A7BFB76FC62E21E" author="Bulanova-Zachvatkina" box="[615,948,1272,1297]" pageId="8" pageNumber="382" refString="Bulanova-Zachvatkina, E. M. (1967) Pancirnyje kleschtschi - Oribatidy. Vysshaya Shkola, Moscow, 254 pp. [in Russian]" type="book" year="1967">Bulanova-Zachvatkina (1967</bibRefCitation>
, 
<bibRefCitation id="EFBB04EAB63BA91F5BDEFB76FC2DE21E" author="Bulanova-Zachvatkina" box="[962,1019,1272,1297]" pageId="8" pageNumber="382" refString="Bulanova-Zachvatkina, E. M. (1975) Nadsemejstvo Belboidea Dubinin, 1954 (= Damaeoidea Balogh, 1961). In: Gilijarov, M. S. &amp; Krivolutskij, D. A. (Eds.), Opredelitel obitaiushtschich v potschve kleschtschei. Nauka, Moskva, pp. 120 - 143 [in Russian]" type="book chapter" year="1975">1975</bibRefCitation>
) and reviewed above. Nevertheless, few differences appeared and raised the question of the identity of our material with original 
<taxonomicName id="4C2A0298B63BA91F5CB9FA92FAB5E23B" box="[1189,1379,1308,1332]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63BA91F5CB9FA92FAB5E23B" box="[1189,1379,1308,1332]" italics="true" pageId="8" pageNumber="382">pseudopapillipes</emphasis>
</taxonomicName>
. The observed discrepancies relate to the sejugal enantiophyses. While in original description there is recorded unspecified presence of paired tubercles in dorsosejugal area, and no ventral tubercles mentioned, in updated version 
<bibRefCitation id="EFBB04EAB63BA91F58EFFA06FD93E2AE" author="Bulanova-Zachvatkina" box="[243,581,1416,1441]" pageId="8" pageNumber="382" refString="Bulanova-Zachvatkina, E. M. (1975) Nadsemejstvo Belboidea Dubinin, 1954 (= Damaeoidea Balogh, 1961). In: Gilijarov, M. S. &amp; Krivolutskij, D. A. (Eds.), Opredelitel obitaiushtschich v potschve kleschtschei. Nauka, Moskva, pp. 120 - 143 [in Russian]" type="book chapter" year="1975">Bulanova-Zachvatkina (1975)</bibRefCitation>
states, that dorsal tubercles should be 
<emphasis id="B95EA509B63BA91F5C18FA07FBC8E2AA" box="[1028,1054,1417,1445]" italics="true" pageId="8" pageNumber="382">B1</emphasis>
and 
<emphasis id="B95EA509B63BA91F5C44FA07FBA4E2AA" box="[1112,1138,1417,1445]" italics="true" pageId="8" pageNumber="382">B2</emphasis>
, i.e anterior and posterior postbothridial tubercle (
<emphasis id="B95EA509B63BA91F59AFFA3EFE1AE2C3" box="[435,460,1456,1484]" italics="true" pageId="8" pageNumber="382">B a</emphasis>
and 
<emphasis id="B95EA509B63BA91F5A0DFA3EFDFCE2C3" box="[529,554,1456,1484]" italics="true" pageId="8" pageNumber="382">B p</emphasis>
as they are named today). In the same place, the presence of posterior ventrosejugal tubercle 
<date id="FF945FDBB63BA91F598AFA59FE79E2FC" box="[406,431,1495,1523]" pageId="8" pageNumber="382">
<emphasis id="B95EA509B63BA91F598AFA59FE79E2FC" box="[406,431,1495,1523]" italics="true" pageId="8" pageNumber="382">V2</emphasis>
</date>
is recorded (
<emphasis id="B95EA509B63BA91F5A5EFA59FD8DE2FC" box="[578,603,1495,1523]" italics="true" pageId="8" pageNumber="382">V p</emphasis>
in recent notation).
