260 lines
48 KiB
XML
260 lines
48 KiB
XML
<document id="3E0BF5AA4E7E89980518E036B0B1EC7B" ID-CLB-Dataset="123829" ID-DOI="10.1111/zoj.12199" ID-GBIF-Dataset="02f79ff2-493a-449e-8206-5662abfa5c52" ID-ISSN="0024-4082" ID-Zenodo-Dep="5331625" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1630169864101" checkinUser="felipe" docAuthor="Lopardo, Lara & Hormiga, Gustavo" docDate="2015" docId="03832D7710BB939EFC28E1BCFDAF5A7D" docLanguage="en" docName="zoj12199.pdf" docOrigin="Zoological Journal of the Linnean Society 173 (3)" docSource="http://dx.doi.org/10.1111/zoj.12199" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Isela Griswold 1985" docType="treatment" docVersion="11" lastPageNumber="780" masterDocId="FFBA550F10469360FFC3E17AFF985E2F" masterDocTitle="Out of the twilight zone: phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea)" masterLastPageNumber="786" masterPageNumber="527" pageNumber="780" updateTime="1699186269595" updateUser="ExternalLinkService">
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<mods:title id="FBF286DBA718D18DDC8A4CA738BD6CA9">Out of the twilight zone: phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea)</mods:title>
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<treatment id="03832D7710BB939EFC28E1BCFDAF5A7D" ID-DOI="http://doi.org/10.5281/zenodo.7004801" ID-GBIF-Taxon="199502761" ID-Zenodo-Dep="7004801" LSID="urn:lsid:plazi:treatment:03832D7710BB939EFC28E1BCFDAF5A7D" httpUri="http://treatment.plazi.org/id/03832D7710BB939EFC28E1BCFDAF5A7D" lastPageId="254" lastPageNumber="780" pageId="253" pageNumber="780">
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<subSubSection id="C330CFEA10BB939DFC28E1BCFB775EF3" box="[1003,1263,198,220]" pageId="253" pageNumber="780" type="nomenclature">
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<paragraph id="8B959C6110BB939DFC28E1BCFB775EF3" blockId="253.[892,1376,198,284]" box="[1003,1263,198,220]" pageId="253" pageNumber="780">
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<heading id="D0DD2B0D10BB939DFC28E1BCFB775EF3" box="[1003,1263,198,220]" centered="true" fontSize="9" level="2" pageId="253" pageNumber="780" reason="2">
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<taxonomicName id="4C2AE7E210BB939DFC28E1BCFB775EF3" ID-CoL="55BP" authority="GRISWOLD 1985" authorityName="Griswold" authorityYear="1985" box="[1003,1263,198,220]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
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<emphasis id="B95E407310BB939DFC28E1BCFBA85EF4" bold="true" box="[1003,1072,198,219]" italics="true" pageId="253" pageNumber="780">ISELA</emphasis>
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GRISWOLD 1985
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C330CFEA10BB939DFCBFE19CFB205F33" pageId="253" pageNumber="780" type="description">
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<paragraph id="8B959C6110BB939DFCBFE19CFB205F33" blockId="253.[892,1376,198,284]" pageId="253" pageNumber="780">
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(
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<figureCitation id="131180E410BB939DFC47E19CFC755ED4" box="[900,1005,230,251]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStartId-0="1.[164,243,1113,1132]" captionStartId-1="5.[164,243,1821,1840]" captionStartId-2="6.[144,223,1558,1577]" captionStartId-3="7.[164,243,1821,1840]" captionStartId-4="8.[144,224,1550,1569]" captionStartId-5="9.[164,243,1510,1529]" captionStartId-6="10.[144,223,1784,1803]" captionStartId-7="11.[164,242,1398,1417]" captionStartId-8="12.[144,222,1555,1574]" captionTargetBox-0="[164,1444,195,1084]" captionTargetBox-1="[229,1380,194,1791]" captionTargetBox-2="[144,1425,195,1528]" captionTargetBox-3="[226,1381,194,1791]" captionTargetBox-4="[144,1424,195,1520]" captionTargetBox-5="[164,1445,195,1480]" captionTargetBox-6="[157,1412,194,1754]" captionTargetBox-7="[164,1445,196,1368]" captionTargetBox-8="[144,1425,194,1525]" captionTargetId-0="figure-354@1.[164,1444,195,1084]" captionTargetId-1="figure-0@5.[229,1380,194,1791]" captionTargetId-2="figure-98@6.[144,1425,195,1528]" captionTargetId-3="figure-0@7.[226,1381,194,1791]" captionTargetId-4="figure-103@8.[144,1424,195,1520]" captionTargetId-5="figure-129@9.[164,1445,195,1480]" captionTargetId-6="figure-0@10.[157,1412,194,1754]" captionTargetId-7="figure-208@11.[164,1445,196,1368]" captionTargetId-8="figure-129@12.[144,1425,194,1525]" captionTargetPageId-0="1" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionText-0="Figure 1. Isela okuncana (Mysmenidae), male left palp: A, D, E, ventral view; B, C, dorsal view; C, D, detail of tip of palp; E, detail of cymbial process. See Appendix 3 for the list of abbreviations." captionText-1="Figure 2. Isela okuncana (Mysmenidae): A, D–F, female; B, C, G, H, male. A, B, carapace, lateral view; C, carapace, ventral view; D, same, detail of labium–sternum junction; E, detail of female palpal tibia; F, detail of mouthparts; G, abdomen, ventral view; H, same detail of pedicel area." captionText-2="Figure 3. Isela okuncana (Mysmenidae), legs: A, B, male; C–F, female; A, left leg I, prolateral view, tibia–metatarsus junction bearing clasping spines; B, detail of metatarsal clasping spine; C, right leg I, femur, ventral surface; D, left leg IV, tarsus, prolateral view; E, right leg I, prolateral view, detail of claws; F, right leg I, prolateral view, detail of tarsal organ." captionText-3="Figure 4. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya, male left palp: A, B, F, prolateral view; C, dorsal view; D, retrolatero–dorsal view; E, J, retrolateral view; G, I, ventral view; F, detail from figure B; I, detail from figure G. See Appendix 3 for the list of abbreviations." captionText-4="Figure 5. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; opisthosoma: A, B, D–F, female; C, male. A, B, epigynum; A, ventral view; B, lateral view. C, abdomen, ventral view; D–F, digested abdomen; D, detail of vulva; E, posterior respiratory system; F, detail of internal posterior tracheae on spiracular area. See Appendix 3 for the list of abbreviations." captionText-5="Figure 6. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; opisthosoma: A–F, female; G, male. A, right anterior lateral spinneret; B, same, left, detail; C, posterior median spinnerets; D, G, posterior lateral spinnerets; E, abdomen, posterior view; F, detail of minor ampullate spigot." captionText-6="Figure 7. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; prosoma: A, C, E–J, female; B, D, male. A, lateral view; B, dorsal view; C, ventral view; D, same, detail of labium–sternum junction; E, mouthparts, anteroventral view; F, same, detail of cheliceral fangs and promarginal teeth; G, same, detail of cheliceral denticles; H, detail of promarginal teeth; I, cheliceral fang and surrounding setae; J, cheliceral retromargin. See Appendix 3 for the list of abbreviations." captionText-7="Figure 8. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; male right leg I: A, metatarsus, dorsal view; B, same, detail of clasping spine; C, same prolateral view; D, tibia, ventral view; E, tarsus, dorsal view; F, same prolateral view." captionText-8="Figure 9. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; legs: A–C, E, female; D, F, male. A–C, E, left leg IV; D, right leg I; F, right leg IV. A, femur, prolateral view; B, same, detail of stridulatory area, C, same, close up; D, tarsus, dorsal view, tarsal organ; E, claws, retrolateral view; F, tibia, prolateral view." figureDoi-0="http://doi.org/10.5281/zenodo.5331627" figureDoi-1="http://doi.org/10.5281/zenodo.5331631" figureDoi-2="http://doi.org/10.5281/zenodo.5331633" figureDoi-3="http://doi.org/10.5281/zenodo.5331635" figureDoi-4="http://doi.org/10.5281/zenodo.5331637" figureDoi-5="http://doi.org/10.5281/zenodo.5331639" figureDoi-6="http://doi.org/10.5281/zenodo.5331641" figureDoi-7="http://doi.org/10.5281/zenodo.5331643" figureDoi-8="http://doi.org/10.5281/zenodo.5331645" httpUri-0="https://zenodo.org/record/5331627/files/figure.png" httpUri-1="https://zenodo.org/record/5331631/files/figure.png" httpUri-2="https://zenodo.org/record/5331633/files/figure.png" httpUri-3="https://zenodo.org/record/5331635/files/figure.png" httpUri-4="https://zenodo.org/record/5331637/files/figure.png" httpUri-5="https://zenodo.org/record/5331639/files/figure.png" httpUri-6="https://zenodo.org/record/5331641/files/figure.png" httpUri-7="https://zenodo.org/record/5331643/files/figure.png" httpUri-8="https://zenodo.org/record/5331645/files/figure.png" pageId="253" pageNumber="780">FIGS 1–9</figureCitation>
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,
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<figureCitation id="131180E410BB939DFC39E19CFBC35ED4" box="[1018,1115,230,251]" captionStart="Figure 128" captionStartId="144.[144,223,1784,1803]" captionTargetBox="[171,1404,198,1751]" captionTargetId="figure-0@144.[162,1404,193,1754]" captionTargetPageId="144" captionText="Figure 128. Mysmenidae female genitalia, cleared: A, Isela okuncana, dorsal view; B, Maymena mayana, dorsal view; C, Kilifina-MYSM-002-KENYA (Isela sp.), from Kwale, Kenya, dorsal view; D, Maymena ambita, ventral view; E, Maymena rica, dorsal view; F, Trogloneta granulum, ventral view; G, Mysmenopsis dipluramigo, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371567" httpUri="https://zenodo.org/record/5371567/files/figure.png" pageId="253" pageNumber="780">128A, C</figureCitation>
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,
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<figureCitation id="131180E410BB939DFBABE19CFB775ED4" box="[1128,1263,230,251]" captionStart="Figure 131" captionStartId="147.[164,242,1731,1750]" captionTargetBox="[219,1397,202,1697]" captionTargetId="figure-0@147.[209,1400,194,1701]" captionTargetPageId="147" captionText="Figure 131. Mysmenidae male left palp (unless otherwise stated), cleared: A, Mysmenopsis penai, retrolateral view, right palp, inverted; B, Mysmenopsis dipluramigo, retrolateral view; C, Mysmenopsis cidrelicola, presumably paralectotype, retrolateral view, right palp, inverted; D, Kilifina-MYSM-002-KENYA (Isela sp.), from Kwale, Kenya, ventral view, right palp, inverted; E, Trogloneta granulum, ventral view; F, Trogloneta cantareira, ventral view; G, Kilifina-MYSM-002- KENYA (Isela sp.), from Kwale, Kenya, dorsal view, right palp, inverted; H, Isela okuncana, ventral view; I, Maymena ambita, ventral view; J, Maymena mayana, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371573" httpUri="https://zenodo.org/record/5371573/files/figure.png" pageId="253" pageNumber="780">131D, G, H</figureCitation>
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,
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<figureCitation id="131180E410BB939DFB3EE19CFAC35ED4" box="[1277,1371,230,251]" captionStart="Figure 140" captionStartId="156.[144,223,1723,1742]" captionTargetBox="[189,1380,194,1691]" captionTargetId="figure-0@156.[189,1380,194,1693]" captionTargetPageId="156" captionText="Figure 140. Composite images of Mysmenidae species: Mysmenopsinae and Maymena. A–C, Isela okuncana; A, female, lateral view; B, female, ventral view; C, male, ventral view. D–F, Kilifina-MYSM-002-KENYA (Isela sp.), from Kwale, Kenya; D, female, lateral view; E, male, lateral view; F, male, ventral view. G, H, Mysmenopsis dipluramigo; G, female, lateral view; H, male, lateral view. I, Mysmenopsis palpalis, female, ventral view. J, K, Mysmenopsis cidrelicola, male paralectotype; J, lateral view; K, ventral view. L, Mysmenopsis penai, female, dorsal view. M–O, Maymena ambita; M, male, lateral view; N, female, lateral view; O, female, dorsal view. Scale bars: A–I, L–O, 0.5 mm; J, K, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5371583" httpUri="https://zenodo.org/record/5371583/files/figure.png" pageId="253" pageNumber="780">140A–F</figureCitation>
