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<mods:title id="25828BC248B96A8EA00C2D4FEFBE16CE">A taxonomic synopsis of Chrysomydas Wilcox, Papavero &amp; Pimentel (Diptera: Mydidae: Mydinae), with first bionomic records for the genus and description of a new species from Brazil</mods:title>
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<taxonomicName id="4C2A4D6AF22FFFD92BF1D66F0A748229" authority="(Olivier, 1811)" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[151,613,1882,1909]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="104" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF22FFFD92BF1D66F0A748229" bold="true" box="[151,613,1882,1909]" pageId="1" pageNumber="104">
<emphasis id="B95EEAFBF22FFFD92BF1D66F09BC8228" bold="true" box="[151,429,1882,1909]" italics="true" pageId="1" pageNumber="104">Chrysomydas nitidulus</emphasis>
(
<bibRefCitation id="EFBB4B18F22FFFD92ADAD66F0A4C8229" author="Olivier, G. A." box="[444,605,1882,1908]" pageId="1" pageNumber="104" pagination="1 - 360" refId="ref7573" refString="Olivier, G. A. (1811) Insectes [i. e., Arthropoda], in Societe de Gens de Lettres, de Savans et d'Artistes, Encyclopedie methodique. Histoire Naturelle 8 (5), 1 - 360." type="journal article" year="1811">Olivier, 1811</bibRefCitation>
)
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<subSubSection id="C3306562F22FFFD92BF1D64B09F582C4" box="[151,484,1918,1945]" pageId="1" pageNumber="104" type="description">
<paragraph id="8B9536E9F22FFFD92BF1D64B09F582C4" blockId="1.[151,613,1882,1945]" box="[151,484,1918,1945]" pageId="1" pageNumber="104">
(
<figureCitation id="13112A6CF22FFFD92BF9D64B08F082C5" box="[159,225,1918,1944]" captionStart="FIGURE 1" captionStartId="3.[151,250,1972,1997]" captionTargetBox="[161,1409,166,1953]" captionTargetId="figure@3.[161,1426,166,1953]" captionTargetPageId="3" captionText="FIGURE 1. Chrysomydas nitidulus. AD: male (UFMG). A. Dorsal habitus; B. Lateral habitus; C. Head, frontal view; D. wing; EH. female (INPA); E. Dorsal habitus; F. Lateral habitus; G. Head, lateral view." httpUri="https://zenodo.org/record/3383564/files/figure.png" pageId="1" pageNumber="104">Figs 1</figureCitation>
AH, 3AG, 4AE, 6)
</paragraph>
</subSubSection>
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<taxonomicName id="4C2A4D6AF22FFFD92BF1D6F709CF8286" authority="Lichtenstein, 1796" authorityName="Lichtenstein" authorityYear="1796" box="[151,478,1986,2011]" class="Insecta" family="Mydidae" genus="Midas" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="104" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="B95EEAFBF22FFFD92BF1D6F709098287" box="[151,280,1986,2010]" italics="true" pageId="1" pageNumber="104">Midas nitida</emphasis>
<bibRefCitation id="EFBB4B18F22FFFD92A78D6F709CF8286" author="Lichtenstein, A. A. H." box="[286,478,1986,2011]" pageId="1" pageNumber="104" refId="ref7394" refString="Lichtenstein, A. A. H. (1796) Catalogus musei zoologici ditissimi Hamburgi III auctionis lege distrahendi. 3. Insecta. Hamburg, 222 pp." type="book" year="1796">Lichtenstein, 1796</bibRefCitation>
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:? (nom. nudum) (in auction-catalog of Holthuysen collection).
</paragraph>
<paragraph id="8B9536E9F22FFFDA2BF1D6D40A948592" blockId="1.[151,1437,1986,2042]" lastBlockId="2.[151,1436,152,361]" lastPageId="2" lastPageNumber="105" pageId="1" pageNumber="104">
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<taxonomicName id="4C2A4D6AF22FFFD92BF1D6D409FD82A4" ID-CoL="4527M" authority="Olivier, 1811: 82" authorityName="Olivier" authorityPageNumber="82" authorityYear="1811" box="[151,492,2016,2041]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="104" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF22FFFD92BF1D6D4092882A4" box="[151,313,2017,2041]" italics="true" pageId="1" pageNumber="104">Mydas nitidulus</emphasis>
<treatmentCitation id="0A8B10F8F22FFFD92A59D6D509FD82A4" author="Olivier, G. A." box="[319,492,2016,2041]" page="82" pageId="1" pageNumber="104" year="1811">
<bibRefCitation id="EFBB4B18F22FFFD92A59D6D509FD82A4" author="Olivier, G. A." box="[319,492,2016,2041]" pageId="1" pageNumber="104" pagination="1 - 360" refId="ref7573" refString="Olivier, G. A. (1811) Insectes [i. e., Arthropoda], in Societe de Gens de Lettres, de Savans et d'Artistes, Encyclopedie methodique. Histoire Naturelle 8 (5), 1 - 360." type="journal article" year="1811">Olivier, 1811: 82</bibRefCitation>
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, 83. Type-locality: “South America”.
<typeStatus id="5491884BF22FFFD92800D6D70B8982A7" box="[870,920,2018,2042]" pageId="1" pageNumber="104">Type</typeStatus>
lost. Refs.—
<treatmentCitation id="0A8B10F8F22FFFD92F7DD6D50D1982A4" author="Wiedemann, C. R. W." box="[1051,1288,2016,2041]" page="626" pageId="1" pageNumber="104" year="1830">
<bibRefCitation id="EFBB4B18F22FFFD92F7DD6D50D1982A4" author="Wiedemann, C. R. W." box="[1051,1288,2016,2041]" pageId="1" pageNumber="104" refId="ref8170" refString="Wiedemann, C. R. W. (1830) Aussereuropaische zweiflu ¨ gelige Insekten. Volume 2. Schulz, Hamm, xii + 684 pp., 5 pls." type="book" year="1830">Wiedemann, 1830: 626</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0A8B10F8F22FFFDA2E28D6D5092185EC" author="Walker, F." lastPageId="2" lastPageNumber="105" page="366" pageId="1" pageNumber="104" year="1854">
<bibRefCitation id="EFBB4B18F22FFFDA2E28D6D5092185EC" author="Walker, F." lastPageId="2" lastPageNumber="105" pageId="1" pageNumber="104" refId="ref7997" refString="Walker, F. (1854) List of the specimens of dipterous insects in the collection of the British Museum. Volume 6. Supplement II. British Museum, London, 206 pp." type="book" year="1854">Walker, 1854: 366</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8B10F8F22CFFDA2A5DD1AD0A0A85EC" author="Gerstaecker, A." box="[315,539,152,177]" page="94" pageId="2" pageNumber="105" year="1868">
<bibRefCitation id="EFBB4B18F22CFFDA2A5DD1AD0A0A85EC" author="Gerstaecker, A." box="[315,539,152,177]" pageId="2" pageNumber="105" pagination="65 - 103" refId="ref7177" refString="Gerstaecker, A. (1868) Systematische Uebersicht der bis jetzt bekannt gewordenen Mydaiden (Mydasii Latr.). Stettiner Entomologische Zeitung, 29, 65 - 103, 1 pl." type="journal article" year="1868">Gerstaecker, 1868: 94</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8B10F8F22CFFDA2940D1AC0ACF85EC" author="Hunter, W. D." box="[550,734,152,177]" page="154" pageId="2" pageNumber="105" year="1901">
<bibRefCitation id="EFBB4B18F22CFFDA2940D1AC0ACF85EC" author="Hunter, W. D." box="[550,734,152,177]" pageId="2" pageNumber="105" pagination="121 - 155" refId="ref7257" refString="Hunter, W. D. (1901) A catalogue of the Diptera of South America. Part II. Transactions of the American Entomological Society, 27, 121 - 155." type="journal article" year="1901">Hunter, 1901: 154</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0A8B10F8F22CFFDA2840D1AC0BCB85EC" author="Kertesz, K." box="[806,986,153,177]" page="39" pageId="2" pageNumber="105" year="1909">
<bibRefCitation id="EFBB4B18F22CFFDA2840D1AC0BCB85EC" author="Kertesz, K." box="[806,986,153,177]" pageId="2" pageNumber="105" refId="ref7346" refString="Kertesz, K. (1909) Catalogus dipterorum hucusque descriptorum. (IV). Oncodidae, Nemestrinidae, Mydaidae, Asilidae. G. Engelmann, Budapest, 348 pp. https: // doi. org / 10.5962 / bhl. title. 515 0" type="book" year="1909">Kertész, 1909: 39</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0A8B10F8F22CFFDA2F44D1AD0CF885EC" author="d'Andretta, M. A. V." box="[1058,1257,152,177]" page="8" pageId="2" pageNumber="105" year="1951">
<bibRefCitation id="EFBB4B18F22CFFDA2F44D1AD0CF885EC" author="d'Andretta, M. A. V." box="[1058,1257,152,177]" pageId="2" pageNumber="105" pagination="1 - 76" refId="ref6544" refString="d'Andretta, M. A. V. (1951) Contribuicao para o conhecimento da familia Mydaidae. Generos: Mydas F., 1794 e Messiasia, gen. n. (Diptera). Papeis avulsos de Zoologia, 10 (1), 1 - 76." type="journal article" year="1951">dAndretta, 1951: 8</bibRefCitation>
</treatmentCitation>
,72 (listed but not treated);
<treatmentCitation id="0A8B10F8F22CFFDA2A44D18D0A528592" author="Papavero, N. &amp; Wilcox, J." box="[290,579,183,208]" page="9" pageId="2" pageNumber="105" year="1968">
<bibRefCitation id="EFBB4B18F22CFFDA2A44D18D0A528592" author="Papavero, N. &amp; Wilcox, J." box="[290,579,183,208]" pageId="2" pageNumber="105" pagination="1 - 19" refId="ref7755" refString="Papavero, N. &amp; Wilcox, J. (1968) 34. Family Mydidae (Mydaidae, Mydasidae). In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Departamento de Zoologia. Secretaria da Agricultura, Sao Paulo, pp. 1 - 19." type="book chapter" year="1968">Papavero &amp; Wilcox, 1968: 9</bibRefCitation>
</treatmentCitation>
(cat.).