</paragraph>
<paragraph id="8B95791BB63BA91F58DBFA70FE69E08C" blockId="8.[151,1437,151,2034]" pageId="8" pageNumber="382">
In present work, we consider (consistently with situation in several 
<taxonomicName id="4C2A0298B63BA91F5BF9FA73FB8BE11A" box="[997,1117,1533,1557]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63BA91F5BF9FA73FB8BE11A" box="[997,1117,1533,1557]" italics="true" pageId="8" pageNumber="382">Metabelba</emphasis>
</taxonomicName>
species) the present dorsal tubercles to be 
<emphasis id="B95EA509B63BA91F5955F9AFFEB3E132" box="[329,357,1569,1597]" italics="true" pageId="8" pageNumber="382">D a</emphasis>
and 
<emphasis id="B95EA509B63BA91F59BCF9AFFE6AE132" box="[416,444,1569,1597]" italics="true" pageId="8" pageNumber="382">D p</emphasis>
, based on their rather central position axial to the level of 
<emphasis id="B95EA509B63BA91F5C74F9AFFBA8E137" box="[1128,1150,1569,1592]" italics="true" pageId="8" pageNumber="382">in</emphasis>
insertions. Nevertheless, homology of these tubercles remains unclear, as they may be more linked to the development of latero 
<emphasis id="B95EA509B63BA91F5D36F9C6FAEEE150" box="[1322,1336,1608,1631]" italics="true" pageId="8" pageNumber="382">–</emphasis>
posterior rather than postero-axial end of bothridial cuticular protuberance. This is supported by the argumentation of Behan 
<bibRefCitation id="EFBB04EAB63BA91F588BF901FE04E1A7" author="Pelletier" box="[151,466,1679,1704]" pageId="8" pageNumber="382" refString="Behan-Pelletier, V. M. &amp; Norton, R. A. (1985) Epidamaeus (Acari: Damaeidae) of subarctic western North America and extreme northeastern USSR. Canadian Entomologist, 117, 277 - 319. http: // dx. doi. org / 10.4039 / ent 117277 - 3" type="journal article" year="1985">Pelletier &amp; Norton (1985)</bibRefCitation>
and by the fact that protuberance on which is positioned bothridium is in 
<taxonomicName id="4C2A0298B63BA91F5D9EF91EFE83E1C3" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63BA91F5D9EF91EFE83E1C3" italics="true" pageId="8" pageNumber="382">N. pseudopapillipes</emphasis>
</taxonomicName>
connected with anterior tubercle of this enantiophysis by oblique ridge (
<figureCitation id="1311659EB63BA91F5CB9F93DFB3AE1C3" box="[1189,1260,1715,1740]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1151]" captionTargetBox="[179,1423,207,1093]" captionTargetId="figure@5.[151,1436,193,1108]" captionTargetPageId="5" captionText="FIGURE 2. Metabelba (Neobelba) pseudopapillipes (Bulanova-Zachvatkina, 1967) — lateral view, details of prodorsum and gnatosoma. A—lateral view of the body, B—prodorsum, C—chelicere, D—palp, E—notogastral setae of individuals collected in Slovakia. Bar representing 100 µm (A, E) or 50 µm (B – D). Details of chelicere (setae cha and chb) and palp (distal part of palp tarsus) further enlarged to allow observation of details. Abbreviations: apd—area prodorsal, dis—discidium, gl—notogastral gland opening. Other markings represent names of setae, tubercles or lyrifissures." httpUri="https://zenodo.org/record/229455/files/figure.png" pageId="8" pageNumber="382">Fig. 2</figureCitation>
B). Shift of the whole enantiophyse in axial direction may be also explained by narrowing of the whole body in the sejugal area. Regarding presence of ventrosejugal posterior tubercle 
<emphasis id="B95EA509B63BA91F5B1FF972FCC2E01C" box="[771,788,1788,1811]" italics="true" pageId="8" pageNumber="382">V</emphasis>
p (corresponding to 
<date id="FF945FDBB63BA91F5BECF972FBDFE017" box="[1008,1033,1788,1816]" pageId="8" pageNumber="382">
<emphasis id="B95EA509B63BA91F5BECF972FBD7E01C" box="[1008,1025,1788,1811]" italics="true" pageId="8" pageNumber="382">V</emphasis>
2
</date>
of Bulanova-Zachvatkina), we note that cuticular triangular thickenings in corresponding area may be interpreted as apophyses, when seen purely in ventral view. Nevertheless, observation from another angle (lateral) shows that there is no protuberance present, and tubercle 
<emphasis id="B95EA509B63BA91F593BF8E5FEEEE08D" box="[295,312,1899,1922]" italics="true" pageId="8" pageNumber="382">V</emphasis>
p is missing.
