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<mods:title id="DF8C02F732A00DB27909916C244E955A">Out of the twilight zone: phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea)</mods:title>
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<treatment id="03832D7710B99260FEC7E245FDD05B24" ID-DOI="http://doi.org/10.5281/zenodo.7007134" ID-GBIF-Taxon="199502760" ID-Zenodo-Dep="7007134" LSID="urn:lsid:plazi:treatment:03832D7710B99260FEC7E245FDD05B24" httpUri="http://treatment.plazi.org/id/03832D7710B99260FEC7E245FDD05B24" lastPageId="256" lastPageNumber="782" pageId="255" pageNumber="782">
<subSubSection id="C330CFEA10B9939FFEC7E245FDBE5D5B" pageId="255" pageNumber="782" type="nomenclature">
<paragraph id="8B959C6110B9939FFEC7E245FDBE5D5B" blockId="255.[260,682,831,884]" pageId="255" pageNumber="782">
<heading id="D0DD2B0D10B9939FFEC7E245FDBE5D5B" centered="true" fontSize="8" level="2" pageId="255" pageNumber="782" reason="8">
<taxonomicName id="4C2AE7E210B9939FFEC7E245FD325D7B" authority="PETRUNKEVITCH, 1928" authorityName="Petrunkevitch" authorityYear="1928" box="[260,682,831,852]" class="Arachnida" family="Mysmenidae" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="subFamily" subFamily="Mysmeninae">MYSMENINAE PETRUNKEVITCH, 1928</taxonomicName>
(CLADE C133)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330CFEA10B9939FFF67E2E7FA9258DB" pageId="255" pageNumber="782" type="discussion">
<paragraph id="8B959C6110B9939FFF67E2E7FEAA5D9D" blockId="255.[164,779,925,1681]" box="[164,306,925,946]" pageId="255" pageNumber="782">
<emphasis id="B95E407310B9939FFF67E2E7FEAA5D9D" box="[164,306,925,946]" italics="true" pageId="255" pageNumber="782">Composition</emphasis>
</paragraph>
<paragraph id="8B959C6110B9939FFF67E2C1FD6158BF" blockId="255.[164,779,925,1681]" pageId="255" pageNumber="782">
This morphologically distinct subfamily is circumscribed here to comprise the following genera: 
<taxonomicName id="4C2AE7E210B9939FFF67E283FEB15A21" authorityName="Baert" authorityYear="1986" box="[164,297,1017,1038]" class="Arachnida" family="Mysmenidae" genus="Anjouanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFF67E283FEB15A21" box="[164,297,1017,1038]" italics="true" pageId="255" pageNumber="782">Anjouanella</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFEF4E283FE585A21" authorityName="Wunderlich" authorityYear="1995" box="[311,448,1017,1038]" class="Arachnida" family="Mysmenidae" genus="Brasilionata" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFEF4E283FE585A21" box="[311,448,1017,1038]" italics="true" pageId="255" pageNumber="782">Brasilionata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFE0DE283FDCF5A21" authorityName="Gertsch &amp; Davis" authorityYear="1936" box="[462,599,1017,1038]" class="Arachnida" family="Mysmenidae" genus="Calodipoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFE0DE283FDCF5A21" box="[462,599,1017,1038]" italics="true" pageId="255" pageNumber="782">Calodipoena</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFDA6E283FC9C5A21" authorityName="Baert &amp; Maelfait" authorityYear="1983" box="[613,772,1017,1038]" class="Arachnida" family="Mysmenidae" genus="Calomyspoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFDA6E283FC9C5A21" box="[613,772,1017,1038]" italics="true" pageId="255" pageNumber="782">Calomyspoena</emphasis>
</taxonomicName>
, 
<emphasis id="B95E407310B9939FFF67E562FF6C5A02" box="[164,244,1048,1069]" italics="true" pageId="255" pageNumber="782">
<collectingRegion id="49EE528310B9939FFF67E562FF6C5A02" box="[164,244,1048,1069]" country="Paraguay" name="Itapua" pageId="255" pageNumber="782">Itapua</collectingRegion>
</emphasis>
, 
<taxonomicName id="4C2AE7E210B9939FFECBE56DFE5F5A03" authorityName="Baert" authorityYear="1982" box="[264,455,1047,1068]" class="Arachnida" family="Mysmenidae" genus="Kekenboschiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFECBE56DFE5F5A03" box="[264,455,1047,1068]" italics="true" pageId="255" pageNumber="782">Kekenboschiella</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFE18E56DFDE65A03" authorityName="Banks" authorityYear="1895" box="[475,638,1047,1068]" class="Arachnida" family="Mysmenidae" genus="Microdipoena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFE18E56DFDE65A03" box="[475,638,1047,1068]" italics="true" pageId="255" pageNumber="782">Microdipoena</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFD51E562FC9B5A02" authorityName="Simon" authorityYear="1894" box="[658,771,1048,1069]" class="Arachnida" family="Mysmenidae" genus="Mysmena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFD51E562FC9B5A02" box="[658,771,1048,1069]" italics="true" pageId="255" pageNumber="782">Mysmena</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFF67E54CFEB05A64" authorityName="Brignoli" authorityYear="1980" box="[164,296,1078,1099]" class="Arachnida" family="Mysmenidae" genus="Mysmenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFF67E54CFEB05A64" box="[164,296,1078,1099]" italics="true" pageId="255" pageNumber="782">Mysmenella</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFEF5E54CFE235A64" authorityName="Thaler" authorityYear="1995" box="[310,443,1078,1099]" class="Arachnida" family="Mysmenidae" genus="Mysmeniola" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFEF5E54CFE235A64" box="[310,443,1078,1099]" italics="true" pageId="255" pageNumber="782">Mysmeniola</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C2AE7E210B9939FFE38E54CFDE85A64" authorityName="Marples" authorityYear="1955" box="[507,624,1078,1099]" class="Arachnida" family="Mysmenidae" genus="Tamasesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFE38E54CFDE85A64" box="[507,624,1078,1099]" italics="true" pageId="255" pageNumber="782">Tamasesia</emphasis>
</taxonomicName>
[
<figureCitation id="131180E410B9939FFDBCE54CFD435A63" box="[639,731,1078,1100]" captionStart="Figure 160" captionStartId="192.[144,223,198,217]" captionTargetId="figure-0@191.[217,1391,194,1815]" captionTargetPageId="191" captionText="Figure 160. Phylogenetic hypothesis rendered by the complete morphological and molecular data set analysed under equal-weights parsimony using the dynamic homology criterion. Cladogram taken from Lopardo et al. (2011: fig. 12). Numbers below each node indicate node numbers. Note that the micropholcommatine Taphiassa punctata and the theridiosomatid Coddingtonia euryopoides were referred to as Parapua punctata and TSMD-002-THAI (respectively) in Lopardo et al. (2011). Family codes used for unidentified species are as follows: ANAP, Anapidae; MYSM, Mysmenidae; SYMP, Symphytognathidae; TSMD, Theridiosomatidae." figureDoi="http://doi.org/10.5281/zenodo.7004753" httpUri="https://zenodo.org/record/7004753/files/figure.png" pageId="255" pageNumber="782">Fig. 160</figureCitation>
, see below and refer to 
<figureCitation id="131180E410B9939FFEB4E52EFE7A5A45" box="[375,482,1108,1130]" captionStart="Figure 161" captionStartId="192.[144,221,1256,1275]" captionTargetBox="[144,1424,433,1225]" captionTargetId="figure-272@192.[144,1425,474,1226]" captionTargetPageId="192" captionText="Figure 161. Hypothesis of relationships within Mysmenidae, taken from the results of the combined analysis from Lopardo et al. (2011) (see Fig. 160). New taxonomic (i.e. generic) status as proposed in our study is assigned to each representative; new combinations are emphasized in bold. Unambiguous morphological character optimizations are shown for every branch in the tree. Numbers below each node indicate node numbers. Empty and filled boxes represent homoplasious and non-homoplasious transformations, respectively. Major clades representing taxonomic decisions discussed in the text are highlighted in grey boxes." figureDoi="http://doi.org/10.5281/zenodo.5371617" httpUri="https://zenodo.org/record/5371617/files/figure.png" pageId="255" pageNumber="782">Fig. 161B</figureCitation>
for synonymies, see below for comments on recently erected Chinese genera not included in the analyses; see main text for the removal and suggested new familial status of 
<taxonomicName id="4C2AE7E210B9939FFDAAE5CAFC9B5AEA" authorityName="Wunderlich" authorityYear="1995" box="[617,771,1200,1221]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFDAAE5CAFC9B5AEA" box="[617,771,1200,1221]" italics="true" pageId="255" pageNumber="782">Crassignatha</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFF67E5B5FF635ACB" authorityName="Simon" authorityYear="1899" box="[164,251,1231,1252]" class="Arachnida" family="Symphytognathidae" genus="Iardinis" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFF67E5B5FF635ACB" box="[164,251,1231,1252]" italics="true" pageId="255" pageNumber="782">Iardinis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFECBE5B5FECE5ACB" authorityName="Miller" authorityYear="1970" box="[264,342,1231,1252]" class="Arachnida" family="Zodariidae" genus="Leviola" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFECBE5B5FECE5ACB" box="[264,342,1231,1252]" italics="true" pageId="255" pageNumber="782">Leviola</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C2AE7E210B9939FFE57E5B5FD945ACB" authorityName="Simon" authorityYear="1895" box="[404,524,1231,1252]" class="Arachnida" family="Mysmenidae" genus="Phricotelus" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFE57E5B5FD945ACB" box="[404,524,1231,1252]" italics="true" pageId="255" pageNumber="782">Phricotelus</emphasis>
</taxonomicName>
; see Miller 
<emphasis id="B95E407310B9939FFD4FE5B5FD225ACB" box="[652,698,1231,1252]" italics="true" pageId="255" pageNumber="782">et al</emphasis>
. (2009) for the removal of 
<taxonomicName id="4C2AE7E210B9939FFE4AE597FDBB5B2D" authorityName="Wunderlich" authorityYear="1995" box="[393,547,1261,1282]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFE4AE597FDBB5B2D" box="[393,547,1261,1282]" italics="true" pageId="255" pageNumber="782">Crassignatha</emphasis>
</taxonomicName>
based on morphology; and see Rix &amp; Harvey (2010) for the transfer of 
<taxonomicName id="4C2AE7E210B9939FFF67E451FEB95B6F" authorityName="Simon" authorityYear="1880" box="[164,289,1323,1344]" class="Arachnida" family="Anapidae" genus="Taphiassa" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFF67E451FEB95B6F" box="[164,289,1323,1344]" italics="true" pageId="255" pageNumber="782">Taphiassa</emphasis>
</taxonomicName>
to 
<taxonomicName id="4C2AE7E210B9939FFE90E451FDFB5B6E" box="[339,611,1323,1345]" class="Arachnida" family="Anapidae" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="subFamily" subFamily="Micropholcommatinae">Micropholcommatinae</taxonomicName>
and its synonymy with 
<taxonomicName id="4C2AE7E210B9939FFEF4E430FE025B70" authorityName="Forster" authorityYear="1959" box="[311,410,1354,1375]" class="Arachnida" family="Anapidae" genus="Parapua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFEF4E430FE025B70" box="[311,410,1354,1375]" italics="true" pageId="255" pageNumber="782">Parapua</emphasis>
</taxonomicName>
]. In our phylogenetic analyses, 
<taxonomicName id="4C2AE7E210B9939FFF67E412FEA95B51" authorityName="Petrunkevitch" authorityYear="1928" box="[164,305,1384,1406]" class="Arachnida" family="Mysmemidae" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="subFamily" subFamily="Mysmeninae">Mysmeninae</taxonomicName>
included the following taxa: 
<taxonomicName id="4C2AE7E210B9939FFDACE412FC9C5B52" authorityName="Banks" authorityYear="1895" box="[623,772,1384,1405]" class="Arachnida" family="Mysmenidae" genus="Microdipoena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFDACE412FC9C5B52" box="[623,772,1384,1405]" italics="true" pageId="255" pageNumber="782">Microdipoena</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFF67E4FCFEB05BB4" authorityName="Thaler" authorityYear="1995" box="[164,296,1414,1435]" class="Arachnida" family="Mysmenidae" genus="Mysmeniola" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFF67E4FCFEB05BB4" box="[164,296,1414,1435]" italics="true" pageId="255" pageNumber="782">Mysmeniola</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFEF6E4FCFE255BB4" authorityName="Wunderlich" authorityYear="1995" box="[309,445,1414,1435]" class="Arachnida" family="Mysmenidae" genus="Brasilionata" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFEF6E4FCFE255BB4" box="[309,445,1414,1435]" italics="true" pageId="255" pageNumber="782">Brasilionata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFE08E4FDFDAC5BB3" authorityName="Simon" authorityYear="1894" box="[459,564,1415,1436]" class="Arachnida" family="Mysmenidae" genus="Mysmena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFE08E4FDFDAC5BB3" box="[459,564,1415,1436]" italics="true" pageId="255" pageNumber="782">Mysmena</emphasis>
</taxonomicName>
(see below for total number of species and generic composition), and the following undescribed species (regarded here as 
<taxonomicName id="4C2AE7E210B9939FFF67E498FEAB5BD7" authorityName="Petrunkevitch" authorityYear="1928" box="[164,307,1506,1528]" class="Arachnida" family="Mysmemidae" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="subFamily" subFamily="Mysmeninae">Mysmeninae</taxonomicName>
<emphasis id="B95E407310B9939FFEF9E499FE575BD8" box="[314,463,1506,1528]" italics="true" pageId="255" pageNumber="782">incertae sedis</emphasis>
): MYSM-001-MAD, MYSM-006-MAD, MYSM-008-ARG, MYSM-009-MAD, MYSM-012-MAD, MYSM-019-MAD, MYSM-020-MAD, MYSM-021-MAD, MYSM-023-MAD, MYSM-026- MAD, MYSM-027-MAD, MYSM-029-MAD, MYSM-031-MAD, MYSM-032-MAD, and MYSM-033-MAD.
