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<document id="2CAF8EE7745EA8E7C7261476837026E7" ID-CLB-Dataset="21179" ID-DOI="10.11646/zootaxa.4615.1.5" ID-GBIF-Dataset="5cad0e06-2a49-4dc6-bfac-9f00391b5d9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995398" ID-ZooBank="2A000D68-CA54-4077-AA8C-50B4455E6A23" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598058885214" checkinUser="felipe" docAuthor="Megna, Yoandri S., Balke, Michael, Apenborn, Rico &amp; Hendrich, Lars" docDate="2019" docId="0382E754E76BFFBFFF22FB40FA78F9E2" docLanguage="en" docName="zootaxa.4615.1.5.pdf" docOrigin="Zootaxa 4615 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydaticus (Prodaticus)" docType="key" docVersion="8" lastPageNumber="128" masterDocId="FFBB9F2CE765FFB0FFB5FFE9FFE5F813" masterDocTitle="A review of Peruvian diving beetles of the genus Hydaticus Leach, 1817, with description of Hydaticus (Prodaticus) panguana sp. nov. and notes on other Neotropical species (Coleoptera: Dytiscidae)" masterLastPageNumber="130" masterPageNumber="113" pageNumber="127" updateTime="1698845201127" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7D667D9660E73309F43395D8E5E4A65B">A review of Peruvian diving beetles of the genus Hydaticus Leach, 1817, with description of Hydaticus (Prodaticus) panguana sp. nov. and notes on other Neotropical species (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="E3C4F52F89481D098EF212DD6821441F">Megna, Yoandri S.</mods:namePart>
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<mods:affiliation id="813627CF67D02FD69DDA35CDDC5D00F2">SNSB-Zoologische Staatssammlung München, Münchhausenstrasse 21, 81247 München, Germany</mods:affiliation>
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<treatment id="0382E754E76BFFBFFF22FB40FA78F9E2" ID-DOI="http://doi.org/10.5281/zenodo.4330435" ID-GBIF-Taxon="167293119" ID-Zenodo-Dep="4330435" LSID="urn:lsid:plazi:treatment:0382E754E76BFFBFFF22FB40FA78F9E2" httpUri="http://treatment.plazi.org/id/0382E754E76BFFBFFF22FB40FA78F9E2" lastPageId="15" lastPageNumber="128" pageId="14" pageNumber="127">
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<paragraph id="8B945642E76BFFBEFF22FB40FB93FCD1" blockId="14.[151,1142,1192,1219]" box="[151,1142,1192,1219]" pageId="14" pageNumber="127">
<heading id="D0DCE12EE76BFFBEFF22FB40FB93FCD1" bold="true" box="[151,1142,1192,1219]" fontSize="11" level="1" pageId="14" pageNumber="127" reason="1">
<emphasis id="B95F8A50E76BFFBEFF22FB40FB93FCD1" bold="true" box="[151,1142,1192,1219]" pageId="14" pageNumber="127">
Key to the species of subgenus
<taxonomicName id="4C2B2DC1E76BFFBEFDADFB41FCCAFCD1" ID-CoL="8NXT4" authority="" box="[536,815,1192,1219]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="127" phylum="Arthropoda" rank="subGenus" subGenus="Prodaticus">
<emphasis id="B95F8A50E76BFFBEFDADFB41FD77FCD0" bold="true" box="[536,658,1192,1219]" italics="true" pageId="14" pageNumber="127">Hydaticus</emphasis>
(
<emphasis id="B95F8A50E76BFFBEFD16FB41FCC0FCD0" bold="true" box="[675,805,1192,1219]" italics="true" pageId="14" pageNumber="127">Prodaticus</emphasis>
)
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of the Neotropical Region
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<paragraph id="8B945642E76BFFBEFF22FB04FA79FD11" blockId="14.[151,1437,1261,2016]" box="[151,1436,1261,1283]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22FB04FA79FD11" box="[151,1436,1261,1283]" pageId="14" pageNumber="127">
1. Elytron with a transverse and subbasal yellowish fascia (
<figureCitation id="13104AC7E76BFFBEFCB7FB04FCD9FD11" box="[770,828,1261,1283]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[391,1187,591,1847]" captionTargetId="figure@2.[340,1247,491,1887]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal view of the colour pattern of a generalized Hydaticus for the Neotropical region. Head (He): Ptbb-posterior transverse black band. Pronotum (Pr): Abb- anterior black band; Pbb-posterior black band; Plaa- posterior lateral angle acute. Scutellum (Sc). Elytron (El): Ai- apex irrorated; Bs- basal spot; Ls- lateral yellow stripes; Ms- middle spot; Sls- Subbasal and lateral yellow stripe; Tbf- transverse basal fasciae; Tpi- third posterior irrorated." figureDoi="http://doi.org/10.5281/zenodo.3995400" httpUri="https://zenodo.org/record/3995400/files/figure.png" pageId="14" pageNumber="127">Fig. 1</figureCitation>