</paragraph>
<paragraph id="8B95791BB63BA91F58DBF81FFB60E0C1" blockId="8.[151,1437,151,2034]" pageId="8" pageNumber="382">
<collectingCountry id="F33D398BB63BA91F58DBF81FFF30E0A6" box="[199,230,1937,1961]" name="American Samoa" pageId="8" pageNumber="382">As</collectingCountry>
all other characters fit exactly the description of Bulanova-Zachvatkina, and no other similar species was found yet, we consider the identity of our individuals with 
<taxonomicName id="4C2A0298B63BA91F5B35F83BFB8CE0C2" box="[809,1114,1973,1997]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63BA91F5B35F83BFB8CE0C2" box="[809,1114,1973,1997]" italics="true" pageId="8" pageNumber="382">Neobelba pseudopapillipes</emphasis>
</taxonomicName>
proved.
</paragraph>
<paragraph id="8B95791BB63BA91E58DBF857FC78E4AB" blockId="8.[151,1437,151,2034]" lastBlockId="9.[151,1437,151,1475]" lastPageId="9" lastPageNumber="383" pageId="8" pageNumber="382">
The species shows important similarities with species of the genus 
<taxonomicName id="4C2A0298B63BA91F5BA0F857FB2FE0FD" authority="Grandjean, 1936" authorityName="Grandjean" authorityYear="1936" box="[956,1273,2009,2034]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63BA91F5BA0F857FBE2E0FE" box="[956,1076,2009,2033]" italics="true" pageId="8" pageNumber="382">Metabelba</emphasis>
Grandjean, 1936
</taxonomicName>
. Development of prodorsum and notogaster and their setae, characters of ventral part of the body as well as the legs are identical or very similar. 
<collectingCountry id="F33D398BB63AA91E5956FF32FEBFE7DB" box="[330,361,188,212]" name="American Samoa" pageId="9" pageNumber="383">As</collectingCountry>
far as we know, the following characters can be considered as unique for 
<taxonomicName id="4C2A0298B63AA91E5CB3FF32FABBE7DB" box="[1199,1389,188,212]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63AA91E5CB3FF32FABBE7DB" box="[1199,1389,188,212]" italics="true" pageId="9" pageNumber="383">pseudopapillipes</emphasis>
</taxonomicName>
: (a) presence of companion seta 
<emphasis id="B95EA509B63AA91E59C0FF6EFE3CE7F7" box="[476,490,224,248]" italics="true" pageId="9" pageNumber="383">d</emphasis>
, coupled with solenidion on tibia IV, (b) presence of distinct prodorsal area (
<emphasis id="B95EA509B63AA91E5D7EFF6EFA5AE7F7" box="[1378,1420,224,248]" italics="true" pageId="9" pageNumber="383">apd</emphasis>
), resembling area porosa, anterior to insertion of interlamellar seta 
<emphasis id="B95EA509B63AA91E5B95FE8BFC49E613" box="[905,927,261,284]" italics="true" pageId="9" pageNumber="383">in</emphasis>
, (c) characteristic form of notogastral setae, bent and thickened at the base in lateral view. Absence or presence of accompanied setae related to tibial solenidia is broadly accepted as important distinguishing character at generic or even supra-generic level; however, this mostly applies to total absence (all tibial solenidia free) than to the absence of individual companion seta. On the other hand, setation of the leg IV seems to be of greater importance than in legs II and III, and absence of companion seta on tibia IV was used as one of the distinguishing characters, 
<emphasis id="B95EA509B63AA91E5BF1FE37FBC0E6DF" box="[1005,1046,441,464]" italics="true" pageId="9" pageNumber="383">e.g.</emphasis>