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</paragraph>
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<emphasis id="B95EEAFBF22CFFDA2BF1D1E3092785B3" box="[151,310,214,238]" italics="true" pageId="2" pageNumber="105">Midas nitidulus</emphasis>
<treatmentCitation id="0A8B10F8F22CFFDA2A5AD1E00A0F85B3" author="Wiedemann, C. R. W." box="[316,542,213,238]" page="36" pageId="2" pageNumber="105" year="1831">
<bibRefCitation id="EFBB4B18F22CFFDA2A5AD1E00A0F85B3" author="Wiedemann, C. R. W." box="[316,542,213,238]" pageId="2" pageNumber="105" pagination="19 - 56" refId="ref8203" refString="Wiedemann, C. R. W. (1831) Monographia generis Midarum. In: Verhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Akademie der Naturforsche, 15 (2), 19 - 56, pls. 52 - 54." type="journal article" year="1831">Wiedemann, 1831: 36</bibRefCitation>
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, 41, pl. 52, fig. 4;
<treatmentCitation id="0A8B10F8F22CFFDA29B1D1E00BBB85B3" author="Westwood, J. O." box="[727,938,213,238]" page="51" pageId="2" pageNumber="105" year="1841">
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.
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</paragraph>
<paragraph id="8B9536E9F22CFFDA2BF1D1C00D998450" blockId="2.[151,1436,152,361]" box="[151,1416,244,269]" pageId="2" pageNumber="105">
<treatmentCitationGroup id="AB3A11C7F22CFFDA2BF1D1C00D998450" box="[151,1416,244,269]" pageId="2" pageNumber="105">
<taxonomicName id="4C2A4D6AF22CFFDA2BF1D1C00A938450" authority=": Wilcox et al., 1989: 131" authorityName="Wilcox" authorityPageNumber="131" authorityYear="1989" box="[151,642,244,269]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF22CFFDA2BF1D1C009918450" box="[151,384,245,269]" italics="true" pageId="2" pageNumber="105">Chrysomydas nitidulus</emphasis>
:
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<bibRefCitation id="EFBB4B18F22CFFDA2AEBD1C10A938450" author="Wilcox, J. &amp; Papavero, N. &amp; Pimentel, T." box="[397,642,244,269]" pageId="2" pageNumber="105" refId="ref8390" refString="Wilcox, J., Papavero, N. &amp; Pimentel, T. (1989) Studies of Mydidae (Diptera). IVb. Mydas and allies in the Americas (Mydinae: Mydini). Museu Paraense Emilio Goeldi, Belem, 139 pp." type="book" year="1989">
Wilcox
<emphasis id="B95EEAFBF22CFFDA2ABBD1C00A1B8450" box="[477,522,245,269]" italics="true" pageId="2" pageNumber="105">et al</emphasis>
., 1989: 131
</bibRefCitation>
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</taxonomicName>
(n. comb.);
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<bibRefCitation id="EFBB4B18F22CFFDA2999D1C00C288450" author="Papavero, N. &amp; Artigas, J. N." box="[767,1081,244,269]" pageId="2" pageNumber="105" pagination="117 - 134" refId="ref7674" refString="Papavero, N. &amp; Artigas, J. N. (1990) Studies of Mydidae (Diptera). VI. Catalogue of the American Species. Gayana Zoologia, 54, 117 - 134." type="journal article" year="1990">Papavero &amp; Artigas, 1990: 125</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0A8B10F8F22CFFDA2FE4D1C00D578450" author="Papavero, N." box="[1154,1350,244,269]" page="15" pageId="2" pageNumber="105" year="2009">
<bibRefCitation id="EFBB4B18F22CFFDA2FE4D1C00D578450" author="Papavero, N." box="[1154,1350,244,269]" pageId="2" pageNumber="105" pagination="1 - 31" refId="ref7618" refString="Papavero, N. (2009) Catalogue of Neotropical Diptera. Mydidae. Neotropical Diptera (14), 1 - 31. Available from: http: // revistas. ffclrp. usp. br / Neotropical _ Diptera / article / viewFile / 220 / 17 5 (accessed 20 August 2018)" type="journal article" year="2009">Papavero, 2009: 15</bibRefCitation>
</treatmentCitation>
(cat.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9536E9F22CFFDA2BF1D0260D768414" blockId="2.[151,1436,152,361]" pageId="2" pageNumber="105">
<treatmentCitationGroup id="AB3A11C7F22CFFDA2BF1D0260D768414" pageId="2" pageNumber="105">
<taxonomicName id="4C2A4D6AF22CFFDA2BF1D0260A0A8471" ID-CoL="4526D" authority="Williston, 1898: 56" authorityName="Williston" authorityPageNumber="56" authorityYear="1898" box="[151,539,275,300]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="claripennis">
<emphasis id="B95EEAFBF22CFFDA2BF1D02609438476" box="[151,338,275,299]" italics="true" pageId="2" pageNumber="105">Mydas claripennis</emphasis>
<treatmentCitation id="0A8B10F8F22CFFDA2A31D0260A0A8471" author="Williston, S. W." box="[343,539,275,300]" page="56" pageId="2" pageNumber="105" year="1898">
<bibRefCitation id="EFBB4B18F22CFFDA2A31D0260A0A8471" author="Williston, S. W." box="[343,539,275,300]" pageId="2" pageNumber="105" pagination="53 - 58" refId="ref8438" refString="Williston, S. W. (1898) Notes and descriptions of Mydaidae. Transactions of the Kansas Academy of Science, (15), 53 - 58. https: // doi. org / 10.2307 / 362381 4" type="journal article" year="1898">Williston, 1898: 56</bibRefCitation>
</treatmentCitation>
</taxonomicName>
. Type-locality:
<collectingCountry id="F33D7679F22CFFDA29D1D0260AE68471" box="[695,759,275,300]" name="Brazil" pageId="2" pageNumber="105">Brazil</collectingCountry>
,
<collectingRegion id="49EEF80BF22CFFDA2867D0210B978476" box="[769,902,275,299]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="105">Mato Grosso</collectingRegion>
, Chapada (dos Guimarães).