. Tbf)............................................. 2
</keyLead>
</paragraph>
<paragraph id="8B945642E76BFFBEFF22FAE2FA79FD33" blockId="14.[151,1437,1261,2016]" box="[151,1436,1290,1313]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22FAE2FA79FD33" box="[151,1436,1290,1313]" pageId="14" pageNumber="127">
- Elytron without a transverse and subbasal yellowish fascia (
<figureCitation id="13104AC7E76BFFBEFC94FAE3FC8AFD32" box="[801,879,1290,1313]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="14" pageNumber="127">Fig. 2A</figureCitation>
)............................................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DF4EE7E76BFFBEFF22FAC1FA79FD49" pageId="14" pageNumber="127">
<paragraph id="8B945642E76BFFBEFF22FAC1FA79FD2E" blockId="14.[151,1437,1261,2016]" box="[151,1436,1320,1342]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22FAC1FA79FD2E" box="[151,1436,1320,1342]" pageId="14" pageNumber="127">
2. Pronotum yellowish with a black band on anterior and posterior margin (
<figureCitation id="13104AC7E76BFFBEFC28FAC1FC33FD2E" box="[925,982,1320,1342]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[391,1187,591,1847]" captionTargetId="figure@2.[340,1247,491,1887]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal view of the colour pattern of a generalized Hydaticus for the Neotropical region. Head (He): Ptbb-posterior transverse black band. Pronotum (Pr): Abb- anterior black band; Pbb-posterior black band; Plaa- posterior lateral angle acute. Scutellum (Sc). Elytron (El): Ai- apex irrorated; Bs- basal spot; Ls- lateral yellow stripes; Ms- middle spot; Sls- Subbasal and lateral yellow stripe; Tbf- transverse basal fasciae; Tpi- third posterior irrorated." figureDoi="http://doi.org/10.5281/zenodo.3995400" httpUri="https://zenodo.org/record/3995400/files/figure.png" pageId="14" pageNumber="127">Fig. 1</figureCitation>
, Abb, Pbb)............................ 4
</keyLead>
</paragraph>
<paragraph id="8B945642E76BFFBEFF22FAAFFA79FD49" blockId="14.[151,1437,1261,2016]" box="[151,1436,1349,1371]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22FAAFFA79FD49" box="[151,1436,1349,1371]" pageId="14" pageNumber="127">
- Pronotum black with broad yellowish lateral margins (
<figureCitation id="13104AC7E76BFFBEFD58FAACFCDDFD48" box="[749,824,1349,1371]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="14" pageNumber="127">Fig. 2B</figureCitation>
)................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DF4EE7E76BFFBEFF22FA8AFA79FDC3" pageId="14" pageNumber="127">
<paragraph id="8B945642E76BFFBEFF22FA8AFA78FD86" blockId="14.[151,1437,1261,2016]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22FA8AFA78FD86" pageId="14" pageNumber="127">
3. Elytron with basal fasciae confluent with sublateral, longitudinal and yellowish stripes; elytron without a subtriangular, postmedial and broad spot..........................................................................
<taxonomicName id="4C2B2DC1E76BFFBEFA9AFA69FA78FD86" ID-CoL="7VG53" authorityName="Tremouilles" authorityYear="1996" box="[1327,1437,1407,1429]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="127" phylum="Arthropoda" rank="species" species="devexus">
<emphasis id="B95F8A50E76BFFBEFA9AFA69FA78FD86" bold="true" box="[1327,1437,1407,1429]" italics="true" pageId="14" pageNumber="127">H. devexus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8B945642E76BFFBEFF22FA77FA79FDC3" blockId="14.[151,1437,1261,2016]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22FA77FA79FDC3" pageId="14" pageNumber="127">
- Elytron usually with basal fasciae not confluent with sublateral, longitudinal and yellowish stripes; elytron with a subtriangular, postmedial and broad spot (
<figureCitation id="13104AC7E76BFFBEFE59FA53FDD2FDC2" box="[492,567,1466,1489]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="14" pageNumber="127">Fig. 2B</figureCitation>
)...............................................................