</paragraph>
<paragraph id="8B959C6110B9939FFF67E7BAFEB958FA" blockId="255.[164,779,1728,1903]" box="[164,289,1728,1749]" pageId="255" pageNumber="782">
<emphasis id="B95E407310B9939FFF67E7BAFEB958FA" box="[164,289,1728,1749]" italics="true" pageId="255" pageNumber="782">Monophyly</emphasis>
</paragraph>
<paragraph id="8B959C6110B9939FFF67E7A4FA9258DB" blockId="255.[164,779,1728,1903]" lastBlockId="255.[826,1442,197,1780]" pageId="255" pageNumber="782">
Relationships within 
<taxonomicName id="4C2AE7E210B9939FFE50E7A4FDBA58DB" authorityName="Petrunkevitch" authorityYear="1928" box="[403,546,1758,1780]" class="Arachnida" family="Mysmemidae" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="subFamily" subFamily="Mysmeninae">Mysmeninae</taxonomicName>
are unstable, except for a few taxa (see below). The following synapomorphies support 
<taxonomicName id="4C2AE7E210B9939FFEC9E661FE38591E" authorityName="Petrunkevitch" authorityYear="1928" box="[266,416,1819,1841]" class="Arachnida" family="Mysmemidae" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="subFamily" subFamily="Mysmeninae">Mysmeninae</taxonomicName>
as we have circumscribed it: respiratory system with anterior tracheae restricted to opisthosoma (
<figureCitation id="131180E410B9939FFE47E623FE695941" box="[388,497,1881,1903]" captionStart="Figure 37" captionStartId="54.[144,224,1265,1284]" captionTargetBox="[144,1425,195,1235]" captionTargetId="figure-266@54.[144,1425,195,1235]" captionTargetPageId="54" captionText="Figure 37. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; female abdomen; A, epigynal area and scapus, ventral view; B, left posterior spinnerets (PS); C, same as A, posterolateral view; D, digested abdomen, detail of vulva; E, right PS. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371439" httpUri="https://zenodo.org/record/5371439/files/figure.png" pageId="255" pageNumber="782">Fig. 37A</figureCitation>
; extending into prosoma in MYSM-005-ARG, MYSM-007- 
<collectingCountry id="F33DDCF110B9939FFAE2E1BCFAFA5EF4" box="[1313,1378,198,219]" name="Mexico" pageId="255" pageNumber="782">MEX</collectingCountry>
, and 
<emphasis id="B95E407310B9939FFCF9E19EFB8A5ED5" box="[826,1042,228,250]" italics="true" pageId="255" pageNumber="782">
<taxonomicName id="4C2AE7E210B9939FFCF9E19EFC7A5ED6" authorityName="Banks" authorityYear="1895" box="[826,994,228,249]" class="Arachnida" family="Mysmenidae" genus="Microdipoena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus" sensu="stricto">Microdipoena</taxonomicName>
<taxonomicNameLabel id="A26DFD0810B9939FFC28E19FFB8A5ED5" box="[1003,1042,229,250]" pageId="255" pageNumber="782" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
); advanced and wide posterior spiracular opening located midway between the spinnerets and epigastric groove (
<figureCitation id="131180E410B9939FFB12E05BFAC15F18" box="[1233,1369,289,311]" captionStart="Figure 24" captionStartId="29.[164,243,1441,1460]" captionTargetBox="[164,1445,194,1409]" captionTargetId="figure-148@29.[164,1445,194,1411]" captionTargetPageId="29" captionText="Figure 24. Microdipoena nyungwe (Mysmenidae), abdomen: A–D, female; E–G, male. A, epigastric furrow, epigynal area and spiracular openings, posterior–ventral view; B, same, detail of epigynal area; C, pedicel area and booklung covers; D, epiandrous spigots, detail from figure F; E, posterior respiratory spiracle and colulus, ventral view; F, pedicel area, booklung covers and epiandrous spigots." figureDoi="http://doi.org/10.5281/zenodo.5331665" httpUri="https://zenodo.org/record/5331665/files/figure.png" pageId="255" pageNumber="782">Fig. 24A, E</figureCitation>
), connected to branched posterior lateral tracheae extending into prosoma (
<figureCitation id="131180E410B9939FFBC4E025FB1E5F5B" box="[1031,1158,351,373]" captionStart="Figure 22" captionStartId="27.[164,243,1669,1688]" captionTargetBox="[164,1445,194,1639]" captionTargetId="figure-0@27.[164,1445,194,1639]" captionTargetPageId="27" captionText="Figure 22. Microdipoena nyungwe (Mysmenidae): A, B, E, female; C, D, F, G, male left palp. A, digested abdomen, respiratory system and vulva; B, same, detail of vulva; C, distal–retrolateral view, detail of tip of embolus and primary cymbial conductor; D, ventral–proximal view; E, tip of palp and tarsal organ; F, dorsal view; G, retrolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331661" httpUri="https://zenodo.org/record/5331661/files/figure.png" pageId="255" pageNumber="782">Fig. 22A, B</figureCitation>
); aciniform gland spigots of posterior spinnerets with two different outlines (
<figureCitation id="131180E410B9939FFC81E0E6FC3E5F9E" box="[834,934,412,434]" captionStart="Figure 19" captionStartId="23.[164,242,1627,1646]" captionTargetBox="[164,1445,194,1595]" captionTargetId="figure-82@23.[164,1445,194,1597]" captionTargetPageId="23" captionText="Figure 19. Microdipoena guttata (Mysmenidae). A, B, D–F, male; C, female. A, prosoma, frontal view; B, C, same, lateral view; D, same, ventral view, detail of labium–sternum junction; E, mouthparts, detail of tip of left chelicera; F, posterior median spinnerets. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371425" httpUri="https://zenodo.org/record/5371425/files/figure.png" pageId="255" pageNumber="782">Figs 19F</figureCitation>
, 
<figureCitation id="131180E410B9939FFC76E0E6FC7E5F9E" box="[949,998,412,433]" captionStart="Figure 33" captionStartId="50.[144,219,1787,1806]" captionTargetBox="[221,1346,194,1755]" captionTargetId="figure-0@50.[221,1346,194,1757]" captionTargetPageId="50" captionText="Figure 33. Mysmena (= Calodipoena) incredula (Mysmenidae); abdomen. A, B, female digested abdomen; C–H, male spinnerets. A, tracheal system and vulva; B, same, detail of vulva; C, spinning field, ventral–posterior view; D, posterior median spinnerets; E, right anterior lateral spinnerets (ALS); F, left ALS; G, right posterior lateral spinnerets; H, same, detail. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371435" httpUri="https://zenodo.org/record/5371435/files/figure.png" pageId="255" pageNumber="782">33D</figureCitation>
, 
<figureCitation id="131180E410B9939FFC35E0E6FB8B5F9E" box="[1014,1043,412,433]" captionStart="Figure 37" captionStartId="54.[144,224,1265,1284]" captionTargetBox="[144,1425,195,1235]" captionTargetId="figure-266@54.[144,1425,195,1235]" captionTargetPageId="54" captionText="Figure 37. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; female abdomen; A, epigynal area and scapus, ventral view; B, left posterior spinnerets (PS); C, same as A, posterolateral view; D, digested abdomen, detail of vulva; E, right PS. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371439" httpUri="https://zenodo.org/record/5371439/files/figure.png" pageId="255" pageNumber="782">37</figureCitation>
,B E); posterior lateral spinnerets with an anterior flat spatulate modified seta [
<figureCitation id="131180E410B9939FFAF4E0C0FCD75FC1" captionStart="Figure 23" captionStartId="28.[144,223,1821,1840]" captionTargetBox="[205,1363,194,1790]" captionTargetId="figure-0@28.[205,1363,194,1791]" captionTargetPageId="28" captionText="Figure 23. Microdipoena nyungwe (Mysmenidae), spinnerets: A, B, female; C–H, male. A, C, anterior lateral spinnerets (ALS); B, E, left posterior lateral spinnerets; D, posterior median spinnerets; F, ALS, detail of major ampullate field; G, H, ALS, detail of intersegmental lobe. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331663" httpUri="https://zenodo.org/record/5331663/files/figure.png" pageId="255" pageNumber="782">Figs 23B, E</figureCitation>
, 
<figureCitation id="131180E410B9939FFCA2E0A3FC255FC1" box="[865,957,473,494]" captionStart="Figure 33" captionStartId="50.[144,219,1787,1806]" captionTargetBox="[221,1346,194,1755]" captionTargetId="figure-0@50.[221,1346,194,1757]" captionTargetPageId="50" captionText="Figure 33. Mysmena (= Calodipoena) incredula (Mysmenidae); abdomen. A, B, female digested abdomen; C–H, male spinnerets. A, tracheal system and vulva; B, same, detail of vulva; C, spinning field, ventral–posterior view; D, posterior median spinnerets; E, right anterior lateral spinnerets (ALS); F, left ALS; G, right posterior lateral spinnerets; H, same, detail. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371435" httpUri="https://zenodo.org/record/5371435/files/figure.png" pageId="255" pageNumber="782">33G, H</figureCitation>
, 
<figureCitation id="131180E410B9939FFC13E0A3FB9B5FC0" box="[976,1027,473,495]" captionStart="Figure 52" captionStartId="69.[164,243,1574,1593]" captionTargetBox="[164,1445,194,1544]" captionTargetId="figure-74@69.[164,1445,194,1544]" captionTargetPageId="69" captionText="Figure 52. A, MYSM-034-MAD (Mysmena, Mysmenidae), female; B–D, MYSM-029-MAD (Mysmeninae, Mysmenidae), female; E, F, MYSM-028-MAD (Mysmena, Mysmenidae), female. A, prosoma, lateral view; B, right anterior lateral spinneret; C, right posterior lateral spinneret; D, right leg I, claws, retrolateral view; E, chelicerae and palps, frontal view; F, epigynal area, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371467" httpUri="https://zenodo.org/record/5371467/files/figure.png" pageId="255" pageNumber="782">52C</figureCitation>
; secondarily absent in 
<taxonomicName id="4C2AE7E210B9939FFAECE0A0FA395FC0" authorityName="Simon" authorityYear="1894" box="[1327,1441,474,495]" class="Arachnida" family="Mysmenidae" genus="Mysmena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFAECE0A0FA395FC0" box="[1327,1441,474,495]" italics="true" pageId="255" pageNumber="782">Mysmena</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C2AE7E210B9939FFC9BE082FC495C22" authorityName="Marples" authorityYear="1955" box="[856,977,504,525]" class="Arachnida" family="Mysmenidae" genus="Tamasesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFC9BE082FC495C22" box="[856,977,504,525]" italics="true" pageId="255" pageNumber="782">Tamasesia</emphasis>
</taxonomicName>
) 
<taxonomicName id="4C2AE7E210B9939FFC27E082FBA75C22" box="[996,1087,504,525]" class="Arachnida" family="Mysmenidae" genus="Tamasesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="species" species="rotunda">