<typeStatus id="5491884BF22CFFDA2FC6D0210CC38471" box="[1184,1234,276,300]" pageId="2" pageNumber="105">Type</typeStatus>
<specimenCount id="9D2CFD60F22CFFDA2FBED0210CFC8476" box="[1240,1261,276,299]" pageId="2" pageNumber="105" type="male"></specimenCount>
, AMNH. Refs.—
<treatmentCitation id="0A8B10F8F22CFFDA2BA1D007096E8417" author="Hunter, W. D." box="[199,383,305,330]" page="154" pageId="2" pageNumber="105" year="1901">
<bibRefCitation id="EFBB4B18F22CFFDA2BA1D007096E8417" author="Hunter, W. D." box="[199,383,305,330]" pageId="2" pageNumber="105" pagination="121 - 155" refId="ref7257" refString="Hunter, W. D. (1901) A catalogue of the Diptera of South America. Part II. Transactions of the American Entomological Society, 27, 121 - 155." type="journal article" year="1901">Hunter, 1901: 154</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0A8B10F8F22CFFDA2AAED0070A6A8417" author="Kertesz, K." box="[456,635,305,330]" page="36" pageId="2" pageNumber="105" year="1909">
<bibRefCitation id="EFBB4B18F22CFFDA2AAED0070A6A8417" author="Kertesz, K." box="[456,635,305,330]" pageId="2" pageNumber="105" refId="ref7346" refString="Kertesz, K. (1909) Catalogus dipterorum hucusque descriptorum. (IV). Oncodidae, Nemestrinidae, Mydaidae, Asilidae. G. Engelmann, Budapest, 348 pp. https: // doi. org / 10.5962 / bhl. title. 515 0" type="book" year="1909">Kertész, 1909: 36</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0A8B10F8F22CFFDA29A2D0040B9D8417" author="d'Andretta, M. A. V." box="[708,908,305,330]" page="8" pageId="2" pageNumber="105" year="1951">
<bibRefCitation id="EFBB4B18F22CFFDA29A2D0040B9D8417" author="d'Andretta, M. A. V." box="[708,908,305,330]" pageId="2" pageNumber="105" pagination="1 - 76" refId="ref6544" refString="d'Andretta, M. A. V. (1951) Contribuicao para o conhecimento da familia Mydaidae. Generos: Mydas F., 1794 e Messiasia, gen. n. (Diptera). Papeis avulsos de Zoologia, 10 (1), 1 - 76." type="journal article" year="1951">dAndretta, 1951: 8</bibRefCitation>
</treatmentCitation>
, 73 (only listed);
<treatmentCitation id="0A8B10F8F22CFFDA2F26D0070D728414" author="Papavero, N. &amp; Wilcox, J." box="[1088,1379,305,330]" page="7" pageId="2" pageNumber="105" year="1968">
<bibRefCitation id="EFBB4B18F22CFFDA2F26D0070D728414" author="Papavero, N. &amp; Wilcox, J." box="[1088,1379,305,330]" pageId="2" pageNumber="105" pagination="1 - 19" refId="ref7755" refString="Papavero, N. &amp; Wilcox, J. (1968) 34. Family Mydidae (Mydaidae, Mydasidae). In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Departamento de Zoologia. Secretaria da Agricultura, Sao Paulo, pp. 1 - 19." type="book chapter" year="1968">Papavero &amp; Wilcox, 1968: 7</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9536E9F22CFFDA2BF1D0640AAE8434" blockId="2.[151,1436,152,361]" box="[151,703,336,361]" pageId="2" pageNumber="105">
<treatmentCitationGroup id="AB3A11C7F22CFFDA2BF1D0640AAE8434" box="[151,703,336,361]" pageId="2" pageNumber="105">
<taxonomicName id="4C2A4D6AF22CFFDA2BF1D0640A3D8434" authority="Grossbeck, 1912: 373" authorityName="Grossbeck" authorityPageNumber="373" authorityYear="1912" box="[151,556,336,361]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="clarifumis">
<emphasis id="B95EEAFBF22CFFDA2BF1D06409598434" box="[151,328,337,361]" italics="true" pageId="2" pageNumber="105">Mydas clarifumis</emphasis>
<treatmentCitation id="0A8B10F8F22CFFDA2A28D0650A3D8434" author="Grossbeck, J. A." box="[334,556,336,361]" page="373" pageId="2" pageNumber="105" year="1912">
<bibRefCitation id="EFBB4B18F22CFFDA2A28D0650A3D8434" author="Grossbeck, J. A." box="[334,556,336,361]" pageId="2" pageNumber="105" pagination="353 - 379" refId="ref7211" refString="Grossbeck, J. A. (1912) Types of insects, except Lepidoptera and Formicidae, in the American Museum of Natural History, additional to those previously listed. Bulletin of the Museum of Natural History, 31, 353 - 379." type="journal article" year="1912">Grossbeck, 1912: 373</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(misspelling).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3306562F22CFFDC2BF1D0A30C308052" lastPageId="4" lastPageNumber="107" pageId="2" pageNumber="105" type="materials_examined">
<paragraph id="8B9536E9F22CFFDA2BF1D0A3094F8743" blockId="2.[151,1437,406,1982]" pageId="2" pageNumber="105">
<emphasis id="B95EEAFBF22CFFDA2BF1D0A3090384EC" bold="true" box="[151,274,406,433]" pageId="2" pageNumber="105">Diagnosis.</emphasis>
Labellum about 1.1x longer than wide. T2T5 with long dense golden recumbent setulae directed laterally. Scutum with distinct lateral white and golden posterior pruinose spots. Male cerci partially covered by epandrial lamellae; distal process of the gonocoxite-hypandrial complex apically wide, truncate, with both dorsal and ventral tips acute.
</paragraph>
<paragraph id="8B9536E9F22CFFDA2BA1D3130BBF863F" blockId="2.[151,1437,406,1982]" pageId="2" pageNumber="105">
<emphasis id="B95EEAFBF22CFFDA2BA1D3130964871C" bold="true" box="[199,373,550,577]" pageId="2" pageNumber="105">Redescription.</emphasis>
Greatest length, excluding antennae
<quantity id="4CD29B0CF22CFFDA2872D3130B81871F" box="[788,912,550,578]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="2" pageNumber="105" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">2528 mm</quantity>
.
<emphasis id="B95EEAFBF22CFFDA28FDD3120C4A871C" bold="true" box="[923,1115,550,577]" pageId="2" pageNumber="105">Male terminalia</emphasis>
(
<figureCitation id="13112A6CF22CFFDA2F0CD3120CBE871C" box="[1130,1199,551,577]" captionStart="FIGURE 3" captionStartId="6.[151,250,1496,1521]" captionTargetBox="[151,1435,174,1457]" captionTargetId="figure@6.[151,1435,174,1457]" captionTargetPageId="6" captionText="FIGURE 3. Male and female terminalia of Chrysomydas. AD. Chrysomydas nitidulus (male, UFMG); A. Epandrium and proctiger, dorsal view; B. Gonocoxite-hypandrial complex and associate structures, dorsal view; C. same, ventral view; D. same, lateral view; EG. Chrysomydas nitidulus (female, INPA); E. Sternite 8; F. Female terminalia, dorsal view; G. same, ventral view; HK. Chrysomydas phoenix sp. nov. (male holotype, MNRJ); H. Epandrium and proctiger, dorsal view; I. Gonocoxite-hypandrial complex and associate structures, dorsal view; J. same, ventral view; K. same, lateral view. Abbreviations: cerc, cercus; ph, phallus; phc, phallic crest; sps, sclerite of spermathecal ducts; S, sternite; S9, genital fork; T, tergite." httpUri="https://zenodo.org/record/3383568/files/figure.png" pageId="2" pageNumber="105">Figs 3</figureCitation>
AD). Black, setulae black. Epandrial lamellae with distinct posterior spur, tip slightly curved. Cercus partially covered by epandrial lamella. Phallus short with bifid dorsal crest. Distal process of gonocoxite-hypandrial complex apically wide, truncate, with both dorsal and ventral tips acute.