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: CLADE C158)
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</paragraph>
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</subSubSection>
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<subSubSection id="C330CFEA10BB939DFCF9E056FB125FF5" pageId="253" pageNumber="780" type="reference_group">
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<paragraph id="8B959C6110BB939DFCF9E056FBFE5F6E" blockId="253.[826,1442,300,475]" box="[826,1126,300,322]" pageId="253" pageNumber="780">
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<taxonomicName id="4C2AE7E210BB939DFCF9E056FBFA5F6E" ID-CoL="55BP" authority="Griswold, 1985: 208" authorityName="Griswold" authorityPageNumber="208" authorityYear="1985" box="[826,1122,300,322]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
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<emphasis id="B95E407310BB939DFCF9E056FCE85F6E" box="[826,880,300,321]" italics="true" pageId="253" pageNumber="780">Isela</emphasis>
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Griswold, 1985: 208
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B959C6110BB939DFC90E031FB125FF5" blockId="253.[826,1442,300,475]" pageId="253" pageNumber="780">
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<taxonomicName id="4C2AE7E210BB939DFC90E031FB665F4F" authority="Baert & Murphy, 1992: 104" authorityName="Baert & Murphy" authorityPageNumber="104" authorityYear="1992" box="[851,1278,331,353]" class="Arachnida" family="Mysmenidae" genus="Kilifina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
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<emphasis id="B95E407310BB939DFC90E031FC335F4F" box="[851,939,331,352]" italics="true" pageId="253" pageNumber="780">Kilifina</emphasis>
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Baert & Murphy, 1992: 104
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</taxonomicName>
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(replacement name for
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<taxonomicName id="4C2AE7E210BB939DFC6AE013FAAD5F50" ID-CoL="588S" authority="Baert & Murphy, 1987: 194" authorityName="Baert & Murphy" authorityPageNumber="194" authorityYear="1987" box="[937,1333,361,383]" class="Insecta" family="Coccidae" genus="Kilifia" kingdom="Animalia" order="Hemiptera" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
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<emphasis id="B95E407310BB939DFC6AE013FC695F51" box="[937,1009,361,382]" italics="true" pageId="253" pageNumber="780">Kilifia</emphasis>
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Baert & Murphy, 1987: 194
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</taxonomicName>
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, preoccupied; type species by monotypy
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<taxonomicName id="4C2AE7E210BB939DFB61E0F2FC745F93" authority="Baert & Murphy, 1987" authorityName="Baert & Murphy" authorityYear="1987" class="Insecta" family="Coccidae" genus="Kilifia" kingdom="Animalia" order="Hemiptera" pageId="253" pageNumber="780" phylum="Arthropoda" rank="species" species="inquilina">
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<emphasis id="B95E407310BB939DFB61E0F2FAC25FB2" box="[1186,1370,392,413]" italics="true" pageId="253" pageNumber="780">Kilifia inquilina</emphasis>
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Baert & Murphy, 1987
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</taxonomicName>
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,
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<typeStatus id="549122C310BB939DFC3BE0DDFBC05F93" box="[1016,1112,423,444]" pageId="253" pageNumber="780" type="paratype">paratype</typeStatus>
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material in IRSN and MRAC, examined).
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<emphasis id="B95E407310BB939DFC02E0BFFB125FF5" bold="true" box="[961,1162,453,474]" pageId="253" pageNumber="780">New synonymy.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C330CFEA10BB939DFCF9E37DFC285C57" pageId="253" pageNumber="780" type="type_taxon">
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<paragraph id="8B959C6110BB939DFCF9E37DFC5B5C33" blockId="253.[826,1441,519,632]" box="[826,963,519,540]" pageId="253" pageNumber="780">
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<emphasis id="B95E407310BB939DFCF9E37DFC5B5C33" box="[826,963,519,540]" italics="true" pageId="253" pageNumber="780">
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<typeStatus id="549122C310BB939DFCF9E37DFCF65C33" box="[826,878,519,540]" pageId="253" pageNumber="780">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph id="8B959C6110BB939DFCF9E35FFC285C57" blockId="253.[826,1441,519,632]" pageId="253" pageNumber="780">
|
||
<taxonomicName id="4C2AE7E210BB939DFCF9E35FFB3E5C14" ID-CoL="3Q6VB" authority="Griswold 1985" authorityName="Griswold" authorityYear="1985" box="[826,1190,549,571]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="species" species="okuncana">
|
||
<emphasis id="B95E407310BB939DFCF9E35FFC755C15" box="[826,1005,549,570]" italics="true" pageId="253" pageNumber="780">Isela okuncana</emphasis>
|
||
Griswold 1985
|
||
</taxonomicName>
|
||