<emphasis id="B95E407310B9939FFC27E082FBA75C22" box="[996,1087,504,525]" italics="true" pageId="255" pageNumber="782">rotunda</emphasis>
</taxonomicName>
], and slim cylindrical spigots (
<figureCitation id="131180E410B9939FFC81E36CFC3D5C04" box="[834,933,534,556]" captionStart="Figure 23" captionStartId="28.[144,223,1821,1840]" captionTargetBox="[205,1363,194,1790]" captionTargetId="figure-0@28.[205,1363,194,1791]" captionTargetPageId="28" captionText="Figure 23. Microdipoena nyungwe (Mysmenidae), spinnerets: A, B, female; C–H, male. A, C, anterior lateral spinnerets (ALS); B, E, left posterior lateral spinnerets; D, posterior median spinnerets; F, ALS, detail of major ampullate field; G, H, ALS, detail of intersegmental lobe. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331663" httpUri="https://zenodo.org/record/5331663/files/figure.png" pageId="255" pageNumber="782">Figs 23B</figureCitation>
, 
<figureCitation id="131180E410B9939FFC73E36CFC645C03" box="[944,1020,534,556]" captionStart="Figure 37" captionStartId="54.[144,224,1265,1284]" captionTargetBox="[144,1425,195,1235]" captionTargetId="figure-266@54.[144,1425,195,1235]" captionTargetPageId="54" captionText="Figure 37. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; female abdomen; A, epigynal area and scapus, ventral view; B, left posterior spinnerets (PS); C, same as A, posterolateral view; D, digested abdomen, detail of vulva; E, right PS. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371439" httpUri="https://zenodo.org/record/5371439/files/figure.png" pageId="255" pageNumber="782">37B, E</figureCitation>
); palpal tibial trichobothria of females lacking (
<figureCitation id="131180E410B9939FFC63E34FFB915C65" box="[928,1033,565,587]" captionStart="Figure 38" captionStartId="55.[164,243,1821,1840]" captionTargetBox="[228,1381,194,1791]" captionTargetId="figure-0@55.[228,1381,194,1791]" captionTargetPageId="55" captionText="Figure 38. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; prosoma: A, B, male; C–H, female; A–C, lateral view; D, carapace, frontal view; E, chelicerae and palps, frontal view; F, mouthparts, ventral view; G, right palp, detail of palpal tip; H, distal promargin of right chelicera." figureDoi="http://doi.org/10.5281/zenodo.5371441" httpUri="https://zenodo.org/record/5371441/files/figure.png" pageId="255" pageNumber="782">Figs 38E</figureCitation>
, 
<figureCitation id="131180E410B9939FFBDBE34FFBD05C64" box="[1048,1096,565,587]" captionStart="Figure 52" captionStartId="69.[164,243,1574,1593]" captionTargetBox="[164,1445,194,1544]" captionTargetId="figure-74@69.[164,1445,194,1544]" captionTargetPageId="69" captionText="Figure 52. A, MYSM-034-MAD (Mysmena, Mysmenidae), female; B–D, MYSM-029-MAD (Mysmeninae, Mysmenidae), female; E, F, MYSM-028-MAD (Mysmena, Mysmenidae), female. A, prosoma, lateral view; B, right anterior lateral spinneret; C, right posterior lateral spinneret; D, right leg I, claws, retrolateral view; E, chelicerae and palps, frontal view; F, epigynal area, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371467" httpUri="https://zenodo.org/record/5371467/files/figure.png" pageId="255" pageNumber="782">52E</figureCitation>
); female copulatory openings within the epigastric furrow (
<figureCitation id="131180E410B9939FFB49E329FB745C47" box="[1162,1260,595,617]" captionStart="Figure 24" captionStartId="29.[164,243,1441,1460]" captionTargetBox="[164,1445,194,1409]" captionTargetId="figure-148@29.[164,1445,194,1411]" captionTargetPageId="29" captionText="Figure 24. Microdipoena nyungwe (Mysmenidae), abdomen: A–D, female; E–G, male. A, epigastric furrow, epigynal area and spiracular openings, posterior–ventral view; B, same, detail of epigynal area; C, pedicel area and booklung covers; D, epiandrous spigots, detail from figure F; E, posterior respiratory spiracle and colulus, ventral view; F, pedicel area, booklung covers and epiandrous spigots." figureDoi="http://doi.org/10.5281/zenodo.5331665" httpUri="https://zenodo.org/record/5331665/files/figure.png" pageId="255" pageNumber="782">Fig. 24A</figureCitation>
; secondarily external in MYSM-023-MAD), membranous atrium (
<figureCitation id="131180E410B9939FFC81E3EBFB995C89" box="[834,1025,657,679]" captionStart="Figure 129" captionStartId="145.[164,243,1731,1750]" captionTargetBox="[210,1396,198,1697]" captionTargetId="figure-0@145.[209,1400,195,1700]" captionTargetPageId="145" captionText="Figure 129. Mysmenidae female genitalia, cleared: A, Microdipoena guttata, ventral view; B, Microdipoena nyungwe, ventral view; C, MYSM-007-MEX (Mysmena), from Chiapas, Mexico, ventral view, arrow to diverticle of copulatory duct; D, Microdipoena elsae, ventral copulatory duct; E, Microdipoena (= Mysmenella) samoensis, syntype, ventral view; F, Microdipoena (= Mysmenella) jobi, paratype, ventral view; G, Mysmena-MYSM-015-MAD (Mysmena), from Antananarivo, Madagascar, dorsal view; H, MYSM-029-MAD (Mysmeninae), from Antsiranana, Madagascar, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371569" httpUri="https://zenodo.org/record/5371569/files/figure.png" pageId="255" pageNumber="782">Figs 129A, E, G</figureCitation>
, 
<figureCitation id="131180E410B9939FFBD3E3EBFBD55C88" box="[1040,1101,657,679]" captionStart="Figure 130" captionStartId="146.[144,222,1760,1779]" captionTargetBox="[190,1379,198,1727]" captionTargetId="figure-0@146.[182,1386,195,1730]" captionTargetPageId="146" captionText="Figure 130. Mysmenidae female genitalia, cleared. A, Mysmena (= Calodipoena) tarautensis, paratype, dorsal view, arrow to broken scapus; B, Mysmena (= Calodipoena) incredula, dorsal view; C, Mysmena leichhardti, from Queensland, Australia, ventral view; D, MYSM-023-MAD (Mysmeninae), from Antananarivo, Madagascar, ventral view; E, Mysmena tasmaniae, ventral view; F, MYSM-005-ARG (Mysmena), from Misiones, Argentina, ventral view; G, MYSM-019-MAD (Mysmeninae), from Toliara, Madagascar, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371571" httpUri="https://zenodo.org/record/5371571/files/figure.png" pageId="255" pageNumber="782">130B</figureCitation>
), and irregular membranous copulatory ducts (
<figureCitation id="131180E410B9939FFBC1E3D5FBF15CEB" box="[1026,1129,687,709]" captionStart="Figure 18" captionStartId="22.[144,222,1539,1558]" captionTargetBox="[144,1424,194,1509]" captionTargetId="figure-83@22.[144,1424,194,1510]" captionTargetPageId="22" captionText="Figure 18. Microdipoena guttata (Mysmenidae): A–F, male left palp; G, female. A, ventral–proximal view; B, retrolateral– distal view; C, dorsal–retrolateral view; D, expanded bulb, prolateral view; E, prolateral view; F, expanded bulb, detail of tip of embolus, retrolateral–distal view; G, digested abdomen, detail of vulva. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331657" httpUri="https://zenodo.org/record/5331657/files/figure.png" pageId="255" pageNumber="782">Figs 18G</figureCitation>
, 
<figureCitation id="131180E410B9939FFBB7E3D5FB3C5CEB" box="[1140,1188,687,708]" captionStart="Figure 27" captionStartId="32.[144,223,1752,1771]" captionTargetBox="[208,1361,194,1721]" captionTargetId="figure-0@32.[208,1361,194,1722]" captionTargetPageId="32" captionText="Figure 27. Microdipoena (= Mysmenella) samoensis (Mysmenidae), syntypes: A–C, F, H, I, male; D, E, G, female. A, left palp, prolateral–ventral view; B, same, retrolateral–ventral view; C, same, detail of tip of embolus and interaction with primary cymbial conductor, prolateral view; D, digested abdomen, vulva, and part of respiratory system; E, same, detail of vulva; F, G, prosoma, lateral view; H, mouthparts, ventral view, detail of retromargin of chelicerae and labium; I, left tibia I, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331671" httpUri="https://zenodo.org/record/5331671/files/figure.png" pageId="255" pageNumber="782">27D</figureCitation>
, 
<figureCitation id="131180E410B9939FFB73E3D5FAD55CEA" box="[1200,1357,687,709]" captionStart="Figure 129" captionStartId="145.[164,243,1731,1750]" captionTargetBox="[210,1396,198,1697]" captionTargetId="figure-0@145.[209,1400,195,1700]" captionTargetPageId="145" captionText="Figure 129. Mysmenidae female genitalia, cleared: A, Microdipoena guttata, ventral view; B, Microdipoena nyungwe, ventral view; C, MYSM-007-MEX (Mysmena), from Chiapas, Mexico, ventral view, arrow to diverticle of copulatory duct; D, Microdipoena elsae, ventral copulatory duct; E, Microdipoena (= Mysmenella) samoensis, syntype, ventral view; F, Microdipoena (= Mysmenella) jobi, paratype, ventral view; G, Mysmena-MYSM-015-MAD (Mysmena), from Antananarivo, Madagascar, dorsal view; H, MYSM-029-MAD (Mysmeninae), from Antsiranana, Madagascar, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371569" httpUri="https://zenodo.org/record/5371569/files/figure.png" pageId="255" pageNumber="782">129A–C, E, H</figureCitation>
, 
<figureCitation id="131180E410B9939FFA99E3D5FCC85CCC" captionStart="Figure 130" captionStartId="146.[144,222,1760,1779]" captionTargetBox="[190,1379,198,1727]" captionTargetId="figure-0@146.[182,1386,195,1730]" captionTargetPageId="146" captionText="Figure 130. Mysmenidae female genitalia, cleared. A, Mysmena (= Calodipoena) tarautensis, paratype, dorsal view, arrow to broken scapus; B, Mysmena (= Calodipoena) incredula, dorsal view; C, Mysmena leichhardti, from Queensland, Australia, ventral view; D, MYSM-023-MAD (Mysmeninae), from Antananarivo, Madagascar, ventral view; E, Mysmena tasmaniae, ventral view; F, MYSM-005-ARG (Mysmena), from Misiones, Argentina, ventral view; G, MYSM-019-MAD (Mysmeninae), from Toliara, Madagascar, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371571" httpUri="https://zenodo.org/record/5371571/files/figure.png" pageId="255" pageNumber="782">130A– G</figureCitation>
; distally sclerotized independently in two clades); males with coiled embolus (