<taxonomicName id="4C2B2DC1E76BFFBEFA9BFA52FA79FDC3" ID-CoL="7VG65" authorityName="Laporte" authorityYear="1835" box="[1326,1436,1466,1488]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="127" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B95F8A50E76BFFBEFA9BFA52FA79FDC3" bold="true" box="[1326,1436,1466,1488]" italics="true" pageId="14" pageNumber="127">H. lateralis</emphasis>
</taxonomicName>
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</keyStep>
<keyStep id="30DF4EE7E76BFFBEFF22FA31FA78FE8E" pageId="14" pageNumber="127">
<paragraph id="8B945642E76BFFBEFF22FA31FA78FE56" blockId="14.[151,1437,1261,2016]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22FA31FA78FE56" pageId="14" pageNumber="127">
4. TL
<quantity id="4CD3FBA7E76BFFBEFEB3FA31FE71FDFE" box="[262,404,1496,1518]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.26" metricValueMin="1.14" pageId="14" pageNumber="127" unit="mm" value="12.0" valueMax="12.6" valueMin="11.4">11.412.6 mm</quantity>
; pronotum with basal black band narrowly extended to anterior margin; elytron without sublateral yellow stripes, and with a large lateral spot just behind middle; apical third largely irrorated; protarsomere I with a series of spines across the dorsal surface; metatibia with bifid setae not parallel to dorsal margin on posterior surface (
<figureCitation id="13104AC7E76BFFBEFB01F9FBFB1AFE3A" box="[1204,1279,1554,1577]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="14" pageNumber="127">Fig. 2C</figureCitation>
)....................................................................................................
<taxonomicName id="4C2B2DC1E76BFFBEFAB6F9D9FA78FE56" ID-CoL="7VGB3" authorityName="Laporte" authorityYear="1835" box="[1283,1437,1583,1605]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="127" phylum="Arthropoda" rank="species" species="subfasciatus">
<emphasis id="B95F8A50E76BFFBEFAB6F9D9FA78FE56" bold="true" box="[1283,1437,1583,1605]" italics="true" pageId="14" pageNumber="127">H. subfasciatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B945642E76BFFBEFF22F9A7FA78FE8E" blockId="14.[151,1437,1261,2016]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22F9A7FA78FE8E" pageId="14" pageNumber="127">
- TL
<quantity id="4CD3FBA7E76BFFBEFEB1F9A4FE63FE70" box="[260,390,1613,1635]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.04" metricValueMin="0.96" pageId="14" pageNumber="127" unit="mm" value="10.0" valueMax="10.4" valueMin="9.6">9.610.4 mm</quantity>
; pronotum with basal black band not extended; elytron with sublateral and lateral yellow stripes, and without lateral spots; apical third largely rufous (
<figureCitation id="13104AC7E76BFFBEFDD0F983FD54FE92" box="[613,689,1642,1665]" captionStart="FIGURE 3" captionStartId="7.[151,250,1982,2007]" captionTargetBox="[246,1342,181,1962]" captionTargetId="figure@7.[246,1342,181,1962]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of H. fractivittis (A) and H. palliatus (B), both from Paraguay, and H. rimosus (C) from Guatemala. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995404" httpUri="https://zenodo.org/record/3995404/files/figure.png" pageId="14" pageNumber="127">Fig. 3D</figureCitation>
); protarsomere I without a series of spines across the dorsal surface; metatibia with bifid setae parallel to dorsal margin on posterior surface.......................................