, between 
<taxonomicName id="4C2A0298B63AA91E5C9BFE36FB1FE6DF" box="[1159,1225,440,464]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5C9BFE36FB1FE6DF" box="[1159,1225,440,464]" italics="true" pageId="9" pageNumber="383">Belba</emphasis>
</taxonomicName>
von Heyden, 1826 and 
<taxonomicName id="4C2A0298B63AA91E58DBFE52FE3BE6FB" authority="Norton, 1979" authorityName="Norton" authorityYear="1979" box="[199,493,476,500]" class="Arachnida" family="Damaeidae" genus="Caenobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E58DBFE52FE98E6FB" box="[199,334,476,500]" italics="true" pageId="9" pageNumber="383">Caenobelba</emphasis>
Norton, 1979
</taxonomicName>
or 
<taxonomicName id="4C2A0298B63AA91E5A08FE52FCF2E6FB" authority="Norton, 1980" authorityName="Norton" authorityYear="1980" box="[532,804,476,500]" class="Arachnida" family="Damaeidae" genus="Dyobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5A08FE52FD57E6FB" box="[532,641,476,500]" italics="true" pageId="9" pageNumber="383">Dyobelba</emphasis>
Norton, 1980
</taxonomicName>
. 
<bibRefCitation id="EFBB04EAB63AA91E5B33FE52FC1AE6FB" author="Subias" box="[815,972,476,501]" pageId="9" pageNumber="383" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1748 - 2002). Graellsia 60, 3 - 305." type="journal article" year="2004">Subías (2004)</bibRefCitation>
placed genus 
<taxonomicName id="4C2A0298B63AA91E5C73FE52FB0CE6FB" box="[1135,1242,476,500]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5C73FE52FB0CE6FB" box="[1135,1242,476,500]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
as a subgenus to genus 
<taxonomicName id="4C2A0298B63AA91E58F9FD8EFE8BE517" box="[229,349,512,536]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E58F9FD8EFE8BE517" box="[229,349,512,536]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
, and later (
<bibRefCitation id="EFBB04EAB63AA91E59F2FE71FD5EE517" author="Subias" box="[494,648,511,536]" pageId="9" pageNumber="383" refString="Subias, L. (2013) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (Excepto fosiles). 540 pp (on-line version). Available from: http: // www. ucm. es / info / zoo / Artropodos / Catalogo. pdf (accessed 24 October 2014)" type="book" year="2013">Subías, 2013</bibRefCitation>
) even synonymized it with 
<taxonomicName id="4C2A0298B63AA91E5BC2FD8EFB80E517" box="[990,1110,512,536]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B95EA509B63AA91E5BC2FD8EFB80E517" box="[990,1110,512,536]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D1872B63AA91E5C7FFD8EFB72E518" box="[1123,1188,512,535]" pageId="9" pageNumber="383" sensu="stricto">s. str.</taxonomicNameLabel>
We assume that the motivation for this move was in great morphological similarity of 
<taxonomicName id="4C2A0298B63AA91E5B8BFDABFBAEE533" box="[919,1144,548,572]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63AA91E5B8BFDABFBAEE533" box="[919,1144,548,572]" italics="true" pageId="9" pageNumber="383">N. pseudopapillipes</emphasis>
</taxonomicName>
and species of the genus 
<taxonomicName id="4C2A0298B63AA91E588BFDC6FED9E56F" box="[151,271,584,608]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E588BFDC6FED9E56F" box="[151,271,584,608]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
. Mourek 