<figureCitation id="131180E410B9939FFB47E396FB745D2E" box="[1156,1260,748,770]" captionStart="Figure 27" captionStartId="32.[144,223,1752,1771]" captionTargetBox="[208,1361,194,1721]" captionTargetId="figure-0@32.[208,1361,194,1722]" captionTargetPageId="32" captionText="Figure 27. Microdipoena (= Mysmenella) samoensis (Mysmenidae), syntypes: A–C, F, H, I, male; D, E, G, female. A, left palp, prolateral–ventral view; B, same, retrolateral–ventral view; C, same, detail of tip of embolus and interaction with primary cymbial conductor, prolateral view; D, digested abdomen, vulva, and part of respiratory system; E, same, detail of vulva; F, G, prosoma, lateral view; H, mouthparts, ventral view, detail of retromargin of chelicerae and labium; I, left tibia I, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331671" httpUri="https://zenodo.org/record/5331671/files/figure.png" pageId="255" pageNumber="782">Figs 27A</figureCitation>
, 
<figureCitation id="131180E410B9939FFB38E396FAD65D2D" box="[1275,1358,748,770]" captionStart="Figure 47" captionStartId="64.[144,221,1188,1207]" captionTargetBox="[144,1424,194,1158]" captionTargetId="figure-244@64.[144,1424,194,1158]" captionTargetPageId="64" captionText="Figure 47. MYSM-020-MAD (Mysmeninae, Mysmenidae) from Toamasina, Madagascar, male. A–E, left palp; A, retrolateral view; B, prolateral–ventral view; C, same, detail of tip of embolus and cymbial conductors; D, same, ventral–retrolateral view; E, same, prolateral view. F, digested abdomen, tracheal system, dorsal view; G, same, detail of posterior tracheal system. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331683" httpUri="https://zenodo.org/record/5331683/files/figure.png" pageId="255" pageNumber="782">47A, B</figureCitation>
, 
<figureCitation id="131180E410B9939FFA9DE396FCD65D0F" captionStart="Figure 132" captionStartId="148.[144,222,1638,1657]" captionTargetBox="[149,1421,202,1603]" captionTargetId="figure-0@148.[144,1424,193,1607]" captionTargetPageId="148" captionText="Figure 132. Mysmenidae male left palp, cleared: A, Microdipoena elsae, prolateral view, arrow to spine of basal prolateral cymbial expansion; B, Microdipoena guttata, prolateral view, arrow to spine of basal prolateral cymbial expansion; C, Microdipoena (= Mysmenella) illectrix, type, prolateral view; D, Microdipoena (= Mysmenella) samoensis, syntype, prolateral view; E, same, retrolateral view; F, Microdipoena (= Mysmenella) jobi, holotype, retrolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331805" httpUri="https://zenodo.org/record/5331805/files/figure.png" pageId="255" pageNumber="782">132D, E</figureCitation>
, 
<figureCitation id="131180E410B9939FFC9FE271FC005D0E" box="[860,920,779,801]" captionStart="Figure 134" captionStartId="150.[144,223,1736,1755]" captionTargetBox="[194,1375,199,1701]" captionTargetId="figure-0@150.[189,1380,195,1706]" captionTargetPageId="150" captionText="Figure 134. Mysmenidae male left palp (unless otherwise stated), cleared: A, Mysmena leucoplagiata, type, prolateral view, right palp, inverted; B, Mysmena leichhardti, from Queensland, Australia, prolateral–ventral view; C, Mysmena tasmaniae, prolateral–ventral view, right palp, inverted; D, Mysmeniola spinifera, holotype, prolateral view, right palp, inverted; E, MYSM-019-MAD (Mysmeninae), from Toliara, Madagascar, prolateral view; F, MYSM-023-MAD (Mysmeninae), from Antananarivo, Madagascar, retrolateral view, right palp, inverted; G, MYSM-020-MAD (Mysmeninae), from Toamasina, Madagascar, dorsal view, right palp, inverted; H, same, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331811" httpUri="https://zenodo.org/record/5331811/files/figure.png" pageId="255" pageNumber="782">134G</figureCitation>
) and secondary (external) cymbial conductor (
<figureCitation id="131180E410B9939FFC81E253FC3F5D10" box="[834,935,809,831]" captionStart="Figure 31" captionStartId="47.[164,243,1483,1502]" captionTargetBox="[164,1444,194,1451]" captionTargetId="figure-129@47.[164,1444,194,1453]" captionTargetPageId="47" captionText="Figure 31. MYSM-007-MEX (Mysmena, Mysmenidae) from Chiapas, Mexico; A–D, male left palp; E–G, female. A, detail of proximal palp, prolateral basal expansion and paracymbium, retroventral view; B, detail of dorsal–retrolateral tip of cymbium; C, same, prolateral view; D, same, retrolateral view; E, cheliceral bases, frontal view; F, abdomen, lateral view; G, abdomen, lateral view, detail of scapus and spinnerets. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371431" httpUri="https://zenodo.org/record/5371431/files/figure.png" pageId="255" pageNumber="782">Figs 31C</figureCitation>
, 
<figureCitation id="131180E410B9939FFC70E250FC7B5D10" box="[947,995,810,831]" captionStart="Figure 40" captionStartId="57.[164,242,1534,1553]" captionTargetBox="[164,1445,195,1504]" captionTargetId="figure-130@57.[164,1445,195,1504]" captionTargetPageId="57" captionText="Figure 40. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; male left palp, details. A, ventral-distal view; B, prolateral view; C, same, close up; D, ventral view; E, retrolateral view; F, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371445" httpUri="https://zenodo.org/record/5371445/files/figure.png" pageId="255" pageNumber="782">40A</figureCitation>
, 
<figureCitation id="131180E410B9939FFC2DE250FB855D10" box="[1006,1053,810,831]" captionStart="Figure 43" captionStartId="60.[144,223,1073,1092]" captionTargetBox="[144,1424,195,1043]" captionTargetId="figure-317@60.[144,1424,195,1043]" captionTargetPageId="60" captionText="Figure 43. Mysmena leichhardti (Mysmenidae) from Queensland, Australia; male left palp, details. A, B, embolus– conductor interaction; C, distal palp, distal–prolateral view; D, dorsal cymbial tip; E, ventral–prolateral cymbial tip. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371451" httpUri="https://zenodo.org/record/5371451/files/figure.png" pageId="255" pageNumber="782">43C</figureCitation>
; secondarily absent in clade C128: 
<taxonomicName id="4C2AE7E210B9939FFCF9E232FC4C5D72" authorityName="Banks" authorityYear="1895" box="[826,980,840,861]" class="Arachnida" family="Mysmenidae" genus="Microdipoena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFCF9E232FC4C5D72" box="[826,980,840,861]" italics="true" pageId="255" pageNumber="782">Microdipoena</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFC20E232FBE95D72" authorityName="Wunderlich" authorityYear="1995" box="[995,1137,840,861]" class="Arachnida" family="Mysmenidae" genus="Brasilionata" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFC20E232FBE95D72" box="[995,1137,840,861]" italics="true" pageId="255" pageNumber="782">Brasilionata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E210B9939FFB43E232FA905D72" authorityName="Thaler" authorityYear="1995" box="[1152,1288,840,861]" class="Arachnida" family="Mysmenidae" genus="Mysmeniola" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFB43E232FA905D72" box="[1152,1288,840,861]" italics="true" pageId="255" pageNumber="782">Mysmeniola</emphasis>
</taxonomicName>
, and MYSM-019-MAD; 
<figureCitation id="131180E410B9939FFC79E21DFB845D52" box="[954,1052,871,893]" captionStart="Figure 22" captionStartId="27.[164,243,1669,1688]" captionTargetBox="[164,1445,194,1639]" captionTargetId="figure-0@27.[164,1445,194,1639]" captionTargetPageId="27" captionText="Figure 22. Microdipoena nyungwe (Mysmenidae): A, B, E, female; C, D, F, G, male left palp. A, digested abdomen, respiratory system and vulva; B, same, detail of vulva; C, distal–retrolateral view, detail of tip of embolus and primary cymbial conductor; D, ventral–proximal view; E, tip of palp and tarsal organ; F, dorsal view; G, retrolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331661" httpUri="https://zenodo.org/record/5331661/files/figure.png" pageId="255" pageNumber="782">Fig. 22F</figureCitation>
). Other synapomorphies include: abdomen with a whitish ventral ring surrounding the spinnerets (
<figureCitation id="131180E410B9939FFC78E2DEFBDC5D96" box="[955,1092,932,954]" captionStart="Figure 142" captionStartId="158.[144,223,1704,1723]" captionTargetBox="[194,1372,193,1673]" captionTargetId="figure-0@158.[194,1372,193,1674]" captionTargetPageId="158" captionText="Figure 142. Composite images of Mysmenidae species: Mysmeninae and Trogloneta. A, B, Microdipoena nyungwe; A, male, lateral view; B, female, ventral view. C, Trogloneta granulum, female, ventral view. D, Mysmena (= Calodipoena) incredula, female, lateral view, inverted. E, Mysmena (= Calodipoena) mootae, female holotype, dorsal view. F, Mysmena (= Calomyspoena) santacruzi, female paratype, dorsal view. G–I, Mysmena (= Itapua) tembei; G, male holotype, lateral view, inverted; H, male holotype, frontal view; I, female paratype, frontal view. J–L, Mysmena (= Kekenboschiella) awari, male paratype; J, lateral view; K, dorsal view; L, ventral view. M, N, Mysmeniola spinifera, male holotype; M, lateral view; N, ventral view. O, Brasilionata arborense, male holotype, dorsal view, abdomen detached. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5371585" httpUri="https://zenodo.org/record/5371585/files/figure.png" pageId="255" pageNumber="782">Figs 142J, L</figureCitation>
, 
<figureCitation id="131180E410B9939FFB93E2DEFB675D96" box="[1104,1279,932,953]" captionStart="Figure 143" captionStartId="159.[164,243,1736,1755]" captionTargetBox="[202,1406,194,1706]" captionTargetId="figure-0@159.[202,1406,194,1706]" captionTargetPageId="159" captionText="Figure 143. Composite images of Mysmeninae species. A–C, Mysmena tasmaniae; A, female, lateral view; B, female, ventral view; C, male, ventral view. D–F, MYSM-019-MAD (Mysmeninae); D, female, lateral view; E, female, ventral view; F, male, lateral view. G–I, MYSM-005-ARG (Mysmena), female; G, lateral view; H, dorsal view; I, ventral view. J–L, MYSM-007-MEX (Mysmena); J, female, lateral view; K, female, dorsal view; L, male, ventral view. M, Mysmena- MYSM-015-MAD (Mysmena), female, lateral view. N, Mysmena leichhardti, female, lateral view. O, MYSM-028-MAD (Mysmena), female prosoma, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5371587" httpUri="https://zenodo.org/record/5371587/files/figure.png" pageId="255" pageNumber="782">143A, B, G, I, L</figureCitation>