<taxonomicName id="4C2B2DC1E76BFFBEFB4AF961FA78FE8E" ID-CoL="7VGBN" baseAuthorityName="Brulle" baseAuthorityYear="1837" box="[1279,1437,1671,1693]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="127" phylum="Arthropoda" rank="species" species="xanthomelas">
<emphasis id="B95F8A50E76BFFBEFB4AF961FA78FE8E" bold="true" box="[1279,1437,1671,1693]" italics="true" pageId="14" pageNumber="127">H. xanthomelas</emphasis>
</taxonomicName>
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</keyStep>
<keyStep id="30DF4EE7E76BFFBEFF22F94CFA79FECB" pageId="14" pageNumber="127">
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<keyLead id="30DAF577E76BFFBEFF22F94CFA79FEA8" box="[151,1436,1701,1723]" pageId="14" pageNumber="127">
5. Pronotum black with yellowish lateral margins (
<figureCitation id="13104AC7E76BFFBEFD05F94CFD1BFEA8" box="[688,766,1701,1723]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="14" pageNumber="127">Fig. 2A</figureCitation>
)...................................................... 6
</keyLead>
</paragraph>
<paragraph id="8B945642E76BFFBEFF22F92AFA79FECB" blockId="14.[151,1437,1261,2016]" box="[151,1436,1730,1752]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22F92AFA79FECB" box="[151,1436,1730,1752]" pageId="14" pageNumber="127">
- Pronotum yellowish with a black band on anterior and posterior margin (
<figureCitation id="13104AC7E76BFFBEFC28F92BFC33FECB" box="[925,982,1730,1752]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[391,1187,591,1847]" captionTargetId="figure@2.[340,1247,491,1887]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal view of the colour pattern of a generalized Hydaticus for the Neotropical region. Head (He): Ptbb-posterior transverse black band. Pronotum (Pr): Abb- anterior black band; Pbb-posterior black band; Plaa- posterior lateral angle acute. Scutellum (Sc). Elytron (El): Ai- apex irrorated; Bs- basal spot; Ls- lateral yellow stripes; Ms- middle spot; Sls- Subbasal and lateral yellow stripe; Tbf- transverse basal fasciae; Tpi- third posterior irrorated." figureDoi="http://doi.org/10.5281/zenodo.3995400" httpUri="https://zenodo.org/record/3995400/files/figure.png" pageId="14" pageNumber="127">Fig. 1</figureCitation>
, Abb, Pbb)............................ 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DF4EE7E76BFFBEFF22F909FA79FF23" pageId="14" pageNumber="127">
<paragraph id="8B945642E76BFFBEFF22F909FA79FEE6" blockId="14.[151,1437,1261,2016]" box="[151,1436,1760,1782]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22F909FA79FEE6" box="[151,1436,1760,1782]" pageId="14" pageNumber="127">
6. Elytron with lateral, longitudinal and yellowish stripes, and without a subtriangular, postmedial and broad spot..
<taxonomicName id="4C2B2DC1E76BFFBEFA95F909FA79FEE6" ID-CoL="7VGBD" authorityName="Tremouilles" authorityYear="1996" box="[1312,1436,1760,1781]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="127" phylum="Arthropoda" rank="species" species="tuyuensis">
<emphasis id="B95F8A50E76BFFBEFA95F909FA79FEE6" bold="true" box="[1312,1436,1760,1781]" italics="true" pageId="14" pageNumber="127">H. tuyuensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B945642E76BFFBEFF22F917FA79FF23" blockId="14.[151,1437,1261,2016]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22F917FA79FF23" pageId="14" pageNumber="127">
- Elytron without lateral, longitudinal and yellowish stripes, and with a subtriangular, postmedial and broad spot (
<figureCitation id="13104AC7E76BFFBEFAA0F914FA86FF00" box="[1301,1379,1789,1811]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="14" pageNumber="127">Fig. 2A</figureCitation>
)......................................................................................
<emphasis id="B95F8A50E76BFFBEFB72F8F2FA79FF23" bold="true" box="[1223,1436,1819,1840]" pageId="14" pageNumber="127">
<taxonomicName id="4C2B2DC1E76BFFBEFB72F8F2FAACFF23" ID-CoL="8LZ8X" authority="Megna &amp; Balke &amp; Apenborn &amp; Hendrich, 2019" authorityName="Megna &amp; Balke &amp; Apenborn &amp; Hendrich" authorityYear="2019" box="[1223,1353,1819,1840]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="127" phylum="Arthropoda" rank="species" species="panguana" status="sp. nov.">
<emphasis id="B95F8A50E76BFFBEFB72F8F2FAACFF23" bold="true" box="[1223,1353,1819,1840]" italics="true" pageId="14" pageNumber="127">H. panguana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C372BE76BFFBEFAE5F8F2FA79FF23" box="[1360,1436,1819,1840]" pageId="14" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DF4EE7E76BFFBEFF22F8D1FA79FFB6" pageId="14" pageNumber="127">
<paragraph id="8B945642E76BFFBEFF22F8D1FA79FF78" blockId="14.[151,1437,1261,2016]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22F8D1FA79FF78" pageId="14" pageNumber="127">
7. Protarsus without accessory medial spinous setae on dorsal surface of basal tarsomere of male; elytron without lateral yellow stripes.....................................................................................