<emphasis id="B95EA509B63AA91E5991FDC7FE1BE56F" box="[397,461,584,608]" italics="true" pageId="9" pageNumber="383">et al.</emphasis>
(2012) considered 
<taxonomicName id="4C2A0298B63AA91E5AA3FDC6FCFCE56F" box="[703,810,584,608]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5AA3FDC6FCFCE56F" box="[703,810,584,608]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
as a separate genus, until more information about morphology of this taxon will be available, and mentioned namely the chaetotaxy of leg femora. To decide about the status of 
<taxonomicName id="4C2A0298B63AA91E593BFD1EFE44E5A7" box="[295,402,656,680]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E593BFD1EFE44E5A7" box="[295,402,656,680]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
it is crucial to weight the value of presence vs. loss of seta 
<emphasis id="B95EA509B63AA91E5C28FD1EFB94E5A7" box="[1076,1090,656,680]" italics="true" pageId="9" pageNumber="383">d</emphasis>
on tibia IV, accompanying the respective solenidion. 
<collectingCountry id="F33D398BB63AA91E5984FD3AFE61E5C3" box="[408,439,692,716]" name="American Samoa" pageId="9" pageNumber="383">As</collectingCountry>
demonstrated above, this character can be used as distinguishing between the genera. This supports the idea that presence or loss of seta 
<emphasis id="B95EA509B63AA91E5AC8FD56FD34E5FF" box="[724,738,728,752]" italics="true" pageId="9" pageNumber="383">d</emphasis>
on tibia IV is rather a generic than a specific character within 
<taxonomicName id="4C2A0298B63AA91E588BFD72FECFE41A" box="[151,281,764,789]" class="Arachnida" family="Damaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="family">Damaeidae</taxonomicName>
. In all cases where it differentiates between the genera, however, this character is combined with some additional distinguishing characters. Two other distinguishing characters of 
<taxonomicName id="4C2A0298B63AA91E5BEFFCAFFB04E437" box="[1011,1234,800,824]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63AA91E5BEFFCAFFB04E437" box="[1011,1234,800,824]" italics="true" pageId="9" pageNumber="383">N. pseudopapillipes</emphasis>
</taxonomicName>
mentioned above can hardly be seen as generic: variability of notogastral setae within the genus 
<taxonomicName id="4C2A0298B63AA91E5C0DFCCAFB5FE453" box="[1041,1161,836,860]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5C0DFCCAFB5FE453" box="[1041,1161,836,860]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
is very broad, and areas similar to prodorsal area 
<emphasis id="B95EA509B63AA91E59A4FCE6FE34E48F" box="[440,482,872,896]" italics="true" pageId="9" pageNumber="383">apd</emphasis>
appear on different places and different species throughout the family. This leaves only single generic character to distinguish 
<taxonomicName id="4C2A0298B63AA91E5A63FC02FD3DE4AB" box="[639,747,908,932]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5A63FC02FD3DE4AB" box="[639,747,908,932]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
from 
<emphasis id="B95EA509B63AA91E5B33FC02FC78E4AB" box="[815,942,908,932]" italics="true" pageId="9" pageNumber="383">
<taxonomicName id="4C2A0298B63AA91E5B33FC02FC7DE4AB" box="[815,939,908,932]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">Metabelba</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B95791BB63AA91E58DBFC3EFB01E3AF" blockId="9.[151,1437,151,1475]" pageId="9" pageNumber="383">