), trichobothria on tibia III and IV between two and three tibia diameters in length (
<figureCitation id="131180E410B9939FFB81E29BFB2E5DD8" box="[1090,1206,993,1015]" captionStart="Figure 26" captionStartId="31.[164,242,1513,1532]" captionTargetBox="[164,1445,194,1482]" captionTargetId="figure-138@31.[164,1445,194,1482]" captionTargetPageId="31" captionText="Figure 26. Microdipoena nyungwe (Mysmenidae), left legs: A, C, D, G, male; B, E, F, female. A, tarsus I, prolateral view; B, claws I, retrolateral view; C, leg I, tibia–metatarsal junction, prolateral view; D, metatarsus I, dorsal view; E, leg I, metatarsus–tarsus junction, dorsal view; F, tarsus IV, prolateral view; G, tibia IV, prolateral view." figureDoi="http://doi.org/10.5281/zenodo.5331669" httpUri="https://zenodo.org/record/5331669/files/figure.png" pageId="255" pageNumber="782">Figs 26G</figureCitation>
, 
<figureCitation id="131180E410B9939FFB0FE29BFA985DD9" box="[1228,1280,993,1014]" captionStart="Figure 29" captionStartId="35.[164,243,1555,1574]" captionTargetBox="[164,1445,194,1525]" captionTargetId="figure-71@35.[164,1445,194,1525]" captionTargetPageId="35" captionText="Figure 29. MYSM-005-ARG (Mysmena, Mysmenidae) from Misiones, Argentina. A–C, E, F, female; D, male. A, digested abdomen, detail of left spermatheca; B, same, detail of posterior tracheal system; C, epigynal area and scapus; D, left femur I, retrolateral view; E, left tibia IV, prolateral view; F, left metatarsus IV, prolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331675" httpUri="https://zenodo.org/record/5331675/files/figure.png" pageId="255" pageNumber="782">29E</figureCitation>
, 
<figureCitation id="131180E410B9939FFAD5E29BFAD25DD9" box="[1302,1354,993,1014]" captionStart="Figure 39" captionStartId="56.[144,222,1789,1808]" captionTargetBox="[189,1380,194,1760]" captionTargetId="figure-0@56.[189,1380,194,1760]" captionTargetPageId="56" captionText="Figure 39. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; abdomen and left legs: A, C, male; B, D–J, female. A, abdomen, ventral view; B, booklung cover; C, leg I, tarsal organ, retrolateral view; D, femur I, retrolateral view; E, metatarsus I, retrolateral view; F, femur IV, retrolateral view; G, tibia IV, retrolateral view; H, tarsus I, retrolateral view; I, tarsus IV, retrolateral view; J, leg IV, metatarsus–tarsus junction, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5371443" httpUri="https://zenodo.org/record/5371443/files/figure.png" pageId="255" pageNumber="782">39G</figureCitation>
; short trichobothria occurs independently within the group), and strongly serrated distal promarginal curved seta (
<figureCitation id="131180E410B9939FFC81E547FC345A7D" box="[834,940,1085,1107]" captionStart="Figure 19" captionStartId="23.[164,242,1627,1646]" captionTargetBox="[164,1445,194,1595]" captionTargetId="figure-82@23.[164,1445,194,1597]" captionTargetPageId="23" captionText="Figure 19. Microdipoena guttata (Mysmenidae). A, B, D–F, male; C, female. A, prosoma, frontal view; B, C, same, lateral view; D, same, ventral view, detail of labium–sternum junction; E, mouthparts, detail of tip of left chelicera; F, posterior median spinnerets. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371425" httpUri="https://zenodo.org/record/5371425/files/figure.png" pageId="255" pageNumber="782">Figs 19E</figureCitation>
, 
<figureCitation id="131180E410B9939FFC7EE547FC685A7D" box="[957,1008,1085,1106]" captionStart="Figure 38" captionStartId="55.[164,243,1821,1840]" captionTargetBox="[228,1381,194,1791]" captionTargetId="figure-0@55.[228,1381,194,1791]" captionTargetPageId="55" captionText="Figure 38. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; prosoma: A, B, male; C–H, female; A–C, lateral view; D, carapace, frontal view; E, chelicerae and palps, frontal view; F, mouthparts, ventral view; G, right palp, detail of palpal tip; H, distal promargin of right chelicera." figureDoi="http://doi.org/10.5281/zenodo.5371441" httpUri="https://zenodo.org/record/5371441/files/figure.png" pageId="255" pageNumber="782">38H</figureCitation>
, 
<figureCitation id="131180E410B9939FFBC2E547FBA95A7D" box="[1025,1073,1085,1106]" captionStart="Figure 42" captionStartId="59.[164,243,1792,1811]" captionTargetBox="[196,1412,194,1762]" captionTargetId="figure-0@59.[196,1412,194,1762]" captionTargetPageId="59" captionText="Figure 42. Mysmena leichhardti (Mysmenidae) from Queensland, Australia: A, B, E, G, H, male; C, D, F, female. A, prosoma, lateral view; B, same, lateral–frontal view; C, epigynal area and scapus, ventral view; D, digested abdomen, detail of vulva, tracheae removed; E, mouthparts, distal view; right femur I, prolateral–dorsal view; G, left femur I, ventral view; H, left metatarsus I, prolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371449" httpUri="https://zenodo.org/record/5371449/files/figure.png" pageId="255" pageNumber="782">42E</figureCitation>
, 
<figureCitation id="131180E410B9939FFB81E547FBEB5A7C" box="[1090,1139,1085,1107]" captionStart="Figure 48" captionStartId="65.[164,243,1558,1577]" captionTargetBox="[164,1445,195,1528]" captionTargetId="figure-103@65.[164,1445,195,1528]" captionTargetPageId="65" captionText="Figure 48. MYSM-020-MAD (Mysmeninae, Mysmenidae) from Toamasina, Madagascar, male: A, epiandrous spigots; B, cheliceral teeth and fang; C, D, right tibia I, detail of ventral paired setae with protruded bases, prolateral view; E, right tibia I, ventral view; F, right tarsus I, detail of tarsal organ, retrolateral view; G, right leg IV, claws." figureDoi="http://doi.org/10.5281/zenodo.5371459" httpUri="https://zenodo.org/record/5371459/files/figure.png" pageId="255" pageNumber="782">48B</figureCitation>
). Ambiguously optimized synapomorphies for this clade include: a posterior tracheal arrangement consisting of lateral tracheae surrounding the minute median apodemes (
<figureCitation id="131180E410B9939FFAEDE5E3FA0C5A80" box="[1326,1428,1177,1199]" captionStart="Figure 29" captionStartId="35.[164,243,1555,1574]" captionTargetBox="[164,1445,194,1525]" captionTargetId="figure-71@35.[164,1445,194,1525]" captionTargetPageId="35" captionText="Figure 29. MYSM-005-ARG (Mysmena, Mysmenidae) from Misiones, Argentina. A–C, E, F, female; D, male. A, digested abdomen, detail of left spermatheca; B, same, detail of posterior tracheal system; C, epigynal area and scapus; D, left femur I, retrolateral view; E, left tibia IV, prolateral view; F, left metatarsus IV, prolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331675" httpUri="https://zenodo.org/record/5331675/files/figure.png" pageId="255" pageNumber="782">Fig. 29B</figureCitation>
); fingerprint cuticular pattern on the piriform field (
<figureCitation id="131180E410B9939FFC81E5ACFC5E5AC4" box="[834,966,1238,1260]" captionStart="Figure 23" captionStartId="28.[144,223,1821,1840]" captionTargetBox="[205,1363,194,1790]" captionTargetId="figure-0@28.[205,1363,194,1791]" captionTargetPageId="28" captionText="Figure 23. Microdipoena nyungwe (Mysmenidae), spinnerets: A, B, female; C–H, male. A, C, anterior lateral spinnerets (ALS); B, E, left posterior lateral spinnerets; D, posterior median spinnerets; F, ALS, detail of major ampullate field; G, H, ALS, detail of intersegmental lobe. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331663" httpUri="https://zenodo.org/record/5331663/files/figure.png" pageId="255" pageNumber="782">Figs 23A, C</figureCitation>
, 
<figureCitation id="131180E410B9939FFC11E5ACFB985AC3" box="[978,1024,1238,1260]" captionStart="Figure 33" captionStartId="50.[144,219,1787,1806]" captionTargetBox="[221,1346,194,1755]" captionTargetId="figure-0@50.[221,1346,194,1757]" captionTargetPageId="50" captionText="Figure 33. Mysmena (= Calodipoena) incredula (Mysmenidae); abdomen. A, B, female digested abdomen; C–H, male spinnerets. A, tracheal system and vulva; B, same, detail of vulva; C, spinning field, ventral–posterior view; D, posterior median spinnerets; E, right anterior lateral spinnerets (ALS); F, left ALS; G, right posterior lateral spinnerets; H, same, detail. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371435" httpUri="https://zenodo.org/record/5371435/files/figure.png" pageId="255" pageNumber="782">33E</figureCitation>
); flagelliform spigots absent in males (
<figureCitation id="131180E410B9939FFC81E58EFC3C5B25" box="[834,932,1268,1290]" captionStart="Figure 23" captionStartId="28.[144,223,1821,1840]" captionTargetBox="[205,1363,194,1790]" captionTargetId="figure-0@28.[205,1363,194,1791]" captionTargetPageId="28" captionText="Figure 23. Microdipoena nyungwe (Mysmenidae), spinnerets: A, B, female; C–H, male. A, C, anterior lateral spinnerets (ALS); B, E, left posterior lateral spinnerets; D, posterior median spinnerets; F, ALS, detail of major ampullate field; G, H, ALS, detail of intersegmental lobe. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331663" httpUri="https://zenodo.org/record/5331663/files/figure.png" pageId="255" pageNumber="782">Fig. 23E</figureCitation>
; secondarily present in 
<taxonomicName id="4C2AE7E210B9939FFB71E58FFA3A5B26" box="[1202,1442,1268,1290]" class="Arachnida" family="Mysmenidae" genus="Mysmena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="species" species="leichhardti">
<emphasis id="B95E407310B9939FFB71E58FFA3A5B26" box="[1202,1442,1268,1290]" italics="true" pageId="255" pageNumber="782">Mysmena leichhardti</emphasis>
</taxonomicName>
and MYSM-005-ARG); orb webs with a proliferation of out-of-plane radii both above and below the orb plane (
<figureCitation id="131180E410B9939FFC81E42AFC485B49" box="[834,976,1360,1382]" captionStart="Figure 147" captionStartId="163.[164,243,1348,1367]" captionTargetBox="[164,1444,195,1318]" captionTargetId="figure-188@163.[164,1444,195,1318]" captionTargetPageId="163" captionText="Figure 147. Webs of Mysmenidae: A, Mysmena tasmaniae, B, MYSM-005-ARG (Mysmena), from Misiones, Argentina, female with egg sac; C, Mysmenidae from Chiapas, Mexico, detail of centre of web, external threads removed to expose the hub; D, Maymena sp. from Misiones, Argentina; E, same, detail of centre of web." figureDoi="http://doi.org/10.5281/zenodo.5331827" httpUri="https://zenodo.org/record/5331827/files/figure.png" pageId="255" pageNumber="782">Fig. 147A, B</figureCitation>