by original designation and monotypy,
|
||
<typeStatus id="549122C310BB939DFBDDE33EFBD75C76" box="[1054,1103,580,601]" pageId="253" pageNumber="780">type</typeStatus>
|
||
material in NMSA and CAS, examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330CFEA10BB939DFCF9E3DEFA3A58DF" pageId="253" pageNumber="780" type="discussion">
|
||
<paragraph id="8B959C6110BB939DFCF9E3DEFBA75C97" blockId="253.[826,1441,675,973]" box="[826,1087,675,697]" pageId="253" pageNumber="780">
|
||
<emphasis id="B95E407310BB939DFCF9E3DEFBA75C97" box="[826,1087,675,697]" italics="true" pageId="253" pageNumber="780">Synonymy justification</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B959C6110BB939DFCF9E3B8FC5A5DE2" blockId="253.[826,1441,675,973]" pageId="253" pageNumber="780">
|
||
Both of these African monotypic genera share extremely peculiar male and female genitalia (see
|
||
<figureCitation id="131180E410BB939DFAEBE39BFCD65D3B" captionStart="Figure 128" captionStartId="144.[144,223,1784,1803]" captionTargetBox="[171,1404,198,1751]" captionTargetId="figure-0@144.[162,1404,193,1754]" captionTargetPageId="144" captionText="Figure 128. Mysmenidae female genitalia, cleared: A, Isela okuncana, dorsal view; B, Maymena mayana, dorsal view; C, Kilifina-MYSM-002-KENYA (Isela sp.), from Kwale, Kenya, dorsal view; D, Maymena ambita, ventral view; E, Maymena rica, dorsal view; F, Trogloneta granulum, ventral view; G, Mysmenopsis dipluramigo, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371567" httpUri="https://zenodo.org/record/5371567/files/figure.png" pageId="253" pageNumber="780">Figs 128A, C</figureCitation>
|
||
,
|
||
<figureCitation id="131180E410BB939DFC98E385FC475D3A" box="[859,991,767,789]" captionStart="Figure 131" captionStartId="147.[164,242,1731,1750]" captionTargetBox="[219,1397,202,1697]" captionTargetId="figure-0@147.[209,1400,194,1701]" captionTargetPageId="147" captionText="Figure 131. Mysmenidae male left palp (unless otherwise stated), cleared: A, Mysmenopsis penai, retrolateral view, right palp, inverted; B, Mysmenopsis dipluramigo, retrolateral view; C, Mysmenopsis cidrelicola, presumably paralectotype, retrolateral view, right palp, inverted; D, Kilifina-MYSM-002-KENYA (Isela sp.), from Kwale, Kenya, ventral view, right palp, inverted; E, Trogloneta granulum, ventral view; F, Trogloneta cantareira, ventral view; G, Kilifina-MYSM-002- KENYA (Isela sp.), from Kwale, Kenya, dorsal view, right palp, inverted; H, Isela okuncana, ventral view; I, Maymena ambita, ventral view; J, Maymena mayana, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371573" httpUri="https://zenodo.org/record/5371573/files/figure.png" pageId="253" pageNumber="780">131D, G, H</figureCitation>
|
||
), form a clade in all morphological and combined analyses, and share several other morphological synapomorphies. The
|
||
<typeStatus id="549122C310BB939DFB5EE247FB575D7D" box="[1181,1231,829,850]" pageId="253" pageNumber="780">type</typeStatus>
|
||
material of both species have been examined as well as a third of undescribed species from
|
||
<collectingCountry id="F33DDCF110BB939DFBAFE200FB205DA0" box="[1132,1208,890,911]" name="Kenya" pageId="253" pageNumber="780">Kenya</collectingCountry>
|
||
, and the conformation of the genital morphology remains consistent across all of them.
|
||
</paragraph>
|
||
<paragraph id="8B959C6110BB939DFCF9E282FB445A21" blockId="253.[826,1441,1016,1221]" box="[826,1244,1016,1038]" pageId="253" pageNumber="780">
|
||
<emphasis id="B95E407310BB939DFCF9E282FB445A21" box="[826,1244,1016,1038]" italics="true" pageId="253" pageNumber="780">Familial placement and composition</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B959C6110BB939DFCF9E56CFAD05AEA" blockId="253.[826,1441,1016,1221]" pageId="253" pageNumber="780">
|
||
<taxonomicName id="4C2AE7E210BB939DFCF9E56CFCE85A04" authorityName="Griswold" authorityYear="1985" box="[826,880,1046,1067]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310BB939DFCF9E56CFCE85A04" box="[826,880,1046,1067]" italics="true" pageId="253" pageNumber="780">Isela</emphasis>
|
||
</taxonomicName>
|
||
is sister to
|
||
<taxonomicName id="4C2AE7E210BB939DFBC4E56DFB065A03" box="[1031,1182,1047,1068]" class="Arachnida" family="Mysmenidae" genus="Mysmenopsis" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310BB939DFBC4E56DFB065A03" box="[1031,1182,1047,1068]" italics="true" pageId="253" pageNumber="780">Mysmenopsis</emphasis>
|
||
</taxonomicName>
|
||
within the subfamily
|
||
<taxonomicName id="4C2AE7E210BB939DFCF9E54FFC775A64" box="[826,1007,1077,1099]" family="Mysmenidae" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" rank="subFamily" subFamily="Mysmenopsinae">Mysmenopsinae</taxonomicName>
|
||
(
|
||
<figureCitation id="131180E410BB939DFC3CE54FFBF35A64" box="[1023,1131,1077,1099]" captionStart="Figure 161" captionStartId="192.[144,221,1256,1275]" captionTargetBox="[144,1424,433,1225]" captionTargetId="figure-272@192.[144,1425,474,1226]" captionTargetPageId="192" captionText="Figure 161. Hypothesis of relationships within Mysmenidae, taken from the results of the combined analysis from Lopardo et al. (2011) (see Fig. 160). New taxonomic (i.e. generic) status as proposed in our study is assigned to each representative; new combinations are emphasized in bold. Unambiguous morphological character optimizations are shown for every branch in the tree. Numbers below each node indicate node numbers. Empty and filled boxes represent homoplasious and non-homoplasious transformations, respectively. Major clades representing taxonomic decisions discussed in the text are highlighted in grey boxes." figureDoi="http://doi.org/10.5281/zenodo.5371617" httpUri="https://zenodo.org/record/5371617/files/figure.png" pageId="253" pageNumber="780">Fig. 161B</figureCitation>