In our view, the only solution of the problem is in placing 
<taxonomicName id="4C2A0298B63AA91E5B9CFC3EFC3DE4C7" box="[896,1003,944,968]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5B9CFC3EFC3DE4C7" box="[896,1003,944,968]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
to broader 
<taxonomicName id="4C2A0298B63AA91E5C6AFC3EFB38E4C7" box="[1142,1262,944,968]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5C6AFC3EFB38E4C7" box="[1142,1262,944,968]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
concept at the subgeneric level. The proposed synonymy of 
<taxonomicName id="4C2A0298B63AA91E5A87FC5AFCD0E4E3" box="[667,774,980,1004]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5A87FC5AFCD0E4E3" box="[667,774,980,1004]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A0298B63AA91E5B23FC5AFC61E4E3" box="[831,951,980,1004]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B95EA509B63AA91E5B23FC5AFC61E4E3" box="[831,951,980,1004]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D1872B63AA91E5BA2FC5BFC2CE4E3" box="[958,1018,981,1004]" pageId="9" pageNumber="383" sensu="stricto">s. str.</taxonomicNameLabel>
must be rejected. Nominal subgenus of 
<taxonomicName id="4C2A0298B63AA91E58AAFC76FEF8E31F" box="[182,302,1016,1040]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E58AAFC76FEF8E31F" box="[182,302,1016,1040]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
has well developed propodolateral apophysis 
<emphasis id="B95EA509B63AA91E5B20FC77FC9CE31F" box="[828,842,1017,1040]" italics="true" pageId="9" pageNumber="383">P</emphasis>
, which is clearly missing in 
<taxonomicName id="4C2A0298B63AA91E5C8FFC76FB28E31F" box="[1171,1278,1016,1040]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5C8FFC76FB28E31F" box="[1171,1278,1016,1040]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
. On the other hand, 
<taxonomicName id="4C2A0298B63AA91E58C7FB93FE6FE33B" box="[219,441,1052,1076]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63AA91E58C7FB93FE6FE33B" box="[219,441,1052,1076]" italics="true" pageId="9" pageNumber="383">N. pseudopapillipes</emphasis>
</taxonomicName>
closely resembles some of the species of recently defined subgenus 
<emphasis id="B95EA509B63AA91E5CAEFB92FAE0E33B" box="[1202,1334,1052,1076]" italics="true" pageId="9" pageNumber="383">Pateribelba</emphasis>
Mourek, Miko &amp; Bernini, 2012. It has to be noted that notation of the prodorsal tubercles in Mourek 
<emphasis id="B95EA509B63AA91E5CBAFBCFFB09E357" box="[1190,1247,1088,1112]" italics="true" pageId="9" pageNumber="383">et al.</emphasis>
(2012) followed the same approach of designation based on the physical position of tubercles (axially to the level of setae 
<emphasis id="B95EA509B63AA91E5D9AFBEBFA4AE373" box="[1414,1436,1125,1148]" italics="true" pageId="9" pageNumber="383">in</emphasis>
insertions), the discussion above on homology of tubercles observed in 
<taxonomicName id="4C2A0298B63AA91E5BA4FB06FBF2E3AF" box="[952,1060,1160,1184]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5BA4FB06FBF2E3AF" box="[952,1060,1160,1184]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
applies as well.