); males with prolateral row of slim setae occupying only distal half of tarsus I (
<figureCitation id="131180E410B9939FFB36E415FAC75BAA" box="[1269,1375,1391,1413]" captionStart="Figure 16" captionStartId="19.[164,244,1787,1806]" captionTargetBox="[241,1368,194,1757]" captionTargetId="figure-0@19.[241,1368,194,1757]" captionTargetPageId="19" captionText="Figure 16. Maymena rica (Mysmenidae), abdomen and legs: A, B, G, H, male; C–F, female. A, epiandrous spigots; B, left anterior lateral spinnerets; C, abdomen, ventral view. D, G, H, left leg I; E, F, left leg IV. D, femur, ventral view; E, tibia, dorsal view; F, claws, retrolateral view; G, tibia–metatarsus junction, prolaterodorsal view; H, tarsus, prolaterodorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371421" httpUri="https://zenodo.org/record/5371421/files/figure.png" pageId="255" pageNumber="782">Figs 16H</figureCitation>
, 
<figureCitation id="131180E410B9939FFAAEE415FA055BAA" box="[1389,1437,1391,1413]" captionStart="Figure 26" captionStartId="31.[164,242,1513,1532]" captionTargetBox="[164,1445,194,1482]" captionTargetId="figure-138@31.[164,1445,194,1482]" captionTargetPageId="31" captionText="Figure 26. Microdipoena nyungwe (Mysmenidae), left legs: A, C, D, G, male; B, E, F, female. A, tarsus I, prolateral view; B, claws I, retrolateral view; C, leg I, tibia–metatarsal junction, prolateral view; D, metatarsus I, dorsal view; E, leg I, metatarsus–tarsus junction, dorsal view; F, tarsus IV, prolateral view; G, tibia IV, prolateral view." figureDoi="http://doi.org/10.5281/zenodo.5331669" httpUri="https://zenodo.org/record/5331669/files/figure.png" pageId="255" pageNumber="782">26A</figureCitation>
, 
<figureCitation id="131180E410B9939FFCF9E4F4FCF25B8C" box="[826,874,1422,1443]" captionStart="Figure 34" captionStartId="51.[164,241,1560,1579]" captionTargetBox="[164,1445,194,1529]" captionTargetId="figure-104@51.[164,1445,194,1530]" captionTargetPageId="51" captionText="Figure 34. Mysmena (= Calodipoena) incredula (Mysmenidae); legs. A, B, female right leg I; C–F, male left legs. A, femoral spot, retrolateral view; B, metatarsal trichobothrium, dorsal view; C, metatarsus I, prolateral view; D, tarsus I, prolateral view; E, femur IV, prolateral view; F, claws IV, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.5331679" httpUri="https://zenodo.org/record/5331679/files/figure.png" pageId="255" pageNumber="782">34D</figureCitation>
, 
<figureCitation id="131180E410B9939FFCB7E4F7FC025B8C" box="[884,922,1421,1443]" captionStart="Figure 45" captionStartId="62.[144,223,1787,1806]" captionTargetBox="[157,1412,193,1757]" captionTargetId="figure-0@62.[157,1412,193,1757]" captionTargetPageId="62" captionText="Figure 45. MYSM-019-MAD (Mysmeninae, Mysmenidae) from Toliara, Madagascar. A–C, male right palp, inverted; A, dorsal view; B, retrolateral view; C, prolateral view. D, E, male left palp, expanded; D, prolateral–ventral view; E, same, detail of embolus and pars pendula; F, female posterior spinnerets; G–I, male left leg I, prolateral view; G, femur and patella; H, metatarsus; I, tarsus. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371455" httpUri="https://zenodo.org/record/5371455/files/figure.png" pageId="255" pageNumber="782">45I</figureCitation>
, 
<figureCitation id="131180E410B9939FFC66E4F7FC4A5B8C" box="[933,978,1421,1443]" captionStart="Figure 50" captionStartId="67.[164,243,1739,1758]" captionTargetBox="[164,1445,195,1709]" captionTargetId="figure-0@67.[164,1445,195,1709]" captionTargetPageId="67" captionText="Figure 50. MYSM-023-MAD (Mysmeninae, Mysmenidae) from Antananarivo, Madagascar: A, D, E, female; B, C, F–H, male. A, B, prosoma, lateral view; C, left chelicera, retromarginal view. D–H, right leg I, prolateral view; D, tarsus– metatarsus junction, prolateral–dorsal view; E, F, tarsus; G, tibia; H, metatarsus." figureDoi="http://doi.org/10.5281/zenodo.5331687" httpUri="https://zenodo.org/record/5331687/files/figure.png" pageId="255" pageNumber="782">50F</figureCitation>
; all along tarsus in 
<taxonomicName id="4C2AE7E210B9939FFB69E4F7FA0B5B8D" box="[1194,1427,1421,1442]" class="Arachnida" family="Mysmenidae" genus="Mysmeniola" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="B95E407310B9939FFB69E4F7FA0B5B8D" box="[1194,1427,1421,1442]" italics="true" pageId="255" pageNumber="782">Mysmeniola spinifera</emphasis>
</taxonomicName>
); cymbial prolateral basal expansion (
<figureCitation id="131180E410B9939FFB1AE4D6FAD85BEE" box="[1241,1344,1452,1474]" captionStart="Figure 27" captionStartId="32.[144,223,1752,1771]" captionTargetBox="[208,1361,194,1721]" captionTargetId="figure-0@32.[208,1361,194,1722]" captionTargetPageId="32" captionText="Figure 27. Microdipoena (= Mysmenella) samoensis (Mysmenidae), syntypes: A–C, F, H, I, male; D, E, G, female. A, left palp, prolateral–ventral view; B, same, retrolateral–ventral view; C, same, detail of tip of embolus and interaction with primary cymbial conductor, prolateral view; D, digested abdomen, vulva, and part of respiratory system; E, same, detail of vulva; F, G, prosoma, lateral view; H, mouthparts, ventral view, detail of retromargin of chelicerae and labium; I, left tibia I, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331671" httpUri="https://zenodo.org/record/5331671/files/figure.png" pageId="255" pageNumber="782">Figs 27A</figureCitation>
, 
<figureCitation id="131180E410B9939FFA8EE4D6FA055BEE" box="[1357,1437,1452,1473]" captionStart="Figure 30" captionStartId="36.[144,221,1103,1122]" captionTargetBox="[144,1424,194,1072]" captionTargetId="figure-359@36.[144,1424,194,1072]" captionTargetPageId="36" captionText="Figure 30. MYSM-007-MEX (Mysmena, Mysmenidae) from Chiapas, Mexico; male left palp. A, retroventral view; B, ventral view; C, prolateral view; D, dorsal view; E, retrolateral view; F, detail of tip of palp, ventral–prolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371429" httpUri="https://zenodo.org/record/5371429/files/figure.png" pageId="255" pageNumber="782">30B, C</figureCitation>
, 
<figureCitation id="131180E410B9939FFCF9E4B0FCF25BCF" box="[826,874,1482,1504]" captionStart="Figure 36" captionStartId="53.[164,239,1113,1132]" captionTargetBox="[164,1444,195,1082]" captionTargetId="figure-314@53.[164,1444,195,1084]" captionTargetPageId="53" captionText="Figure 36. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; male left palp; A, retrolateral view; B, distal view; C, prolateral view; D, ventral view; E, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331681" httpUri="https://zenodo.org/record/5331681/files/figure.png" pageId="255" pageNumber="782">36C</figureCitation>
, 
<figureCitation id="131180E410B9939FFCB6E4B1FC3C5BCE" box="[885,932,1483,1505]" captionStart="Figure 47" captionStartId="64.[144,221,1188,1207]" captionTargetBox="[144,1424,194,1158]" captionTargetId="figure-244@64.[144,1424,194,1158]" captionTargetPageId="64" captionText="Figure 47. MYSM-020-MAD (Mysmeninae, Mysmenidae) from Toamasina, Madagascar, male. A–E, left palp; A, retrolateral view; B, prolateral–ventral view; C, same, detail of tip of embolus and cymbial conductors; D, same, ventral–retrolateral view; E, same, prolateral view. F, digested abdomen, tracheal system, dorsal view; G, same, detail of posterior tracheal system. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331683" httpUri="https://zenodo.org/record/5331683/files/figure.png" pageId="255" pageNumber="782">47B</figureCitation>
); embolus with pars pendula (
<figureCitation id="131180E410B9939FFB39E4B1FAFA5BCF" box="[1274,1378,1483,1505]" captionStart="Figure 32" captionStartId="48.[144,223,1491,1510]" captionTargetBox="[144,1424,194,1461]" captionTargetId="figure-124@48.[144,1424,194,1461]" captionTargetPageId="48" captionText="Figure 32. Mysmena (= Calodipoena) incredula (Mysmenidae); male palp. A–C, E–H, left palp; D, right palp, subtly expanded, inverted. A, D retrolateral view; B, ventral–prolateral view; C, detail of proximal palp, retrolateral view; E, detail of tibia and prolateral basal expansion, ventral view; F, same, close up; G, detail of tip of palp, ventral view; H, same, retrolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371433" httpUri="https://zenodo.org/record/5371433/files/figure.png" pageId="255" pageNumber="782">Figs 32H</figureCitation>
, 
<figureCitation id="131180E410B9939FFAADE4B0FA055BCF" box="[1390,1437,1482,1504]" captionStart="Figure 36" captionStartId="53.[164,239,1113,1132]" captionTargetBox="[164,1444,195,1082]" captionTargetId="figure-314@53.[164,1444,195,1084]" captionTargetPageId="53" captionText="Figure 36. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; male left palp; A, retrolateral view; B, distal view; C, prolateral view; D, ventral view; E, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331681" httpUri="https://zenodo.org/record/5331681/files/figure.png" pageId="255" pageNumber="782">36B</figureCitation>
, 
<figureCitation id="131180E410B9939FFCF9E493FC005BD1" box="[826,920,1513,1534]" captionStart="Figure 132" captionStartId="148.[144,222,1638,1657]" captionTargetBox="[149,1421,202,1603]" captionTargetId="figure-0@148.[144,1424,193,1607]" captionTargetPageId="148" captionText="Figure 132. Mysmenidae male left palp, cleared: A, Microdipoena elsae, prolateral view, arrow to spine of basal prolateral cymbial expansion; B, Microdipoena guttata, prolateral view, arrow to spine of basal prolateral cymbial expansion; C, Microdipoena (= Mysmenella) illectrix, type, prolateral view; D, Microdipoena (= Mysmenella) samoensis, syntype, prolateral view; E, same, retrolateral view; F, Microdipoena (= Mysmenella) jobi, holotype, retrolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331805" httpUri="https://zenodo.org/record/5331805/files/figure.png" pageId="255" pageNumber="782">132C–F</figureCitation>