|
||
). Currently, and as defined here, the genus
|
||
<taxonomicName id="4C2AE7E210BB939DFC29E52EFB865A46" authorityName="Griswold" authorityYear="1985" box="[1002,1054,1108,1129]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310BB939DFC29E52EFB865A46" box="[1002,1054,1108,1129]" italics="true" pageId="253" pageNumber="780">Isela</emphasis>
|
||
</taxonomicName>
|
||
comprises two species (
|
||
<taxonomicName id="4C2AE7E210BB939DFAE1E52EFA395A46" authorityName="Griswold" authorityYear="1985" box="[1314,1441,1108,1129]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="species" species="okuncana">
|
||
<emphasis id="B95E407310BB939DFAE1E52EFA395A46" box="[1314,1441,1108,1129]" italics="true" pageId="253" pageNumber="780">I. okuncana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2AE7E210BB939DFCB3E509FC6A5AA8" authorityName="Lopardo & Hormiga" authorityYear="2015" box="[880,1010,1138,1160]" class="Insecta" family="Coccidae" genus="Kilifia" kingdom="Animalia" order="Hemiptera" pageId="253" pageNumber="780" phylum="Arthropoda" rank="species" species="inquilina" status="comb. nov.">
|
||
<emphasis id="B95E407310BB939DFCB3E509FC6A5AA8" box="[880,1010,1138,1160]" italics="true" pageId="253" pageNumber="780">I. inquilina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26DFD0810BB939DFC3BE508FBE95AA7" box="[1016,1137,1138,1160]" pageId="253" pageNumber="780" rank="species">comb. nov.</taxonomicNameLabel>
|
||
), and is here represented by one described plus one undescribed species:
|
||
<taxonomicName id="4C2AE7E210BB939DFCF9E5CAFC585AEB" authorityName="Griswold" authorityYear="1985" box="[826,960,1199,1221]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="species" species="okuncana">
|
||
<emphasis id="B95E407310BB939DFCF9E5CAFC585AEB" box="[826,960,1199,1221]" italics="true" pageId="253" pageNumber="780">I. okuncana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B95E407310BB939DFC3CE5D5FBC65AEB" box="[1023,1118,1199,1220]" italics="true" pageId="253" pageNumber="780">Kilifina-</emphasis>
|
||
MYSM-002-
|
||
<collectingCountry id="F33DDCF110BB939DFB25E5CAFADC5AEA" box="[1254,1348,1200,1221]" name="Kenya" pageId="253" pageNumber="780">KENYA</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B959C6110BB939DFCF9E58BFC6F5B29" blockId="253.[826,1442,1265,1776]" box="[826,1015,1265,1286]" pageId="253" pageNumber="780">
|
||
<emphasis id="B95E407310BB939DFCF9E58BFC6F5B29" box="[826,1015,1265,1286]" italics="true" pageId="253" pageNumber="780">Synapomorphies</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B959C6110BB939DFCF9E475FA3A58DF" blockId="253.[826,1442,1265,1776]" pageId="253" pageNumber="780">
|
||
Morphological synapomorphies of
|
||
<taxonomicName id="4C2AE7E210BB939DFB70E475FB7F5B0B" authorityName="Griswold" authorityYear="1985" box="[1203,1255,1295,1316]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310BB939DFB70E475FB7F5B0B" box="[1203,1255,1295,1316]" italics="true" pageId="253" pageNumber="780">Isela</emphasis>
|
||
</taxonomicName>
|
||
include: females with coiled tubuliform spermathecae (
|
||
<figureCitation id="131180E410BB939DFB35E454FAC95B6B" box="[1270,1361,1326,1348]" captionStart="Figure 5" captionStartId="8.[144,224,1550,1569]" captionTargetBox="[144,1424,195,1520]" captionTargetId="figure-103@8.[144,1424,195,1520]" captionTargetPageId="8" captionText="Figure 5. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; opisthosoma: A, B, D–F, female; C, male. A, B, epigynum; A, ventral view; B, lateral view. C, abdomen, ventral view; D–F, digested abdomen; D, detail of vulva; E, posterior respiratory system; F, detail of internal posterior tracheae on spiracular area. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331637" httpUri="https://zenodo.org/record/5331637/files/figure.png" pageId="253" pageNumber="780">Figs 5D</figureCitation>
|
||
,
|
||
<figureCitation id="131180E410BB939DFA9CE454FCD45B4C" captionStart="Figure 128" captionStartId="144.[144,223,1784,1803]" captionTargetBox="[171,1404,198,1751]" captionTargetId="figure-0@144.[162,1404,193,1754]" captionTargetPageId="144" captionText="Figure 128. Mysmenidae female genitalia, cleared: A, Isela okuncana, dorsal view; B, Maymena mayana, dorsal view; C, Kilifina-MYSM-002-KENYA (Isela sp.), from Kwale, Kenya, dorsal view; D, Maymena ambita, ventral view; E, Maymena rica, dorsal view; F, Trogloneta granulum, ventral view; G, Mysmenopsis dipluramigo, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371567" httpUri="https://zenodo.org/record/5371567/files/figure.png" pageId="253" pageNumber="780">128A, C</figureCitation>
|
||
) and without flagelliform gland spigots (
|
||
<figureCitation id="131180E410BB939DFAC8E436FAC65B4D" box="[1291,1374,1356,1378]" captionStart="Figure 6" captionStartId="9.[164,243,1510,1529]" captionTargetBox="[164,1445,195,1480]" captionTargetId="figure-129@9.[164,1445,195,1480]" captionTargetPageId="9" captionText="Figure 6. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; opisthosoma: A–F, female; G, male. A, right anterior lateral spinneret; B, same, left, detail; C, posterior median spinnerets; D, G, posterior lateral spinnerets; E, abdomen, posterior view; F, detail of minor ampullate spigot." figureDoi="http://doi.org/10.5281/zenodo.5331639" httpUri="https://zenodo.org/record/5331639/files/figure.png" pageId="253" pageNumber="780">Fig. 6D</figureCitation>
|
||
); both sexes with median trichobothria on metatarsus I (
|
||