<taxonomicName id="4C2B2DC1E76BFFBEFAAFF8BFFA79FF78" ID-CoL="7VG5J" authorityName="Guignot" authorityYear="1951" box="[1306,1436,1877,1899]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="127" phylum="Arthropoda" rank="species" species="fractivittis">
<emphasis id="B95F8A50E76BFFBEFAAFF8BFFA79FF78" bold="true" box="[1306,1436,1877,1899]" italics="true" pageId="14" pageNumber="127">H. fractivittis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B945642E76BFFBEFF22F89AFA79FFB6" blockId="14.[151,1437,1261,2016]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22F89AFA79FFB6" pageId="14" pageNumber="127">
- Protarsus with accessory medial spinous setae on dorsal surface of basal tarsomere of male; elytron with lateral yellow stripes (
<figureCitation id="13104AC7E76BFFBEFF5CF879FEC7FFB6" box="[233,290,1936,1958]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[391,1187,591,1847]" captionTargetId="figure@2.[340,1247,491,1887]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal view of the colour pattern of a generalized Hydaticus for the Neotropical region. Head (He): Ptbb-posterior transverse black band. Pronotum (Pr): Abb- anterior black band; Pbb-posterior black band; Plaa- posterior lateral angle acute. Scutellum (Sc). Elytron (El): Ai- apex irrorated; Bs- basal spot; Ls- lateral yellow stripes; Ms- middle spot; Sls- Subbasal and lateral yellow stripe; Tbf- transverse basal fasciae; Tpi- third posterior irrorated." figureDoi="http://doi.org/10.5281/zenodo.3995400" httpUri="https://zenodo.org/record/3995400/files/figure.png" pageId="14" pageNumber="127">Fig. 1</figureCitation>
, Ls)........................................................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DF4EE7E76BFFBFFF22F844FA79F8DF" lastPageId="15" lastPageNumber="128" pageId="14" pageNumber="127">
<paragraph id="8B945642E76BFFBEFF22F844FA79FFF3" blockId="14.[151,1437,1261,2016]" pageId="14" pageNumber="127">
<keyLead id="30DAF577E76BFFBEFF22F844FA79FFF3" pageId="14" pageNumber="127">
8. Distribution:
<collectingCountry id="F33C16D2E76BFFBEFED6F844FE21FFD0" box="[355,452,1965,1987]" name="Colombia" pageId="14" pageNumber="127">Colombia</collectingCountry>
,
<collectingCountry id="F33C16D2E76BFFBEFE78F844FDFAFFD0" box="[461,543,1965,1987]" name="Ecuador" pageId="14" pageNumber="127">Ecuador</collectingCountry>
,
<collectingCountry id="F33C16D2E76BFFBEFD9DF844FD80FFD0" box="[552,613,1965,1987]" name="Brazil" pageId="14" pageNumber="127">Brazil</collectingCountry>
,
<collectingCountry id="F33C16D2E76BFFBEFDDAF847FD2DFFD0" box="[623,712,1966,1987]" name="Paraguay" pageId="14" pageNumber="127">Paraguay</collectingCountry>
and
<collectingCountry id="F33C16D2E76BFFBEFD40F844FCB0FFD0" box="[757,853,1965,1987]" name="Argentina" pageId="14" pageNumber="127">Argentina</collectingCountry>
median lobe of aedeagus rounded apically in lateral view... ................................................................................................... 9
</keyLead>
</paragraph>
<paragraph id="8B945642E76AFFBFFF22FF73FA79F8DF" blockId="15.[151,1437,153,498]" pageId="15" pageNumber="128">
<keyLead id="30DAF577E76AFFBFFF22FF73FA79F8DF" pageId="15" pageNumber="128">
- Distribution:
<collectingCountry id="F33C16D2E76AFFBFFED6FF73FE70F8BC" box="[355,405,154,175]" name="United States of America" pageId="15" pageNumber="128">USA</collectingCountry>
, Central America and West Indies; median lobe of aedeagus angulate or truncated apically in lateral view. .................................................................................................. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DF4EE7E76AFFBFFF22FF3DFA79F951" pageId="15" pageNumber="128">
<paragraph id="8B945642E76AFFBFFF22FF3DFA79F915" blockId="15.[151,1437,153,498]" pageId="15" pageNumber="128">
<keyLead id="30DAF577E76AFFBFFF22FF3DFA79F915" pageId="15" pageNumber="128">
9. TL
<quantity id="4CD3FBA7E76AFFBFFEB1FF3DFE74F8F9" box="[260,401,212,234]" metricMagnitude="-2" metricUnit="m" metricValue="1.165" metricValueMax="1.25" metricValueMin="1.08" pageId="15" pageNumber="128" unit="mm" value="11.65" valueMax="12.5" valueMin="10.8">10.812.5 mm</quantity>
; epipenite with apex rounded; median lobe of aedeagus in apical third greatly widened and rounded in dorsal view..................................................................................