<emphasis id="B95EEAFBF22CFFDA29E9D3A60B7887F0" bold="true" box="[655,873,658,685]" pageId="2" pageNumber="105">Female terminalia</emphasis>
(
<figureCitation id="13112A6CF22CFFDA281FD3A60BAF87F0" box="[889,958,659,685]" captionStart="FIGURE 3" captionStartId="6.[151,250,1496,1521]" captionTargetBox="[151,1435,174,1457]" captionTargetId="figure@6.[151,1435,174,1457]" captionTargetPageId="6" captionText="FIGURE 3. Male and female terminalia of Chrysomydas. AD. Chrysomydas nitidulus (male, UFMG); A. Epandrium and proctiger, dorsal view; B. Gonocoxite-hypandrial complex and associate structures, dorsal view; C. same, ventral view; D. same, lateral view; EG. Chrysomydas nitidulus (female, INPA); E. Sternite 8; F. Female terminalia, dorsal view; G. same, ventral view; HK. Chrysomydas phoenix sp. nov. (male holotype, MNRJ); H. Epandrium and proctiger, dorsal view; I. Gonocoxite-hypandrial complex and associate structures, dorsal view; J. same, ventral view; K. same, lateral view. Abbreviations: cerc, cercus; ph, phallus; phc, phallic crest; sps, sclerite of spermathecal ducts; S, sternite; S9, genital fork; T, tergite." httpUri="https://zenodo.org/record/3383568/files/figure.png" pageId="2" pageNumber="105">Figs 3</figureCitation>
EG). Black, setulae black. T9+10 anteroposteriorly narrowed. S10 with narrow membranous medial region. S8 distally with lateral and medial lobes; lateral lobes with oval sclerotized area, medial lobe largely membranous. Genital fork (S9) composed of two separated sclerites convergent anteriorly; anterior tips of sclerites bifid. Sclerite at base of spermathecal ducts slightly wider than cercus. Three spermathecae present; capsule of lateral spermathecae about as wide as spermathecal duct, with acute apex; capsule of medial spermatheca with roundish dilated apex.
</paragraph>
<paragraph id="8B9536E9F22CFFDA2BA1D25E0CCF836C" blockId="2.[151,1437,406,1982]" pageId="2" pageNumber="105">
<emphasis id="B95EEAFBF22CFFDA2BA1D25E091486D8" bold="true" box="[199,261,875,901]" pageId="2" pageNumber="105">Pupa</emphasis>
(
<figureCitation id="13112A6CF22CFFDA2A7ED25E097286D8" box="[280,355,875,901]" captionStart="FIGURE 4" captionStartId="7.[151,250,1854,1879]" captionTargetBox="[151,1435,697,1804]" captionTargetId="figure@7.[151,1435,697,1804]" captionTargetPageId="7" captionText="FIGURE 4. Pupal exuviae of Chrysomydas. AE. Chrysomydas nitidulus (female, INPA). FJ. Chrysomydas phoenix sp. nov. (male, MNRJ). Abbreviations: aap, anterior antennal process; abs, abdominal spiracles; amsp, anterior mesothoracic spines; dpp, dorsal posterolateral process; ls, labral sheath; ls1, leg sheath 1; ls2, leg sheath 2; ls3, leg sheath 3; ms, maxillary sheath; pap, posterior antennal processes; pmc, posterior mesothoracic callosity; prs, proboscidal sheath; ps; palpal sheath; pths, prothoracic spiracle; vc, ventral callosity; vpp, ventral posterolateral process; ws, wing sheath." httpUri="https://zenodo.org/record/3383570/files/figure.png" pageId="2" pageNumber="105">Figs 4</figureCitation>
AE). Greatest length, not including anterior antennal or dorsal posterolateral processes
<quantity id="4CD29B0CF22CFFDA2E0DD25E08D786F7" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="2" pageNumber="105" unit="mm" value="21.3">21.3 mm</quantity>
. Greatest width of thorax
<quantity id="4CD29B0CF22CFFDA2A8CD2BA0A5286F7" box="[490,579,911,938]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="105" unit="mm" value="4.9">4.9 mm</quantity>
. Greatest width of abdomen
<quantity id="4CD29B0CF22CFFDA28E0D2BB0BCE86F7" box="[902,991,910,938]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="105" unit="mm" value="5.2">5.2 mm</quantity>
. Body yellowish brown. Spines reddish brown. Anterior antennal processes spiniform, reddish on proximal two thirds, black on distal third, rugose, curved outwards. Posterior antennal processes reddish, rugose, with two separate posteriorly curved spines. Maxillary sheath rugose. Proboscidal sheath short, rugose, truncate. Prothoracic spiracle laterally at anterior margin of thorax on highly rugose, raised area. Anterior lateral mesothoracic callosity reddish, flat, rugose. Wing sheath with small basal reclinate spine. Thoracic area above wing sheath smoothly transversely rugose. Leg sheaths rugose. Leg sheath 3 reaching to middle of abdominal segment 2. Spiracles of abdominal segments 17 situated along midline laterally, raised, shining, reddish. Spiracle of segment 8 not as conspicuous as remaining abdominal spiracles. Abdominal segment 1 anteriorly with 13 flattened proclinate spines, posterolaterally with 89 reclinate spines, some of them bifid. Abdominal segments 27 each with 3958 posterior flattened erect to reclinate spines. Abdominal segment 2 with 58 posterior flattened erect to reclinate spines, with 21 dorsal, 910 on each lateral, and 18 ventral. Abdominal segment 3 with 52 posterior flattened erect to reclinate spines, with 19 dorsal, 78 on each lateral, and 18 ventral. Abdominal segment 4 with 52 posterior flattened erect to reclinate spines, with 18 dorsal, 78 on each lateral, and 19 ventral. Abdominal segment 5 with 49 posterior flattened erect to reclinate spines, with 17 dorsal, 78 on each lateral, and 17 ventral. Abdominal segment 6 with 46 posterior flattened erect to reclinate spines, with 15 dorsal, 78 on each lateral, and 16 ventral. Abdominal segment 7 with 39 posterior flattened erect to reclinate spines, with 14 dorsal, 6 on each lateral, and 13 ventral. Abdominal segment 8 dorsally with 7 reclinate short spines, laterally with 1 long and 12 minute reclinate spine on each of two raised tubercular areas on each side, and 10 erect ventral spines. Abdominal segment 9 with pair of divergent long posterior spiniform processes, ventrally curved.
</paragraph>
<paragraph id="8B9536E9F22CFFDA2BA1D70E097D8327" blockId="2.[151,1437,406,1982]" pageId="2" pageNumber="105">
<emphasis id="B95EEAFBF22CFFDA2BA1D70E09FF8308" bold="true" box="[199,494,1594,1621]" pageId="2" pageNumber="105">Type material examined.</emphasis>
<taxonomicName id="4C2A4D6AF22CFFDA2A93D70E0AB98308" authorityName="Olivier" authorityYear="1811" box="[501,680,1595,1621]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF22CFFDA2A93D70E0AB98308" box="[501,680,1595,1621]" italics="true" pageId="2" pageNumber="105">Mydas nitidulus</emphasis>
</taxonomicName>
:
<typeStatus id="5491884BF22CFFDA29D1D70E0B088308" box="[695,793,1595,1621]" pageId="2" pageNumber="105" type="holotype">holotype</typeStatus>
lost (type depository not stated by Olivier; see the Remarks section below).
</paragraph>
<paragraph id="8B9536E9F22CFFDA2BA1D7B60C47839C" blockId="2.[151,1437,406,1982]" pageId="2" pageNumber="105">
<taxonomicName id="4C2A4D6AF22CFFDA2BA1D7B609E783C3" authority=": Images" authorityName="Images" box="[199,502,1667,1694]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="claripennis">
<emphasis id="B95EEAFBF22CFFDA2BA1D7B6098683C0" box="[199,407,1667,1693]" italics="true" pageId="2" pageNumber="105">Mydas claripennis</emphasis>
: Images
</taxonomicName>
of the male
<typeStatus id="5491884BF22CFFDA29E7D7B60AF283C0" box="[641,739,1667,1693]" pageId="2" pageNumber="105" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2A4D6AF22CFFDA2861D7B10BB783C0" box="[775,934,1667,1694]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="claripennis">
<emphasis id="B95EEAFBF22CFFDA2861D7B10BB783C0" box="[775,934,1667,1694]" italics="true" pageId="2" pageNumber="105">C. claripennis</emphasis>
</taxonomicName>
taken by Dr. Torsten Dikow.