<figureCitation id="131180E410BB939DFC81E4F0FC005B8F" box="[834,920,1418,1440]" captionStart="Figure 8" captionStartId="11.[164,242,1398,1417]" captionTargetBox="[164,1445,196,1368]" captionTargetId="figure-208@11.[164,1445,196,1368]" captionTargetPageId="11" captionText="Figure 8. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; male right leg I: A, metatarsus, dorsal view; B, same, detail of clasping spine; C, same prolateral view; D, tibia, ventral view; E, tarsus, dorsal view; F, same prolateral view." figureDoi="http://doi.org/10.5281/zenodo.5331643" httpUri="https://zenodo.org/record/5331643/files/figure.png" pageId="253" pageNumber="780">Fig. 8A</figureCitation>
|
||
); and males with secondary external cymbial conductor (
|
||
<figureCitation id="131180E410BB939DFC00E4D2FB855B91" box="[963,1053,1448,1470]" captionStart="Figure 4" captionStartId="7.[164,243,1821,1840]" captionTargetBox="[226,1381,194,1791]" captionTargetId="figure-0@7.[226,1381,194,1791]" captionTargetPageId="7" captionText="Figure 4. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya, male left palp: A, B, F, prolateral view; C, dorsal view; D, retrolatero–dorsal view; E, J, retrolateral view; G, I, ventral view; F, detail from figure B; I, detail from figure G. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331635" httpUri="https://zenodo.org/record/5331635/files/figure.png" pageId="253" pageNumber="780">Fig. 4G</figureCitation>
|
||
); internal cymbial tarsal organ (
|
||
<figureCitation id="131180E410BB939DFC81E4BDFC175BF2" box="[834,911,1479,1501]" captionStart="Figure 4" captionStartId="7.[164,243,1821,1840]" captionTargetBox="[226,1381,194,1791]" captionTargetId="figure-0@7.[226,1381,194,1791]" captionTargetPageId="7" captionText="Figure 4. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya, male left palp: A, B, F, prolateral view; C, dorsal view; D, retrolatero–dorsal view; E, J, retrolateral view; G, I, ventral view; F, detail from figure B; I, detail from figure G. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331635" httpUri="https://zenodo.org/record/5331635/files/figure.png" pageId="253" pageNumber="780">Fig. 4I</figureCitation>
|
||
); coiled embolus (
|
||
<figureCitation id="131180E410BB939DFBA2E4BDFB265BF3" box="[1121,1214,1479,1501]" captionStart="Figure 4" captionStartId="7.[164,243,1821,1840]" captionTargetBox="[226,1381,194,1791]" captionTargetId="figure-0@7.[226,1381,194,1791]" captionTargetPageId="7" captionText="Figure 4. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya, male left palp: A, B, F, prolateral view; C, dorsal view; D, retrolatero–dorsal view; E, J, retrolateral view; G, I, ventral view; F, detail from figure B; I, detail from figure G. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331635" httpUri="https://zenodo.org/record/5331635/files/figure.png" pageId="253" pageNumber="780">Figs 4H</figureCitation>
|
||
,
|
||
<figureCitation id="131180E410BB939DFB0FE4BDFA935BF2" box="[1228,1291,1479,1501]" captionStart="Figure 131" captionStartId="147.[164,242,1731,1750]" captionTargetBox="[219,1397,202,1697]" captionTargetId="figure-0@147.[209,1400,194,1701]" captionTargetPageId="147" captionText="Figure 131. Mysmenidae male left palp (unless otherwise stated), cleared: A, Mysmenopsis penai, retrolateral view, right palp, inverted; B, Mysmenopsis dipluramigo, retrolateral view; C, Mysmenopsis cidrelicola, presumably paralectotype, retrolateral view, right palp, inverted; D, Kilifina-MYSM-002-KENYA (Isela sp.), from Kwale, Kenya, ventral view, right palp, inverted; E, Trogloneta granulum, ventral view; F, Trogloneta cantareira, ventral view; G, Kilifina-MYSM-002- KENYA (Isela sp.), from Kwale, Kenya, dorsal view, right palp, inverted; H, Isela okuncana, ventral view; I, Maymena ambita, ventral view; J, Maymena mayana, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371573" httpUri="https://zenodo.org/record/5371573/files/figure.png" pageId="253" pageNumber="780">131H</figureCitation>
|
||
); and two or more prolateral tibial trichobothria on male palp (
|
||
<figureCitation id="131180E410BB939DFC81E77EFC015836" box="[834,921,1540,1562]" captionStart="Figure 1" captionStartId="1.[164,243,1113,1132]" captionTargetBox="[164,1444,195,1084]" captionTargetId="figure-354@1.[164,1444,195,1084]" captionTargetPageId="1" captionText="Figure 1. Isela okuncana (Mysmenidae), male left palp: A, D, E, ventral view; B, C, dorsal view; C, D, detail of tip of palp; E, detail of cymbial process. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331627" httpUri="https://zenodo.org/record/5331627/files/figure.png" pageId="253" pageNumber="780">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="131180E410BB939DFC65E77EFC505835" box="[934,968,1540,1562]" captionStart="Figure 4" captionStartId="7.[164,243,1821,1840]" captionTargetBox="[226,1381,194,1791]" captionTargetId="figure-0@7.[226,1381,194,1791]" captionTargetPageId="7" captionText="Figure 4. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya, male left palp: A, B, F, prolateral view; C, dorsal view; D, retrolatero–dorsal view; E, J, retrolateral view; G, I, ventral view; F, detail from figure B; I, detail from figure G. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331635" httpUri="https://zenodo.org/record/5331635/files/figure.png" pageId="253" pageNumber="780">4E</figureCitation>
|
||
). Ambiguously optimized synapomorphies for this genus include: posterior respiratory system with median and single lateral tracheae (
|
||
<figureCitation id="131180E410BB939DFB06E73BFA8D5878" box="[1221,1301,1601,1623]" captionStart="Figure 5" captionStartId="8.[144,224,1550,1569]" captionTargetBox="[144,1424,195,1520]" captionTargetId="figure-103@8.[144,1424,195,1520]" captionTargetPageId="8" captionText="Figure 5. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; opisthosoma: A, B, D–F, female; C, male. A, B, epigynum; A, ventral view; B, lateral view. C, abdomen, ventral view; D–F, digested abdomen; D, detail of vulva; E, posterior respiratory system; F, detail of internal posterior tracheae on spiracular area. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331637" httpUri="https://zenodo.org/record/5331637/files/figure.png" pageId="253" pageNumber="780">Fig. 5E</figureCitation>