<taxonomicName id="4C2B2DC1E76AFFBFFB4DFF1BFA79F915" ID-CoL="7VG5W" authorityName="Tremouilles" authorityYear="1996" box="[1272,1436,241,263]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="inexspectatus">
<emphasis id="B95F8A50E76AFFBFFB4DFF1BFA79F915" bold="true" box="[1272,1436,241,263]" italics="true" pageId="15" pageNumber="128">H. inexspectatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B945642E76AFFBFFF22FEE6FA79F951" blockId="15.[151,1437,153,498]" pageId="15" pageNumber="128">
<keyLead id="30DAF577E76AFFBFFF22FEE6FA79F951" pageId="15" pageNumber="128">
- TL 13.0
<quantity id="4CD3FBA7E76AFFBFFE8FFEE6FE74F937" box="[314,401,271,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="15" pageNumber="128" unit="mm" value="13.6">13.6 mm</quantity>
; epipenite with apex acute; median lobe of aedeagus in apical third subparallel and slightly rounded in dorsal view.......................................................................................
<taxonomicName id="4C2B2DC1E76AFFBFFA9DFEC5FA79F951" ID-CoL="7VG6Y" authorityName="Aube" authorityYear="1838" box="[1320,1436,300,322]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="palliatus">
<emphasis id="B95F8A50E76AFFBFFA9DFEC5FA79F951" bold="true" box="[1320,1436,300,322]" italics="true" pageId="15" pageNumber="128">H. palliatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DF4EE7E76AFFBFFF22FEA3FA78F9E2" pageId="15" pageNumber="128">
<paragraph id="8B945642E76AFFBFFF22FEA3FA79F989" blockId="15.[151,1437,153,498]" pageId="15" pageNumber="128">
<keyLead id="30DAF577E76AFFBFFF22FEA3FA79F989" pageId="15" pageNumber="128">
10. Median lobe of aedeagus angulate at apex in lateral view; each elytron with a definite and regular submarginal yellow stripe, more or less recurved at the base, without inward extensions; posterior black band of pronotum usually restricted to basal third; anterior disc of metatibiae generally with more than 10 large spines..................................
<taxonomicName id="4C2B2DC1E76AFFBFFB4FFE6DFA79F989" ID-CoL="7VG4K" baseAuthorityName="Say" baseAuthorityYear="1830" box="[1274,1436,388,410]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="bimarginatus">
<emphasis id="B95F8A50E76AFFBFFB4FFE6DFA79F989" bold="true" box="[1274,1436,388,410]" italics="true" pageId="15" pageNumber="128">H. bimarginatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B945642E76AFFBFFF22FE4BFA78F9E2" blockId="15.[151,1437,153,498]" pageId="15" pageNumber="128">
<keyLead id="30DAF577E76AFFBFFF22FE4BFA78F9E2" pageId="15" pageNumber="128">
- Median lobe of aedeagus truncate apically in lateral view; each elytron with a submarginal stripe usually broad, sometimes with inward extensions (
<figureCitation id="13104AC7E76AFFBFFE28FE56FE0CF9C6" box="[413,489,447,469]" captionStart="FIGURE 3" captionStartId="7.[151,250,1982,2007]" captionTargetBox="[246,1342,181,1962]" captionTargetId="figure@7.[246,1342,181,1962]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of H. fractivittis (A) and H. palliatus (B), both from Paraguay, and H. rimosus (C) from Guatemala. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995404" httpUri="https://zenodo.org/record/3995404/files/figure.png" pageId="15" pageNumber="128">Fig. 3C</figureCitation>
); posterior black band of pronotum often extended to or almost to anterior margin; anterior disc of metatibiae generally with less than 10 large spines....................................................
<taxonomicName id="4C2B2DC1E76AFFBFFA9AFE35FA78F9E2" ID-CoL="7VG9K" authorityName="Aube" authorityYear="1838" box="[1327,1437,476,497]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="rimosus">
<emphasis id="B95F8A50E76AFFBFFA9AFE35FA78F9E2" bold="true" box="[1327,1437,476,497]" italics="true" pageId="15" pageNumber="128">H. rimosus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>