<collectingCountry id="F33D7679F22CFFDA2F88D7B60D4783C0" box="[1262,1366,1667,1693]" name="Brazil" pageId="2" pageNumber="105">BRAZIL</collectingCountry>
,
<collectingRegion id="49EEF80BF22CFFDA2E07D7B108FA839C" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="105">Mato Grosso</collectingRegion>
: Chapada [dos Guimarães,
<geoCoordinate id="EE1E502EF22CFFDA297BD7930A69839C" box="[541,632,1702,1729]" degrees="15" direction="south" minutes="27" orientation="latitude" pageId="2" pageNumber="105" precision="925" value="-15.45">15°27S</geoCoordinate>
,
<geoCoordinate id="EE1E502EF22CFFDA29E2D7930AFC839C" box="[644,749,1702,1729]" degrees="55" direction="west" minutes="44" orientation="longitude" pageId="2" pageNumber="105" precision="925" value="-55.733334">55°44W</geoCoordinate>
], no date, H.H. Smith (
<collectionCode id="ED3BAE2CF22CFFDA2888D7920C5B839C" box="[1006,1098,1703,1729]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="105">AMNH</collectionCode>
).
</paragraph>
<paragraph id="8B9536E9F22CFFDA2BA1D7FF0B368228" blockId="2.[151,1437,406,1982]" pageId="2" pageNumber="105">
<emphasis id="B95EEAFBF22CFFDA2BA1D7FF0A2083B8" bold="true" box="[199,561,1738,1765]" pageId="2" pageNumber="105">Additional material examined.</emphasis>
<materialsCitation id="3B423CB4F22CFFDA2950D7FF0ABB8270" ID-GBIF-Occurrence="2416711349" collectingDate="2017-04-10" collectingDateMax="2017-05-29" collectingDateMin="2017-04-10" collectionCode="INPA" collectorName="Ramal Agua Branca II" country="Brazil" latitude="-2.864722" location="Rodovia" municipality="Manaus" pageId="2" pageNumber="105" specimenCode="AM 010" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas">
<emphasis id="B95EEAFBF22CFFDA2950D7FF0AB083B8" bold="true" box="[566,673,1738,1765]" pageId="2" pageNumber="105">
<collectingCountry id="F33D7679F22CFFDA2950D7FF0AB083B8" box="[566,673,1738,1765]" name="Brazil" pageId="2" pageNumber="105">BRAZIL</collectingCountry>
</emphasis>
,
<emphasis id="B95EEAFBF22CFFDA29C8D7FF0B2583B8" bold="true" box="[686,820,1738,1765]" pageId="2" pageNumber="105">
<collectingRegion id="49EEF80BF22CFFDA29C8D7FF0B3E83B8" box="[686,815,1738,1765]" country="Brazil" name="Amazonas" pageId="2" pageNumber="105">Amazonas</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BF1AC93F22CFFDA285FD7F90B8583BB" box="[825,916,1740,1766]" pageId="2" pageNumber="105">Manaus</collectingMunicipality>
,
<location id="8EF56032F22CFFDA28F9D7FE0BEF83B8" LSID="urn:lsid:plazi:treatment:038387FFF22FFFDC2BF1D66F0C308052:8EF56032F22CFFDA28F9D7FE0BEF83B8" box="[927,1022,1739,1765]" country="Brazil" latitude="-2.864722" municipality="Manaus" name="Rodovia" pageId="2" pageNumber="105" stateProvince="Amazonas">Rodovia</location>
<specimenCode id="DB8C9E92F22CFFDA2F67D7FE0C7083B8" box="[1025,1121,1739,1765]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="2" pageNumber="105" type="Museum">AM 010</specimenCode>
, km 35,
<collectorName id="26DF533FF22CFFDA2FDAD7FE08BD8257" pageId="2" pageNumber="105">Ramal Água Branca II</collectorName>
,
<location id="8EF56032F22CFFDA2BD1D7DA099F8254" LSID="urn:lsid:plazi:treatment:038387FFF22FFFDC2BF1D66F0C308052:8EF56032F22CFFDA2BD1D7DA099F8254" box="[183,398,1775,1801]" country="Brazil" latitude="-2.864722" municipality="Manaus" name="Sitio Vida Tropical" pageId="2" pageNumber="105" stateProvince="Amazonas">Sítio Vida Tropical</location>
,
<geoCoordinate id="EE1E502EF22CFFDA2AFED7DB0A1C8254" box="[408,525,1774,1801]" degrees="2" direction="south" minutes="51" orientation="latitude" pageId="2" pageNumber="105" precision="15" seconds="53" value="-2.864722">2°5153″S</geoCoordinate>
,
<geoCoordinate id="EE1E502EF22CFFDA297ED7DB0ABC8254" box="[536,685,1774,1801]" pageId="2" pageNumber="105">59°5600″W</geoCoordinate>
,
<date id="FF941029F22CFFDA29DED7DA0B3F8254" box="[696,814,1775,1801]" pageId="2" pageNumber="105" value="2017-04-10">
<collectingDate id="EFD0E9C1F22CFFDA29DED7DA0B3F8254" box="[696,814,1775,1801]" pageId="2" pageNumber="105" value="2017-04-10">10.iv.2017</collectingDate>
</date>
(larva collected in coconut trunk),
<date id="FF941029F22CFFDA2FD3D7DA0D328254" box="[1205,1315,1775,1801]" pageId="2" pageNumber="105" value="2017-05-29">
<collectingDate id="EFD0E9C1F22CFFDA2FD3D7DA0D328254" box="[1205,1315,1775,1801]" pageId="2" pageNumber="105" value="2017-05-29">29.v.2017</collectingDate>
</date>
(emergen- ce of female imago in laboratory), (
<specimenCount id="9D2CFD60F22CFFDA2941D6260A418273" box="[551,592,1811,1838]" pageId="2" pageNumber="105" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED3BAE2CF22CFFDA2938D6260AB38270" box="[606,674,1811,1837]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="105">INPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B423CB4F22CFFDA29D2D6270CD2820C" ID-GBIF-Occurrence="2382327815" collectingDate="2016-11-19" collectionCode="UFMG" collectorName="M. F. Vasconcelos" country="Brazil" elevation="750" latitude="-20.0" location="Fazenda Bocaina" longLatPrecision="21" longitude="-43.47111" municipality="Santa Barbara" pageId="2" pageNumber="105" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Minas Gerais">
<emphasis id="B95EEAFBF22CFFDA29D2D6270B4F8270" bold="true" box="[692,862,1810,1837]" pageId="2" pageNumber="105">
<collectingRegion id="49EEF80BF22CFFDA29D2D6270B488270" box="[692,857,1810,1837]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="105">Minas Gerais</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BF1AC93F22CFFDA2803D6260C178270" box="[869,1030,1811,1837]" pageId="2" pageNumber="105">Santa Bárbara</collectingMunicipality>
,
<location id="8EF56032F22CFFDA2F77D6260CC58270" LSID="urn:lsid:plazi:treatment:038387FFF22FFFDC2BF1D66F0C308052:8EF56032F22CFFDA2F77D6260CC58270" box="[1041,1236,1811,1837]" country="Brazil" latitude="-20.0" longLatPrecision="21" longitude="-43.47111" municipality="Santa Barbara" name="Fazenda Bocaina" pageId="2" pageNumber="105" stateProvince="Minas Gerais">Fazenda Bocaina</location>
, pomar [orchard],
<geoCoordinate id="EE1E502EF22CFFDA2BF1D603090B820C" box="[151,282,1846,1873]" degrees="20" direction="south" minutes="00" orientation="latitude" pageId="2" pageNumber="105" precision="15" seconds="00" value="-20.0">20°0000″S</geoCoordinate>
,
<geoCoordinate id="EE1E502EF22CFFDA2A4FD60309A4820C" box="[297,437,1846,1873]" degrees="43" direction="west" minutes="28" orientation="longitude" pageId="2" pageNumber="105" precision="15" seconds="16" value="-43.47111">43°2816″W</geoCoordinate>
,
<date id="FF941029F22CFFDA2AA2D6020A2F820C" box="[452,574,1847,1873]" pageId="2" pageNumber="105" value="2016-11-19">
<collectingDate id="EFD0E9C1F22CFFDA2AA2D6020A2F820C" box="[452,574,1847,1873]" pageId="2" pageNumber="105" value="2016-11-19">19.xi.2016</collectingDate>
</date>
, alt.