|
||
); males with twisted metatarsal clasping spine (
|
||
<figureCitation id="131180E410BB939DFB74E71AFA94585A" box="[1207,1292,1632,1654]" captionStart="Figure 3" captionStartId="6.[144,223,1558,1577]" captionTargetBox="[144,1425,195,1528]" captionTargetId="figure-98@6.[144,1425,195,1528]" captionTargetPageId="6" captionText="Figure 3. Isela okuncana (Mysmenidae), legs: A, B, male; C–F, female; A, left leg I, prolateral view, tibia–metatarsus junction bearing clasping spines; B, detail of metatarsal clasping spine; C, right leg I, femur, ventral surface; D, left leg IV, tarsus, prolateral view; E, right leg I, prolateral view, detail of claws; F, right leg I, prolateral view, detail of tarsal organ." figureDoi="http://doi.org/10.5281/zenodo.5331633" httpUri="https://zenodo.org/record/5331633/files/figure.png" pageId="253" pageNumber="780">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="131180E410BB939DFAD4E71AFACE585A" box="[1303,1366,1632,1653]" captionStart="Figure 8" captionStartId="11.[164,242,1398,1417]" captionTargetBox="[164,1445,196,1368]" captionTargetId="figure-208@11.[164,1445,196,1368]" captionTargetPageId="11" captionText="Figure 8. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya; male right leg I: A, metatarsus, dorsal view; B, same, detail of clasping spine; C, same prolateral view; D, tibia, ventral view; E, tarsus, dorsal view; F, same prolateral view." figureDoi="http://doi.org/10.5281/zenodo.5331643" httpUri="https://zenodo.org/record/5331643/files/figure.png" pageId="253" pageNumber="780">8B, C</figureCitation>
|
||
,
|
||
<figureCitation id="131180E410BB939DFAA1E71AFCD458BB" captionStart="Figure 140" captionStartId="156.[144,223,1723,1742]" captionTargetBox="[189,1380,194,1691]" captionTargetId="figure-0@156.[189,1380,194,1693]" captionTargetPageId="156" captionText="Figure 140. Composite images of Mysmenidae species: Mysmenopsinae and Maymena. A–C, Isela okuncana; A, female, lateral view; B, female, ventral view; C, male, ventral view. D–F, Kilifina-MYSM-002-KENYA (Isela sp.), from Kwale, Kenya; D, female, lateral view; E, male, lateral view; F, male, ventral view. G, H, Mysmenopsis dipluramigo; G, female, lateral view; H, male, lateral view. I, Mysmenopsis palpalis, female, ventral view. J, K, Mysmenopsis cidrelicola, male paralectotype; J, lateral view; K, ventral view. L, Mysmenopsis penai, female, dorsal view. M–O, Maymena ambita; M, male, lateral view; N, female, lateral view; O, female, dorsal view. Scale bars: A–I, L–O, 0.5 mm; J, K, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5371583" httpUri="https://zenodo.org/record/5371583/files/figure.png" pageId="253" pageNumber="780">140E, F</figureCitation>
|
||
); epiandrous fusules dispersed in a row; palpal tibial bearing spine-like strong setae (
|
||
<figureCitation id="131180E410BB939DFB73E7E7FAB4589D" box="[1200,1324,1693,1715]" captionStart="Figure 1" captionStartId="1.[164,243,1113,1132]" captionTargetBox="[164,1444,195,1084]" captionTargetId="figure-354@1.[164,1444,195,1084]" captionTargetPageId="1" captionText="Figure 1. Isela okuncana (Mysmenidae), male left palp: A, D, E, ventral view; B, C, dorsal view; C, D, detail of tip of palp; E, detail of cymbial process. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331627" httpUri="https://zenodo.org/record/5331627/files/figure.png" pageId="253" pageNumber="780">Figs 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="131180E410BB939DFAF9E7E7FAE5589D" box="[1338,1405,1693,1714]" captionStart="Figure 4" captionStartId="7.[164,243,1821,1840]" captionTargetBox="[226,1381,194,1791]" captionTargetId="figure-0@7.[226,1381,194,1791]" captionTargetPageId="7" captionText="Figure 4. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya, male left palp: A, B, F, prolateral view; C, dorsal view; D, retrolatero–dorsal view; E, J, retrolateral view; G, I, ventral view; F, detail from figure B; I, detail from figure G. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331635" httpUri="https://zenodo.org/record/5331635/files/figure.png" pageId="253" pageNumber="780">4A, E</figureCitation>
|
||
); a prolateral apical secondary cymbial conductor (
|
||
<figureCitation id="131180E410BB939DFA8AE7C6FCD458DF" captionStart="Figure 4" captionStartId="7.[164,243,1821,1840]" captionTargetBox="[226,1381,194,1791]" captionTargetId="figure-0@7.[226,1381,194,1791]" captionTargetPageId="7" captionText="Figure 4. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya, male left palp: A, B, F, prolateral view; C, dorsal view; D, retrolatero–dorsal view; E, J, retrolateral view; G, I, ventral view; F, detail from figure B; I, detail from figure G. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331635" httpUri="https://zenodo.org/record/5331635/files/figure.png" pageId="253" pageNumber="780">Fig. 4G, F</figureCitation>
|
||
); and a row of small setae on cymbial fold (
|
||
<figureCitation id="131180E410BB939DFAFCE7A0FA0D58DF" box="[1343,1429,1754,1776]" captionStart="Figure 4" captionStartId="7.[164,243,1821,1840]" captionTargetBox="[226,1381,194,1791]" captionTargetId="figure-0@7.[226,1381,194,1791]" captionTargetPageId="7" captionText="Figure 4. Kilifina-MYSM-002-KENYA (Isela sp., Mysmenidae) from Kwale, Kenya, male left palp: A, B, F, prolateral view; C, dorsal view; D, retrolatero–dorsal view; E, J, retrolateral view; G, I, ventral view; F, detail from figure B; I, detail from figure G. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331635" httpUri="https://zenodo.org/record/5331635/files/figure.png" pageId="253" pageNumber="780">Fig. 4G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330CFEA10BB939EFCF9E666FDAF5A7D" lastPageId="254" lastPageNumber="781" pageId="253" pageNumber="780" type="diagnosis">