, 
<figureCitation id="131180E410B9939FFC6AE493FC715BD1" box="[937,1001,1513,1534]" captionStart="Figure 133" captionStartId="149.[164,242,1693,1712]" captionTargetBox="[227,1382,195,1662]" captionTargetId="figure-0@149.[221,1386,194,1664]" captionTargetPageId="149" captionText="Figure 133. Mysmenidae male left palp (unless otherwise stated), cleared. A, MYSM-005-ARG (Mysmena), from Misiones, Argentina, dorsal view, right palp, inverted, arrow to cymbial groove (CyG); B, same, ventral view; C, MYSM-007-MEX (Mysmena), from Chiapas, Mexico, retrolateral view, arrows to CyG; D, Mysmena (= Kekenboschiella) awari, paratype, prolateral view; E, same, dorsal–retrolateral view; F, Mysmena (= Kekenboschiella) marijkeae, holotype, retrolateral view, right palp, inverted; G, Brasilionata arborense, holotype, retrolateral view, right palp, inverted; H, Mysmena (= Tamasesia) rotunda, type, prolateral view; I, same, retrolateral view, schematic drawing. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331809" httpUri="https://zenodo.org/record/5331809/files/figure.png" pageId="255" pageNumber="782">133C</figureCitation>
, secondarily absent in MYSM-005- ARG); palpal tibial rim setae longer and arranged distally in a row or two [
<figureCitation id="131180E410B9939FFBFFE75CFB3E5814" box="[1084,1190,1574,1596]" captionStart="Figure 18" captionStartId="22.[144,222,1539,1558]" captionTargetBox="[144,1424,194,1509]" captionTargetId="figure-83@22.[144,1424,194,1510]" captionTargetPageId="22" captionText="Figure 18. Microdipoena guttata (Mysmenidae): A–F, male left palp; G, female. A, ventral–proximal view; B, retrolateral– distal view; C, dorsal–retrolateral view; D, expanded bulb, prolateral view; E, prolateral view; F, expanded bulb, detail of tip of embolus, retrolateral–distal view; G, digested abdomen, detail of vulva. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331657" httpUri="https://zenodo.org/record/5331657/files/figure.png" pageId="255" pageNumber="782">Figs 18A</figureCitation>
, 
<figureCitation id="131180E410B9939FFB70E75CFB7A5814" box="[1203,1250,1574,1595]" captionStart="Figure 32" captionStartId="48.[144,223,1491,1510]" captionTargetBox="[144,1424,194,1461]" captionTargetId="figure-124@48.[144,1424,194,1461]" captionTargetPageId="48" captionText="Figure 32. Mysmena (= Calodipoena) incredula (Mysmenidae); male palp. A–C, E–H, left palp; D, right palp, subtly expanded, inverted. A, D retrolateral view; B, ventral–prolateral view; C, detail of proximal palp, retrolateral view; E, detail of tibia and prolateral basal expansion, ventral view; F, same, close up; G, detail of tip of palp, ventral view; H, same, retrolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371433" httpUri="https://zenodo.org/record/5371433/files/figure.png" pageId="255" pageNumber="782">32E</figureCitation>
, 
<figureCitation id="131180E410B9939FFB32E75CFABA5813" box="[1265,1314,1574,1596]" captionStart="Figure 36" captionStartId="53.[164,239,1113,1132]" captionTargetBox="[164,1444,195,1082]" captionTargetId="figure-314@53.[164,1444,195,1084]" captionTargetPageId="53" captionText="Figure 36. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; male left palp; A, retrolateral view; B, distal view; C, prolateral view; D, ventral view; E, dorsal view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5331681" httpUri="https://zenodo.org/record/5331681/files/figure.png" pageId="255" pageNumber="782">36D</figureCitation>
, 
<figureCitation id="131180E410B9939FFAECE75CFAC75814" box="[1327,1375,1574,1595]" captionStart="Figure 42" captionStartId="59.[164,243,1792,1811]" captionTargetBox="[196,1412,194,1762]" captionTargetId="figure-0@59.[196,1412,194,1762]" captionTargetPageId="59" captionText="Figure 42. Mysmena leichhardti (Mysmenidae) from Queensland, Australia: A, B, E, G, H, male; C, D, F, female. A, prosoma, lateral view; B, same, lateral–frontal view; C, epigynal area and scapus, ventral view; D, digested abdomen, detail of vulva, tracheae removed; E, mouthparts, distal view; right femur I, prolateral–dorsal view; G, left femur I, ventral view; H, left metatarsus I, prolateral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371449" httpUri="https://zenodo.org/record/5371449/files/figure.png" pageId="255" pageNumber="782">42B</figureCitation>
, 
<figureCitation id="131180E410B9939FFAAEE75CFA055813" box="[1389,1437,1574,1596]" captionStart="Figure 45" captionStartId="62.[144,223,1787,1806]" captionTargetBox="[157,1412,193,1757]" captionTargetId="figure-0@62.[157,1412,193,1757]" captionTargetPageId="62" captionText="Figure 45. MYSM-019-MAD (Mysmeninae, Mysmenidae) from Toliara, Madagascar. A–C, male right palp, inverted; A, dorsal view; B, retrolateral view; C, prolateral view. D, E, male left palp, expanded; D, prolateral–ventral view; E, same, detail of embolus and pars pendula; F, female posterior spinnerets; G–I, male left leg I, prolateral view; G, femur and patella; H, metatarsus; I, tarsus. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371455" httpUri="https://zenodo.org/record/5371455/files/figure.png" pageId="255" pageNumber="782">45C</figureCitation>
; secondarily irregular in 
<taxonomicName id="4C2AE7E210B9939FFBA4E73FFAF05875" box="[1127,1384,1605,1626]" class="Arachnida" family="Mysmenidae" genus="Mysmeniola" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="B95E407310B9939FFBA4E73FFAF05875" box="[1127,1384,1605,1626]" italics="true" pageId="255" pageNumber="782">Mysmeniola spinifera</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2AE7E210B9939FFCF9E719FC4D5857" authorityName="Banks" authorityYear="1895" box="[826,981,1635,1656]" class="Arachnida" family="Mysmenidae" genus="Microdipoena" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFCF9E719FC4D5857" box="[826,981,1635,1656]" italics="true" pageId="255" pageNumber="782">Microdipoena</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C2AE7E210B9939FFC3EE719FB1D5857" authorityName="Brignoli" authorityYear="1980" box="[1021,1157,1635,1656]" class="Arachnida" family="Mysmenidae" genus="Mysmenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407310B9939FFC3EE719FB1D5857" box="[1021,1157,1635,1656]" italics="true" pageId="255" pageNumber="782">Mysmenella</emphasis>
</taxonomicName>
) 
<taxonomicName id="4C2AE7E210B9939FFB55E719FB585857" box="[1174,1216,1635,1656]" class="Arachnida" family="Mysmenidae" genus="Mysmenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="species" species="jobi">
<emphasis id="B95E407310B9939FFB55E719FB585857" box="[1174,1216,1635,1656]" italics="true" pageId="255" pageNumber="782">jobi</emphasis>
</taxonomicName>
]; and females with weakly modified epigynal area (i.e. epigynum absent; 
<figureCitation id="131180E410B9939FFCF9E7DBFC055899" box="[826,925,1697,1719]" captionStart="Figure 14" captionStartId="17.[164,243,1036,1055]" captionTargetBox="[164,1444,196,1004]" captionTargetId="figure-351@17.[164,1444,196,1006]" captionTargetPageId="17" captionText="Figure 14. Maymena rica (Mysmenidae): A, B, D, male left palp; C, E, female. A, dorsal view; B, same, prolateral view, detail of tip of embolus and housing cymbial conductors; C, epigynum, ventral view; D, same as B, dorsal view; E, digested abdomen, detail of vulva. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371417" httpUri="https://zenodo.org/record/5371417/files/figure.png" pageId="255" pageNumber="782">Figs 14C</figureCitation>
, 
<figureCitation id="131180E410B9939FFC6BE7DBFC4E5899" box="[936,982,1697,1718]" captionStart="Figure 37" captionStartId="54.[144,224,1265,1284]" captionTargetBox="[144,1425,195,1235]" captionTargetId="figure-266@54.[144,1425,195,1235]" captionTargetPageId="54" captionText="Figure 37. Mysmena-MYSM-015-MAD (Mysmena, Mysmenidae) from Antananarivo, Madagascar; female abdomen; A, epigynal area and scapus, ventral view; B, left posterior spinnerets (PS); C, same as A, posterolateral view; D, digested abdomen, detail of vulva; E, right PS. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371439" httpUri="https://zenodo.org/record/5371439/files/figure.png" pageId="255" pageNumber="782">37A</figureCitation>
, 
<figureCitation id="131180E410B9939FFC22E7DBFB965898" box="[993,1038,1697,1719]" captionStart="Figure 52" captionStartId="69.[164,243,1574,1593]" captionTargetBox="[164,1445,194,1544]" captionTargetId="figure-74@69.[164,1445,194,1544]" captionTargetPageId="69" captionText="Figure 52. A, MYSM-034-MAD (Mysmena, Mysmenidae), female; B–D, MYSM-029-MAD (Mysmeninae, Mysmenidae), female; E, F, MYSM-028-MAD (Mysmena, Mysmenidae), female. A, prosoma, lateral view; B, right anterior lateral spinneret; C, right posterior lateral spinneret; D, right leg I, claws, retrolateral view; E, chelicerae and palps, frontal view; F, epigynal area, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371467" httpUri="https://zenodo.org/record/5371467/files/figure.png" pageId="255" pageNumber="782">52F</figureCitation>
; modified copulatory area in MYSM-023-MAD, 
<figureCitation id="131180E410B9939FFC02E7C5FBCB58FA" box="[961,1107,1727,1749]" captionStart="Figure 49" captionStartId="66.[144,223,1765,1784]" captionTargetBox="[201,1366,195,1735]" captionTargetId="figure-0@66.[201,1366,195,1735]" captionTargetPageId="66" captionText="Figure 49. MYSM-023-MAD (Mysmeninae, Mysmenidae) from Antananarivo, Madagascar: A, D, E, female; B, C, F–H, male. A, digested abdomen, detail of vulva; B, left palp, prolateral view, bulb displaced; C, same, detail of cymbial tip, retrolateral–dorsal view; D, abdomen, detail of epigynal area and respiratory spiracles, posterior view; E, same, ventral view; F, colulus and spinnerets, ventral view; G, posterior median spinnerets; H, prosoma, detail of sternum, labium and maxillae, ventral view. See Appendix 3 for the list of abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5371461" httpUri="https://zenodo.org/record/5371461/files/figure.png" pageId="255" pageNumber="782">Fig. 49D, E</figureCitation>
). A total of 163 molecular synapomorphies support this subfamily.