<quantity id="4CD29B0CF22CFFDA291DD6030AD6820F" box="[635,711,1846,1874]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="105" unit="m" value="750.0">
<elevation id="0007D1DAF22CFFDA291DD6030AD6820F" box="[635,711,1846,1874]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="105" unit="m" value="750.0">750 m</elevation>
</quantity>
,
<collectorName id="26DF533FF22CFFDA29B1D60D0B8E820C" box="[727,927,1847,1874]" pageId="2" pageNumber="105">M.F. Vasconcelos</collectorName>
leg. (
<specimenCount id="9D2CFD60F22CFFDA288FD6020C07820F" box="[1001,1046,1847,1874]" pageId="2" pageNumber="105" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D2CFD60F22CFFDA2F40D6020C45820F" box="[1062,1108,1847,1874]" pageId="2" pageNumber="105" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED3BAE2CF22CFFDA2F03D6020CAA820C" box="[1125,1211,1847,1873]" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="2" pageNumber="105">UFMG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B423CB4F22CFFDA2FB7D6030B328228" ID-GBIF-Occurrence="2382327813" collectingDate="2015-01-18" collectionCode="MZUSP" collectorName="G. Ide" country="Brazil" latitude="-23.4" location="Aruja" longLatPrecision="1256" longitude="-46.3" pageId="2" pageNumber="105" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo">
<emphasis id="B95EEAFBF22CFFDA2FB7D6030D5A820C" bold="true" box="[1233,1355,1846,1873]" pageId="2" pageNumber="105">
<collectingRegion id="49EEF80BF22CFFDA2FB7D6030D5A820C" box="[1233,1355,1846,1873]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="105">São Paulo</collectingRegion>
</emphasis>
:
<location id="8EF56032F22CFFDA2E3AD6020D8D820C" LSID="urn:lsid:plazi:treatment:038387FFF22FFFDC2BF1D66F0C308052:8EF56032F22CFFDA2E3AD6020D8D820C" box="[1372,1436,1847,1873]" country="Brazil" latitude="-23.4" longLatPrecision="1256" longitude="-46.3" name="Aruja" pageId="2" pageNumber="105" stateProvince="Sao Paulo">Arujá</location>
[
<geoCoordinate id="EE1E502EF22CFFDA2BF8D66F08ED8228" box="[158,252,1882,1909]" degrees="23" direction="south" minutes="24" orientation="latitude" pageId="2" pageNumber="105" precision="925" value="-23.4">23°24S</geoCoordinate>
,
<geoCoordinate id="EE1E502EF22CFFDA2A61D66F09618228" box="[263,368,1882,1909]" degrees="46" direction="west" minutes="18" orientation="longitude" pageId="2" pageNumber="105" precision="925" value="-46.3">46°18W</geoCoordinate>
],
<date id="FF941029F22CFFDA2AE4D66F09FF8228" box="[386,494,1882,1909]" pageId="2" pageNumber="105" value="2015-01-18">
<collectingDate id="EFD0E9C1F22CFFDA2AE4D66F09FF8228" box="[386,494,1882,1909]" pageId="2" pageNumber="105" value="2015-01-18">18.i.2015</collectingDate>
</date>
,
<collectorName id="26DF533FF22CFFDA2A9CD66E0A518228" box="[506,576,1883,1909]" pageId="2" pageNumber="105">G. Ide</collectorName>
col. (
<specimenCount id="9D2CFD60F22CFFDA29E5D66E0ABC822B" box="[643,685,1883,1910]" pageId="2" pageNumber="105" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED3BAE2CF22CFFDA29DFD66E0B0D8228" box="[697,796,1883,1909]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="105">MZUSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9536E9F22CFFDA2BA1D64B0B1582E0" blockId="2.[151,1437,406,1982]" pageId="2" pageNumber="105">
<emphasis id="B95EEAFBF22CFFDA2BA1D64B094882C4" bold="true" box="[199,345,1918,1945]" pageId="2" pageNumber="105">Distribution</emphasis>
(
<figureCitation id="13112A6CF22CFFDA2A0CD64A09BE82C4" box="[362,431,1918,1945]" captionStart="FIGURE 5" captionStartId="8.[151,250,1395,1420]" captionTargetBox="[151,1435,337,1376]" captionTargetId="figure@8.[151,1435,337,1376]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map for species of Chrysomydas in South America." httpUri="https://zenodo.org/record/3383572/files/figure.png" pageId="2" pageNumber="105">Fig. 5</figureCitation>
).
<collectingCountry id="F33D7679F22CFFDA2AA3D64A0A1882C4" box="[453,521,1919,1945]" name="Brazil" pageId="2" pageNumber="105">Brazil</collectingCountry>
(
<collectingRegion id="49EEF80BF22CFFDA297CD64A0A8482C4" box="[538,661,1919,1945]" country="Brazil" name="Amazonas" pageId="2" pageNumber="105">Amazonas</collectingRegion>
,
<collectingRegion id="49EEF80BF22CFFDA29C6D6B50B2482C4" box="[672,821,1919,1946]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="105">Mato Grosso</collectingRegion>
,
<collectingRegion id="49EEF80BF22CFFDA2824D64A0BCD82C4" box="[834,988,1919,1945]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="105">Minas Gerais</collectingRegion>
,
<collectingRegion id="49EEF80BF22CFFDA288ED64A0C4F82C7" box="[1000,1118,1919,1946]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="105">São Paulo</collectingRegion>
),
<collectingCountry id="F33D7679F22CFFDA2F14D64A0D0882C4" box="[1138,1305,1919,1945]" name="French Guiana" pageId="2" pageNumber="105">French Guiana</collectingCountry>
(Cayenne),
<collectingCountry id="F33D7679F22CFFDA2BF1D69608FF82E0" box="[151,238,1955,1981]" name="Guyana" pageId="2" pageNumber="105">Guyana</collectingCountry>
(
<collectingCountry id="F33D7679F22CFFDA2B9BD696096182E0" box="[253,368,1955,1981]" name="Guyana" pageId="2" pageNumber="105">Demerara</collectingCountry>
, Georgetown, Warratuk),
<collectingCountry id="F33D7679F22CFFDA29F7D6960B1182E0" box="[657,768,1955,1981]" name="Suriname" pageId="2" pageNumber="105">Suriname</collectingCountry>
.
</paragraph>
<caption id="DF556661F22DFFDB2BF1D6810C0282AC" httpUri="https://zenodo.org/record/3383564/files/figure.png" pageId="3" pageNumber="106" startId="3.[151,250,1972,1997]" targetBox="[161,1409,166,1953]" targetPageId="3">
<paragraph id="8B9536E9F22DFFDB2BF1D6810C0282AC" blockId="3.[151,1436,1972,2033]" pageId="3" pageNumber="106">
<emphasis id="B95EEAFBF22DFFDB2BF1D68109078290" bold="true" box="[151,278,1972,1997]" pageId="3" pageNumber="106">FIGURE 1.</emphasis>
<taxonomicName id="4C2A4D6AF22DFFDB2A79D6800A1B8290" box="[287,522,1973,1997]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="106" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF22DFFDB2A79D6800A1B8290" box="[287,522,1973,1997]" italics="true" pageId="3" pageNumber="106">Chrysomydas nitidulus</emphasis>
</taxonomicName>
.
<emphasis id="B95EEAFBF22DFFDB297FD6810A5B8290" bold="true" box="[537,586,1972,1997]" pageId="3" pageNumber="106">AD</emphasis>
: male (UFMG).