</paragraph>
<paragraph id="8B95791BB63AA91E58DBFB22FBA9E2CD" blockId="9.[151,1437,151,1475]" pageId="9" pageNumber="383">
Particularly interesting is the common presence of such subtle characters like the presence of 2 pairs of light spots on the notogaster near the insertions of 
<emphasis id="B95EA509B63AA91E5A91FB5EFD4BE3E3" box="[653,669,1232,1260]" italics="true" pageId="9" pageNumber="383">l a</emphasis>
and 
<emphasis id="B95EA509B63AA91E5ACEFB5EFD37E3E3" box="[722,737,1232,1260]" italics="true" pageId="9" pageNumber="383">l p</emphasis>
in most of 
<emphasis id="B95EA509B63AA91E5B42FB5EFC34E3E7" box="[862,994,1232,1256]" italics="true" pageId="9" pageNumber="383">Pateribelba</emphasis>
species studied and in 
<taxonomicName id="4C2A0298B63AA91E5CFEFB5EFA9BE3E7" box="[1250,1357,1232,1256]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5CFEFB5EFA9BE3E7" box="[1250,1357,1232,1256]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
; or the form and size of setae on leg segments (
<emphasis id="B95EA509B63AA91E5A78FB76FD5AE200" box="[612,652,1272,1295]" italics="true" pageId="9" pageNumber="383">e.g.</emphasis>
dorsal setae on trochanters, femora and genua or length and form of solenidia except solenidion of tibia IV) very similar 
<emphasis id="B95EA509B63AA91E5B36FA95FC84E23D" box="[810,850,1307,1330]" italics="true" pageId="9" pageNumber="383">e.g.</emphasis>
in both 
<taxonomicName id="4C2A0298B63AA91E5BD7FA95FAECE23C" authority="Strenzke, 1950" authorityName="Strenzke" authorityYear="1950" box="[971,1338,1306,1331]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="sphagni" subGenus="P.">
<emphasis id="B95EA509B63AA91E5BD7FA95FBAFE23C" box="[971,1145,1307,1331]" italics="true" pageId="9" pageNumber="383">M.(P.) sphagni</emphasis>
Strenzke, 1950
</taxonomicName>
and 
<taxonomicName id="4C2A0298B63AA91E5D9EFA95FE83E258" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="pseudopapillipes">
<emphasis id="B95EA509B63AA91E5D9EFA95FE83E258" italics="true" pageId="9" pageNumber="383">N. pseudopapillipes</emphasis>
</taxonomicName>
. These common characters further support inclusion of 
<taxonomicName id="4C2A0298B63AA91E5BF1FAB1FB8EE258" box="[1005,1112,1343,1367]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5BF1FAB1FB8EE258" box="[1005,1112,1343,1367]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
into 
<taxonomicName id="4C2A0298B63AA91E5C87FAB1FAC5E258" box="[1179,1299,1343,1367]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5C87FAB1FAC5E258" box="[1179,1299,1343,1367]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
. 
<collectingCountry id="F33D398BB63AA91E5D39FAB1FA92E258" box="[1317,1348,1343,1367]" name="American Samoa" pageId="9" pageNumber="383">As</collectingCountry>
argued above, we consider the presence of companion seta on tibia IV to be a character at generic level, and therefore reject the synonymy with 
<emphasis id="B95EA509B63AA91E59A7FA09FDE9E290" box="[443,575,1415,1439]" italics="true" pageId="9" pageNumber="383">Pateribelba</emphasis>
and support the original proposal of 
<bibRefCitation id="EFBB04EAB63AA91E5BC3FA08FBA8E290" author="Subias" box="[991,1150,1414,1439]" pageId="9" pageNumber="383" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1748 - 2002). Graellsia 60, 3 - 305." type="journal article" year="2004">Subías (2004)</bibRefCitation>
, i.e. to place 
<taxonomicName id="4C2A0298B63AA91E5D0EFA09FAA8E290" box="[1298,1406,1415,1439]" class="Arachnida" family="Damaeidae" genus="Neobelba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5D0EFA09FAA8E290" box="[1298,1406,1415,1439]" italics="true" pageId="9" pageNumber="383">Neobelba</emphasis>
</taxonomicName>
as a separate subgenus of the genus 
<taxonomicName id="4C2A0298B63AA91E5A10FA25FD52E2CC" box="[524,644,1451,1475]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA509B63AA91E5A10FA25FD52E2CC" box="[524,644,1451,1475]" italics="true" pageId="9" pageNumber="383">Metabelba</emphasis>
</taxonomicName>
, with close relation to subgenus 
<emphasis id="B95EA509B63AA91E5BE8FA25FBAEE2CC" box="[1012,1144,1451,1475]" italics="true" pageId="9" pageNumber="383">Pateribelba</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>