</paragraph>
</subSubSection>
<subSubSection id="C330CFEA10B99260FCF9E666FDD05B24" lastPageId="256" lastPageNumber="783" pageId="255" pageNumber="782" type="diagnosis">
<paragraph id="8B959C6110B9939FFCF9E666FC32591E" blockId="255.[826,1441,1820,1902]" box="[826,938,1820,1841]" pageId="255" pageNumber="782">
<emphasis id="B95E407310B9939FFCF9E666FC32591E" box="[826,938,1820,1841]" italics="true" pageId="255" pageNumber="782">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B959C6110B99260FCF9E640FDD05B24" blockId="255.[826,1441,1820,1902]" lastBlockId="256.[144,759,198,1291]" lastPageId="256" lastPageNumber="783" pageId="255" pageNumber="782">
<taxonomicName id="4C2AE7E210B9939FFCF9E640FC54597F" authorityName="Petrunkevitch" authorityYear="1928" box="[826,972,1850,1872]" class="Arachnida" family="Mysmemidae" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="782" phylum="Arthropoda" rank="subFamily" subFamily="Mysmeninae">Mysmeninae</taxonomicName>
differs from all other mysmenid genera and subfamilies by the following unique combination of features: the characteristic architecture of their orb webs with a proliferation of out-of-plane radii both above and below the orb plane; anterior tracheae restricted to opisthosoma (extending into prosoma in MYSM-005-ARG, MYSM-007- 
<collectingCountry id="F33DDCF111469260FE45E03AFE595F7A" box="[390,449,320,341]" name="Mexico" pageId="256" pageNumber="783">MEX</collectingCountry>
, and 
<emphasis id="B95E407311469260FE3FE03AFD205F79" box="[508,696,320,342]" italics="true" pageId="256" pageNumber="783">
<taxonomicName id="4C2AE7E211469260FE3FE03AFD095F7A" authorityName="Banks" authorityYear="1895" box="[508,657,320,341]" class="Arachnida" family="Mysmenidae" genus="Microdipoena" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus" sensu="stricto">Microdipoena</taxonomicName>
<taxonomicNameLabel id="A26DFD0811469260FD55E03BFD205F79" box="[662,696,321,342]" pageId="256" pageNumber="783" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
), and an advanced and wide posterior spiracular opening located midway between the spinnerets and epigastric groove, connected to branched posterior lateral tracheae extending into prosoma and surrounding the minute median apodemes; anterior lateral spinnerets with fingerprint cuticular pattern on the piriform field, posterior spinnerets with aciniform gland spigots of two different outlines, and posterior lateral spinnerets with an anterior flat spatulate modified seta [as in 
<taxonomicName id="4C2AE7E211469260FD46E32EFD685C46" authorityName="Gertsch" authorityYear="1960" box="[645,752,596,617]" class="Arachnida" family="Mysmenidae" genus="Maymena" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407311469260FD46E32EFD685C46" box="[645,752,596,617]" italics="true" pageId="256" pageNumber="783">Maymena</emphasis>
</taxonomicName>
, secondarily absent in 
<taxonomicName id="4C2AE7E211469260FE42E309FE725CA7" authorityName="Simon" authorityYear="1894" box="[385,490,627,648]" class="Arachnida" family="Mysmenidae" genus="Mysmena" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407311469260FE42E309FE725CA7" box="[385,490,627,648]" italics="true" pageId="256" pageNumber="783">Mysmena</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C2AE7E211469260FDCDE309FD1A5CA7" authorityName="Marples" authorityYear="1955" box="[526,642,627,648]" class="Arachnida" family="Mysmenidae" genus="Tamasesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407311469260FDCDE309FD1A5CA7" box="[526,642,627,648]" italics="true" pageId="256" pageNumber="783">Tamasesia</emphasis>
</taxonomicName>
) 
<taxonomicName id="4C2AE7E211469260FD52E308FD715CA8" box="[657,745,626,647]" class="Arachnida" family="Mysmenidae" genus="Tamasesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="species" species="rotunda">
<emphasis id="B95E407311469260FD52E308FD715CA8" box="[657,745,626,647]" italics="true" pageId="256" pageNumber="783">rotunda</emphasis>
</taxonomicName>
], slim cylindrical spigots, and flagelliform spigots absent in males (secondarily present in males of 
<taxonomicName id="4C2AE7E211469260FD49E3CAFE895CCC" class="Arachnida" family="Mysmenidae" genus="Mysmena" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="species" species="leichhardti">
<emphasis id="B95E407311469260FD49E3CAFE895CCC" italics="true" pageId="256" pageNumber="783">Mysmena leichhardti</emphasis>
</taxonomicName>
and MYSM-005-ARG); females without epigynum and with the copulatory openings within the epigastric furrow (secondarily external in MYSM-023- MAD), a membranous internal atrium and irregular membranous copulatory ducts (distally sclerotized independently in two clades); males with secondary (external) cymbial conductor (secondarily absent in clade C128: 
<taxonomicName id="4C2AE7E211469260FF1EE2DEFEE15D96" authorityName="Banks" authorityYear="1895" box="[221,377,932,953]" class="Arachnida" family="Mysmenidae" genus="Microdipoena" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407311469260FF1EE2DEFEE15D96" box="[221,377,932,953]" italics="true" pageId="256" pageNumber="783">Microdipoena</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E211469260FE4FE2DEFD845D96" authorityName="Wunderlich" authorityYear="1995" box="[396,540,932,953]" class="Arachnida" family="Mysmenidae" genus="Brasilionata" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407311469260FE4FE2DEFD845D96" box="[396,540,932,953]" italics="true" pageId="256" pageNumber="783">Brasilionata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE7E211469260FDECE2DEFD215D96" authorityName="Thaler" authorityYear="1995" box="[559,697,932,953]" class="Arachnida" family="Mysmenidae" genus="Mysmeniola" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407311469260FDECE2DEFD215D96" box="[559,697,932,953]" italics="true" pageId="256" pageNumber="783">Mysmeniola</emphasis>
</taxonomicName>
, and MYSM-019-MAD), cymbial prolateral basal expansion, coiled embolus with pars pendula (secondarily absent in MYSM-005-ARG), palpal tibial rim setae long and arranged distally in a row or two [secondarily irregular in 
<taxonomicName id="4C2AE7E211469260FED6E547FD8C5A7D" box="[277,532,1085,1106]" class="Arachnida" family="Mysmenidae" genus="Mysmeniola" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="B95E407311469260FED6E547FD8C5A7D" box="[277,532,1085,1106]" italics="true" pageId="256" pageNumber="783">Mysmeniola spinifera</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2AE7E211469260FD94E547FD6E5A7D" authorityName="Banks" authorityYear="1895" box="[599,758,1085,1106]" class="Arachnida" family="Mysmenidae" genus="Microdipoena" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407311469260FD94E547FD6E5A7D" box="[599,758,1085,1106]" italics="true" pageId="256" pageNumber="783">Microdipoena</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C2AE7E211469260FF6EE526FEAB5A5E" authorityName="Brignoli" authorityYear="1980" box="[173,307,1116,1137]" class="Arachnida" family="Mysmenidae" genus="Mysmenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="genus">
<emphasis id="B95E407311469260FF6EE526FEAB5A5E" box="[173,307,1116,1137]" italics="true" pageId="256" pageNumber="783">Mysmenella</emphasis>
</taxonomicName>
) 
<taxonomicName id="4C2AE7E211469260FE80E526FEF45A5E" box="[323,364,1116,1137]" class="Arachnida" family="Mysmenidae" genus="Mysmenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="species" species="jobi">
<emphasis id="B95E407311469260FE80E526FEF45A5E" box="[323,364,1116,1137]" italics="true" pageId="256" pageNumber="783">jobi</emphasis>
</taxonomicName>
], a prolateral row of slim setae occupying only distal half of tarsus I (all along tarsus in 
<taxonomicName id="4C2AE7E211469260FF73E5E3FE305A81" box="[176,424,1177,1198]" class="Arachnida" family="Mysmenidae" genus="Mysmeniola" kingdom="Animalia" order="Araneae" pageId="256" pageNumber="783" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="B95E407311469260FF73E5E3FE305A81" box="[176,424,1177,1198]" italics="true" pageId="256" pageNumber="783">Mysmeniola spinifera</emphasis>
</taxonomicName>
); and also a strongly serrated distal promarginal curved seta, abdomen with a whitish ventral ring around the spinnerets, trichobothria on tibia III and IV of medium length.
</paragraph>
</subSubSection>
</treatment>
</document>