<emphasis id="B95EEAFBF22DFFDB2998D6810B068290" bold="true" box="[766,791,1972,1997]" pageId="3" pageNumber="106">A.</emphasis>
Dorsal habitus;
<emphasis id="B95EEAFBF22DFFDB28A0D6800BCC8290" bold="true" box="[966,989,1973,1997]" pageId="3" pageNumber="106">B.</emphasis>
Lateral habitus;
<emphasis id="B95EEAFBF22DFFDB2FF6D6810CB98290" bold="true" box="[1168,1192,1972,1997]" pageId="3" pageNumber="106">C.</emphasis>
Head, frontal view;
<emphasis id="B95EEAFBF22DFFDB2EE2D6800D8D8290" bold="true" box="[1412,1436,1973,1997]" pageId="3" pageNumber="106">D.</emphasis>
wing;
<emphasis id="B95EEAFBF22DFFDB2BB1D6EC091E82AC" bold="true" box="[215,271,2009,2033]" pageId="3" pageNumber="106">EH.</emphasis>
female (INPA);
<emphasis id="B95EEAFBF22DFFDB2AD1D6EC09DF82AC" bold="true" box="[439,462,2009,2033]" pageId="3" pageNumber="106">E.</emphasis>
Dorsal habitus;
<emphasis id="B95EEAFBF22DFFDB2913D6EC0A9982AC" bold="true" box="[629,648,2009,2033]" pageId="3" pageNumber="106">F.</emphasis>
Lateral habitus;
<emphasis id="B95EEAFBF22DFFDB2852D6ED0B5F82AC" bold="true" box="[820,846,2008,2033]" pageId="3" pageNumber="106">G.</emphasis>
Head, lateral view.
</paragraph>
</caption>
<paragraph id="8B9536E9F22AFFDC2BA1D1AD0CC184F3" blockId="4.[151,1437,152,1295]" pageId="4" pageNumber="107">
<emphasis id="B95EEAFBF22AFFDC2BA1D1AD095285EE" bold="true" box="[199,323,152,179]" pageId="4" pageNumber="107">Bionomics</emphasis>
. Two third-instar larvae (L3) were collected on the same decaying trunk of a coconut palm (
<taxonomicName id="4C2A4D6AF22AFFDC2E30D1AC08E5858A" authorityName="Linneo" authorityYear="1753" class="Liliopsida" family="Arecaceae" genus="Cocos" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="107" phylum="Tracheophyta" rank="species" species="nucifera">
<emphasis id="B95EEAFBF22AFFDC2E30D1AC08E5858A" italics="true" pageId="4" pageNumber="107">Cocos nucifera</emphasis>
</taxonomicName>
L.), and one of those larvae completed its development in the laboratory. The mydid larva was fed
<taxonomicName id="4C2A4D6AF22AFFDC2E3FD18908DE85A7" authorityName="Leach" authorityYear="1815" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="family">Elateridae</taxonomicName>
and
<taxonomicName id="4C2A4D6AF22AFFDC2A61D1D5098A85A7" authorityName="Latreille" authorityYear="1802" box="[263,411,224,250]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
larvae (
<figureCitation id="13112A6CF22AFFDC2A90D1D50A2885A6" box="[502,569,224,251]" captionStart="FIGURE 5" captionStartId="8.[151,250,1395,1420]" captionTargetBox="[151,1435,337,1376]" captionTargetId="figure@8.[151,1435,337,1376]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map for species of Chrysomydas in South America." httpUri="https://zenodo.org/record/3383572/files/figure.png" pageId="4" pageNumber="107">Fig. 5</figureCitation>
) and also earthworms. The larva pupated after about 30 days, with the pupal stage lasting 19 days. The pupa underwent color change in the pre-emergence period, from light yellow to brown. On the nineteenth day, the pupa became immobile and rigid, non-responsive to mechanical stimulation, with ventral portion upward, and the female imago emerged (Video: https://www.youtube.com/watch?v=j18XUCOkMSQ). The adult remained motionless for about thirty minutes, and grew darker, eliminating the meconium and stretching its wings to make its first flight. The adult fly was then preserved and added to the
<collectionCode id="ED3BAE2CF22AFFDC2F70D0A00C4284F2" box="[1046,1107,405,431]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="4" pageNumber="107">INPA</collectionCode>
collection.
</paragraph>
<paragraph id="8B9536E9F22AFFDC2BA1D08C0D5787F7" blockId="4.[151,1437,152,1295]" pageId="4" pageNumber="107">
<emphasis id="B95EEAFBF22AFFDC2BA1D08C092A848E" bold="true" box="[199,315,441,467]" pageId="4" pageNumber="107">Remarks.</emphasis>
The species
<taxonomicName id="4C2A4D6AF22AFFDC2AB7D08C0A44848F" box="[465,597,440,467]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF22AFFDC2AB7D08C0A44848F" box="[465,597,440,467]" italics="true" pageId="4" pageNumber="107">C. nitidulus</emphasis>
</taxonomicName>
has a confusing history, and its
<typeStatus id="5491884BF22AFFDC28A1D08D0C38848F" box="[967,1065,440,466]" pageId="4" pageNumber="107" type="holotype">holotype</typeStatus>
has been considered lost for decades (
<bibRefCitation id="EFBB4B18F22AFFDC2B83D0E909AD84AA" author="Wilcox, J. &amp; Papavero, N. &amp; Pimentel, T." box="[229,444,476,503]" pageId="4" pageNumber="107" refId="ref8390" refString="Wilcox, J., Papavero, N. &amp; Pimentel, T. (1989) Studies of Mydidae (Diptera). IVb. Mydas and allies in the Americas (Mydinae: Mydini). Museu Paraense Emilio Goeldi, Belem, 139 pp." type="book" year="1989">Wilcox et al. 1989</bibRefCitation>
;
<bibRefCitation id="EFBB4B18F22AFFDC2AAFD0E80A6484AA" author="Papavero, N." box="[457,629,477,503]" pageId="4" pageNumber="107" pagination="1 - 31" refId="ref7618" refString="Papavero, N. (2009) Catalogue of Neotropical Diptera. Mydidae. Neotropical Diptera (14), 1 - 31. Available from: http: // revistas. ffclrp. usp. br / Neotropical _ Diptera / article / viewFile / 220 / 17 5 (accessed 20 August 2018)" type="journal article" year="2009">Papavero 2009</bibRefCitation>
;
<bibRefCitation id="EFBB4B18F22AFFDC29E4D0E80BB484AA" author="Papavero, N. &amp; Artigas, J. N." box="[642,933,476,503]" pageId="4" pageNumber="107" pagination="117 - 134" refId="ref7674" refString="Papavero, N. &amp; Artigas, J. N. (1990) Studies of Mydidae (Diptera). VI. Catalogue of the American Species. Gayana Zoologia, 54, 117 - 134." type="journal article" year="1990">Papavero &amp; Artigas 1990</bibRefCitation>
;
<bibRefCitation id="EFBB4B18F22AFFDC28D4D0E80C8A84AA" author="Papavero, N. &amp; Pimentel, T. &amp; Rafael, J. A." box="[946,1179,476,503]" pageId="4" pageNumber="107" pagination="1 - 15" refId="ref7712" refString="Papavero, N., Pimentel, T. &amp; Rafael, J. A. (2002) Insecta - Diptera - Mydidae (Mydaidae, Mydasidae). Fauna da Amazonia brasileira, Belem, 2, 1 - 15." type="journal article" year="2002">Papavero et al. 2002</bibRefCitation>
). Lichtenstein was the first to study a specimen of this species at the Holthuysen collection; however, in the Holthuysen collection catalog (
<bibRefCitation id="EFBB4B18F22AFFDC2BF9D311097D8762" author="Lichtenstein, A. A. H." box="[159,364,548,575]" pageId="4" pageNumber="107" refId="ref7394" refString="Lichtenstein, A. A. H. (1796) Catalogus musei zoologici ditissimi Hamburgi III auctionis lege distrahendi. 3. Insecta. Hamburg, 222 pp." type="book" year="1796">Lichtenstein 1796</bibRefCitation>
) edited in
<collectingRegion id="49EEF80BF22AFFDC2A81D3110A428763" box="[487,595,548,574]" country="Germany" name="Hamburg" pageId="4" pageNumber="107">Hamburg</collectingRegion>
, he referred to this species as
<taxonomicName id="4C2A4D6AF22AFFDC28CFD3110C528763" box="[937,1091,548,574]" class="Insecta" family="Mydidae" genus="Midas" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B95EEAFBF22AFFDC28CFD3110C528763" box="[937,1091,548,574]" italics="true" pageId="4" pageNumber="107">Midas nitidus</emphasis>
</taxonomicName>
(nom. nud.). This species was only formally described by
<bibRefCitation id="EFBB4B18F22AFFDC2AAFD37D0A7B873E" author="Olivier, G. A." box="[457,618,584,611]" pageId="4" pageNumber="107" pagination="1 - 360" refId="ref7573" refString="Olivier, G. A. (1811) Insectes [i. e., Arthropoda], in Societe de Gens de Lettres, de Savans et d'Artistes, Encyclopedie methodique. Histoire Naturelle 8 (5), 1 - 360." type="journal article" year="1811">Olivier (1811)</bibRefCitation>
, who did not designate type specimens. It is possible that he described the species based on the same specimen studied by Lichtenstein, since Olivier named the species almost with the same name:
<taxonomicName id="4C2A4D6AF22AFFDC2B85D3A5099A87F7" box="[227,395,656,682]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="B95EEAFBF22AFFDC2B85D3A5099A87F7" box="[227,395,656,682]" italics="true" pageId="4" pageNumber="107">Mydas nitidula</emphasis>
</taxonomicName>
(instead of
<taxonomicName id="4C2A4D6AF22AFFDC2976D3A40A9987F7" box="[528,648,656,682]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="nitudus">
<emphasis id="B95EEAFBF22AFFDC2976D3A40A9987F7" box="[528,648,656,682]" italics="true" pageId="4" pageNumber="107">M. nitudus</emphasis>
</taxonomicName>
). The short original description by Olivier is translated below:
</paragraph>
<paragraph id="8B9536E9F22AFFDC2BA1D3810D718666" blockId="4.[151,1437,152,1295]" pageId="4" pageNumber="107">
“The antennae are black. The head and the thorax are black, and covered, in some places, with red hair. The abdomen is black, with a golden-green spot on each side of the second, third, fourth, and fifth segments; a few red hairs are seen on the first. The wings are transparent, with a slight brown tint on the outer edge. The legs are black, the hind legs are little enlarged, and have short spines on their inner surface. It is found in South America” (
<bibRefCitation id="EFBB4B18F22AFFDC2FFCD2150D428666" author="Olivier, G. A." box="[1178,1363,800,827]" pageId="4" pageNumber="107" pagination="1 - 360" refId="ref7573" refString="Olivier, G. A. (1811) Insectes [i. e., Arthropoda], in Societe de Gens de Lettres, de Savans et d'Artistes, Encyclopedie methodique. Histoire Naturelle 8 (5), 1 - 360." type="journal article" year="1811">Olivier 1811: 83</bibRefCitation>
).
</paragraph>
<paragraph id="8B9536E9F22AFFDC2BA1D27109C2816A" blockId="4.[151,1437,152,1295]" pageId="4" pageNumber="107">
Decades later,
<bibRefCitation id="EFBB4B18F22AFFDC2A09D2710A598602" author="Wiedemann, C. R. W." box="[367,584,836,863]" pageId="4" pageNumber="107" refId="ref8170" refString="Wiedemann, C. R. W. (1830) Aussereuropaische zweiflu ¨ gelige Insekten. Volume 2. Schulz, Hamm, xii + 684 pp., 5 pls." type="book" year="1830">Wiedemann (1830)</bibRefCitation>
redescribed the species based on the specimen studied by Lichtenstein, an “Exemplum vetustissimum” from
<collectingCountry id="F33D7679F22AFFDC297FD25D0A9986DF" box="[537,648,872,898]" name="Suriname" pageId="4" pageNumber="107">Suriname</collectingCountry>
. He cites the name given by Olivier with two corrections: the spelling of the genus name (
<taxonomicName id="4C2A4D6AF22AFFDC2A1DD2B909D086FB" box="[379,449,908,934]" class="Insecta" family="Mydidae" genus="Midas" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAFBF22AFFDC2A1DD2B909D086FB" box="[379,449,908,934]" italics="true" pageId="4" pageNumber="107">Midas</emphasis>
</taxonomicName>
for
<taxonomicName id="4C2A4D6AF22AFFDC2A95D2B90A2F86FB" box="[499,574,908,934]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAFBF22AFFDC2A95D2B90A2F86FB" box="[499,574,908,934]" italics="true" pageId="4" pageNumber="107">Mydas</emphasis>
</taxonomicName>
) and the gender concordance of the specific epithet (
<taxonomicName id="4C2A4D6AF22AFFDC2FC7D2B90CE986FB" box="[1185,1272,908,934]" class="Insecta" family="Mydidae" genus="Mydas" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="B95EEAFBF22AFFDC2FC7D2B90CE986FB" box="[1185,1272,908,934]" italics="true" pageId="4" pageNumber="107">nitidula</emphasis>
</taxonomicName>
for
<taxonomicName id="4C2A4D6AF22AFFDC2E4CD2B90D9D86FB" box="[1322,1420,908,934]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF22AFFDC2E4CD2B90D9D86FB" box="[1322,1420,908,934]" italics="true" pageId="4" pageNumber="107">nitidulus</emphasis>
</taxonomicName>
). The description by Wiedemann fits the specimen formerly studied by Lichtenstein in all details mentioned in the description, even the abdominal color, and absence of the abdominal sternites (J. Pohl, pers. comm.). This specimen is now deposited in the Museum für Naturkunde,
<collectingRegion id="49EEF80BF22AFFDC29A5D2CD0B1D814F" box="[707,780,1016,1042]" country="Germany" name="Berlin" pageId="4" pageNumber="107">Berlin</collectingRegion>
,
<collectingCountry id="F33D7679F22AFFDC2871D2CC0B91814E" box="[791,896,1017,1043]" name="Germany" pageId="4" pageNumber="107">Germany</collectingCountry>
, and the labels are as follows (no
<typeStatus id="5491884BF22AFFDC2F9CD2CC0D3B814E" box="[1274,1322,1017,1043]" pageId="4" pageNumber="107">type</typeStatus>
label present) (J. Pohl, pers. comm.):
</paragraph>
<paragraph id="8B9536E9F22AFFDC2BA1D57509448122" blockId="4.[151,1437,152,1295]" pageId="4" pageNumber="107">
<collectingCountry id="F33D7679F22AFFDC2BB7D57509228107" box="[209,307,1088,1114]" name="Suriname" pageId="4" pageNumber="107">Surinam</collectingCountry>
/?”; “1178”; “nitidulus / Ol. Wd. /
<taxonomicName id="4C2A4D6AF22AFFDC29DBD5750B5F8107" authority="Licht" authorityName="Licht" box="[701,846,1088,1114]" class="Insecta" family="Mydidae" genus="Midas" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="nitidus">nitidus Licht</taxonomicName>
* / Mus Holth. / Kirst. ” [line breaks in the labels are indicated with /].
</paragraph>
<paragraph id="8B9536E9F22AFFDC2BA1D5BD0C308052" blockId="4.[151,1437,152,1295]" pageId="4" pageNumber="107">
Although there is a good possibility that this specimen is indeed the
<typeStatus id="5491884BF22AFFDC28A8D5BD0C2181FF" box="[974,1072,1160,1186]" pageId="4" pageNumber="107" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2A4D6AF22AFFDC2F31D5BC0CCB81FF" box="[1111,1242,1160,1187]" class="Insecta" family="Mydidae" genus="Chrysomydas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B95EEAFBF22AFFDC2F31D5BC0CCB81FF" box="[1111,1242,1160,1187]" italics="true" pageId="4" pageNumber="107">C. nitidulus</emphasis>
</taxonomicName>
, the status of the specimen remains unclear because there are no labels indicating that it was actually examined by Olivier (there are only labels by Lichtenstein and Wiedemann). We believe that this fact, more than the lack of type labels, prevents us from determining the type status of the specimen until new evidence emerges.
</paragraph>
</subSubSection>
</treatment>
</document>