|
||
<paragraph id="8B959C6110BB939DFCF9E666FC32591E" blockId="253.[826,1441,1820,1903]" box="[826,938,1820,1841]" pageId="253" pageNumber="780">
|
||
<emphasis id="B95E407310BB939DFCF9E666FC32591E" box="[826,938,1820,1841]" italics="true" pageId="253" pageNumber="780">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B959C6110BB939EFCF9E640FDAF5A7D" blockId="253.[826,1441,1820,1903]" lastBlockId="254.[144,759,198,1107]" lastPageId="254" lastPageNumber="781" pageId="253" pageNumber="780">
|
||
<taxonomicName id="4C2AE7E210BB939DFCF9E640FCEC5960" authorityName="Griswold" authorityYear="1985" box="[826,884,1850,1871]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="780" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310BB939DFCF9E640FCEC5960" box="[826,884,1850,1871]" italics="true" pageId="253" pageNumber="780">Isela</emphasis>
|
||
</taxonomicName>
|
||
differs from all other mysmenid genera by the spine-like strong setae on male palpal tibial, a secondary cymbial conductor located prolaterally– apically; females without flagelliform gland spigots and with coiled tubuliform spermathecae; posterior median and single lateral tracheae; and trichobothrium located medially on metatarsus I. In addition, the following combination of male features is unique for
|
||
<taxonomicName id="4C2AE7E210B8939EFD9FE025FD0A5F5B" authorityName="Griswold" authorityYear="1985" box="[604,658,351,372]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="254" pageNumber="781" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310B8939EFD9FE025FD0A5F5B" box="[604,658,351,372]" italics="true" pageId="254" pageNumber="781">Isela</emphasis>
|
||
</taxonomicName>
|
||
: twisted metatarsal clasping spine; internal cymbial tarsal organ; a row of small setae on cymbial fold; coiled embolus; and epiandrous fusules dispersed in a row. Originally,
|
||
<taxonomicName id="4C2AE7E210B8939EFF19E0A3FE895FC1" authorityName="Griswold" authorityYear="1985" box="[218,273,473,494]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="254" pageNumber="781" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310B8939EFF19E0A3FE895FC1" box="[218,273,473,494]" italics="true" pageId="254" pageNumber="781">Isela</emphasis>
|
||
</taxonomicName>
|
||
was diagnosed by the carapace depression separating the posterior median eyes (PME) from the anterior eye row (AER); the male palpal tibia large, cup-shaped, and with stout dorsoapical spines; by the morphology of cephalothorax; and by the general morphology of male and female genitalia (Griswold, 1985). Baert & Murphy (1987) also noted the general resemblance of
|
||
<taxonomicName id="4C2AE7E210B8939EFF3FE3D5FECA5CEB" authorityName="Baert & Murphy" authorityYear="1992" box="[252,338,687,708]" class="Arachnida" family="Mysmenidae" genus="Kilifina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="254" pageNumber="781" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310B8939EFF3FE3D5FECA5CEB" box="[252,338,687,708]" italics="true" pageId="254" pageNumber="781">Kilifina</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C2AE7E210B8939EFE56E3D5FE535CEB" authorityName="Griswold" authorityYear="1985" box="[405,459,687,708]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="254" pageNumber="781" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310B8939EFE56E3D5FE535CEB" box="[405,459,687,708]" italics="true" pageId="254" pageNumber="781">Isela</emphasis>
|
||
</taxonomicName>
|
||
in terms of carapace morphology, the separation of the PME from the AER, and the swollen femora I, but stated that the differences between these two genera were based on the male palpal morphology, which were diagnostic for
|
||
<taxonomicName id="4C2AE7E210B8939EFD89E250FD385D10" authorityName="Baert & Murphy" authorityYear="1992" box="[586,672,810,831]" class="Arachnida" family="Mysmenidae" genus="Kilifina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="254" pageNumber="781" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310B8939EFD89E250FD385D10" box="[586,672,810,831]" italics="true" pageId="254" pageNumber="781">Kilifina</emphasis>
|
||
</taxonomicName>
|
||
. As the male and female genitalia of these two species are identical in their general morphology, the differences between them are here attributed to be at species, and not generic, level. Interestingly, the only diagnostic feature shared between the current and the previous diagnoses for
|
||
<taxonomicName id="4C2AE7E210B8939EFF38E29BFEA85DD9" authorityName="Griswold" authorityYear="1985" box="[251,304,993,1014]" class="Arachnida" family="Mysmenidae" genus="Isela" kingdom="Animalia" order="Araneae" pageId="254" pageNumber="781" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310B8939EFF38E29BFEA85DD9" box="[251,304,993,1014]" italics="true" pageId="254" pageNumber="781">Isela</emphasis>
|
||
</taxonomicName>
|
||
(including
|
||
<taxonomicName id="4C2AE7E210B8939EFE73E29BFD9D5DD9" authorityName="Baert & Murphy" authorityYear="1992" box="[432,517,993,1014]" class="Arachnida" family="Mysmenidae" genus="Kilifina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="254" pageNumber="781" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E407310B8939EFE73E29BFD9D5DD9" box="[432,517,993,1014]" italics="true" pageId="254" pageNumber="781">Kilifina</emphasis>
|
||
</taxonomicName>
|
||
) are the strong setae on the male palpal tibia. Remaining characteristics are shared in varying degrees with other mysmenid genera, and are not diagnostic of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |