202 lines
19 KiB
XML
202 lines
19 KiB
XML
<document id="314EAAE03F1CC2C13D3BE71DCBD3ED1A" ID-CLB-Dataset="276483" ID-DOI="10.1111/j.1096-3642.2009.00552.x" ID-GBIF-Dataset="3be82545-1797-46a0-95fc-3e0fc44c9af0" ID-ISSN="0024-4082" ID-Zenodo-Dep="5438122" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630440497297" checkinUser="valdenar" docAuthor="Zhang, Kuiyan, Yang, Ding, Ren, Dong & Shih, Chungkun" docDate="2010" docId="038296595830FF8EFF19939AFE325100" docLanguage="en" docName="j.1096-3642.2009.00552.x.pdf" docOrigin="Zoological Journal of the Linnean Society 158 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00552.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Uranorhagio daohugouensis Zhang & Yang & Ren & Shih 2010" docType="treatment" docVersion="5" lastPageNumber="566" masterDocId="FFBBEE215832FF8DFFBB9529FFCA5766" masterDocTitle="An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)" masterLastPageNumber="572" masterPageNumber="563" pageNumber="565" updateTime="1699284570084" updateUser="ExternalLinkService">
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<mods:titleInfo id="DD6A062B6E07FDA13587365BA58430E1">
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<mods:title id="E017B2CE7CC7C5F5EE2C15CE4DA17DDE">An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)</mods:title>
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</mods:titleInfo>
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<mods:name id="68FE6650F8264CC79641F52F9EAE547D" type="personal">
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<mods:roleTerm id="0E7D1B33D4677F684E7614A35A7BD47C">Author</mods:roleTerm>
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<mods:namePart id="A9E66E98A95DD4CEBEB11540FCF41340">Zhang, Kuiyan</mods:namePart>
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</mods:name>
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<mods:name id="516F8D37DCF6A43DE4C51184AA8597AE" type="personal">
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<mods:role id="5B744BFDCCCBFAC5DEFBF6A4D89A3AF4">
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<mods:roleTerm id="97AD118A61594EF8AEA40B9876F21228">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FC694E599D7A0CF63AD36B267C1E3A82">Yang, Ding</mods:namePart>
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</mods:name>
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<mods:name id="7B9AA3D5D19DA9F099A3778DE99989F0" type="personal">
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<mods:roleTerm id="4EF3ADC3C4BC7BB60B9467FAE4A41AD7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="985BBB5B4D6C1CD8F7B9B463396F3951">Ren, Dong</mods:namePart>
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</mods:name>
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<mods:name id="3F4BF956B35405A35A955A74ADBBAE8F" type="personal">
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<mods:role id="6C833A2070828EC9613A921F5F149976">
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<mods:roleTerm id="CD2A2B1AFF98E05B9DFEB5B9F6B25DF3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0CE321B10EAA27B6794F6C90CFE34307">Shih, Chungkun</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="86CF611E23893B42D93865EEACBBF07B">text</mods:typeOfResource>
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<mods:title id="6F5879D08A6839D5642A373550D6C56A">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="FCFC959E7AB2BA4E9F3534B70E5FFE4D">
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<mods:date id="5397ABAB570ACB297BA80CC992A7CF63">2010</mods:date>
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<mods:detail id="F25BCE7E84AA776E0CF42DD8D52477D2" type="pubDate">
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<mods:number id="1BFFA728612319A448684E054EB5E27F">2010-02-26</mods:number>
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</mods:detail>
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<mods:detail id="642F10358C1EEA48935E8C0A5B517355" type="volume">
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<mods:number id="86E8740228E43FDE7A3F2BDC748439A5">158</mods:number>
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</mods:detail>
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<mods:detail id="BC11A07E48B3913764AE68CC34DAB8B9" type="issue">
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<mods:number id="FFCF2BBFFF38F5EB8594941428AD974E">3</mods:number>
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<mods:extent id="9651D605BAC5B419B7D3DC1BCD124DA4" unit="page">
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<mods:start id="68228EF25A0E65E972DFA5DFC1C95935">563</mods:start>
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<mods:end id="B234E533217C64728481573B1F719467">572</mods:end>
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<mods:location id="EB32B0CE227570F1C787CE1417630FB2">
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<mods:url id="ADD6B9CEDF1F087DA12285947EEC3114">http://dx.doi.org/10.1111/j.1096-3642.2009.00552.x</mods:url>
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</mods:location>
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<mods:classification id="EE44F614F5746991A92C97F7ABE7EDD2">journal article</mods:classification>
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<mods:identifier id="BEAFBA7064B8163A83AE60A614C5A9D1" type="CLB-Dataset">276483</mods:identifier>
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<mods:identifier id="6DCAE45490AA88A89C4B7F907AA0A71C" type="DOI">10.1111/j.1096-3642.2009.00552.x</mods:identifier>
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<mods:identifier id="D7C092CB7DF1E36B2668640BFFE7F754" type="GBIF-Dataset">3be82545-1797-46a0-95fc-3e0fc44c9af0</mods:identifier>
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<mods:identifier id="58C27E79E3AFA1BE7F4D52A072AE9014" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="EE5E61CC9635AF80D17A0E973DC47AA1" type="Zenodo-Dep">5438122</mods:identifier>
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<treatment id="038296595830FF8EFF19939AFE325100" ID-GBIF-Taxon="217238225" LSID="urn:lsid:plazi:treatment:038296595830FF8EFF19939AFE325100" httpUri="http://treatment.plazi.org/id/038296595830FF8EFF19939AFE325100" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565">
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<subSubSection id="C33174C45830FF8FFF19939AFD4C518E" pageId="2" pageNumber="565" type="nomenclature">
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<paragraph id="8B94274F5830FF8FFF19939AFD4C518E" blockId="2.[162,740,1714,1768]" pageId="2" pageNumber="565">
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<heading id="D0DC90235830FF8FFF19939AFD4C518E" centered="true" fontSize="9" level="2" pageId="2" pageNumber="565" reason="4">
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<taxonomicName id="4C2B5CCC5830FF8FFF19939AFDE951A1" authority="Zhang & Yang & Ren & Shih, 2010" authorityName="Zhang & Yang & Ren & Shih" authorityYear="2010" box="[162,547,1715,1736]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="daohugouensis" status="sp. nov.">
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<emphasis id="B95FFB5D5830FF8FFF19939AFDE951A1" bold="true" box="[162,547,1715,1736]" italics="true" pageId="2" pageNumber="565">URANORHAGIO DAOHUGOUENSIS</emphasis>
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</taxonomicName>
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ZHANG, YANG & REN
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<emphasis id="B95FFB5D5830FF8FFE8C93FFFDCF5181" bold="true" box="[311,517,1746,1767]" pageId="2" pageNumber="565">GEN. ET SP. NOV.</emphasis>
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(
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<figureCitation id="13103BCA5830FF8FFDA393FFFD945181" box="[536,606,1746,1767]" captionStart="Figure 1" captionStartId="2.[144,223,1189,1208]" captionTargetBox="[143,1424,664,1158]" captionTargetId="figure-483@2.[723,1424,664,1158]" captionTargetPageId="2" captionText="Figure 1. Uranorhagio daohugouensis sp. nov. Holotype, body with a wing in lateral view. A, photograph. B, camera lucida drawing." figureDoi="http://doi.org/10.5281/zenodo.5438124" httpUri="https://zenodo.org/record/5438124/files/figure.png" pageId="2" pageNumber="565">FIGS 1</figureCitation>
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,
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<figureCitation id="13103BCA5830FF8FFDCB93FBFDB4518E" box="[624,638,1746,1768]" captionStart="Figure 2" captionStartId="3.[164,243,831,850]" captionTargetBox="[164,1442,197,800]" captionTargetId="figure-568@3.[743,1444,197,802]" captionTargetPageId="3" captionText="Figure 2. Uranorhagio daohugouensis sp. nov. Paratype, body with a wing in lateral view. A, photograph. B, camera lucida drawing." figureDoi="http://doi.org/10.5281/zenodo.5438126" httpUri="https://zenodo.org/record/5438126/files/figure.png" pageId="2" pageNumber="565">2</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33174C45830FF8EFF2B93D0FE325100" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565" type="etymology">
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<paragraph id="8B94274F5830FF8FFF2B93D0FD66504B" blockId="2.[144,759,1785,1837]" pageId="2" pageNumber="565">
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<emphasis id="B95FFB5D5830FF8FFF2B93D0FEC75068" box="[144,269,1785,1806]" italics="true" pageId="2" pageNumber="565">Etymology:</emphasis>
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The specific name refers to the
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<typeStatus id="549099ED5830FF8FFD3B93D0FD7B5068" box="[640,689,1785,1806]" pageId="2" pageNumber="565">type</typeStatus>
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locality: Daohugou Village,
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<collectingRegion id="49EFE9AD5830FF8FFE279231FD9B504B" box="[412,593,1815,1837]" country="China" name="Nei Mongol" pageId="2" pageNumber="565">Inner Mongolia</collectingRegion>
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,
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<collectingCountry id="F33C67DF5830FF8FFDE4923EFD62504B" box="[607,680,1815,1837]" name="China" pageId="2" pageNumber="565">China</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B94274F5830FF8FFF2B9270FD7B5008" blockId="2.[144,689,1881,1903]" box="[144,689,1881,1903]" pageId="2" pageNumber="565">
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<emphasis id="B95FFB5D5830FF8FFF2B9270FECC5008" box="[144,262,1881,1902]" italics="true" pageId="2" pageNumber="565">Diagnosis:</emphasis>
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The same as
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<taxonomicName id="4C2B5CCC5830FF8FFE159270FD8A5008" authority="Zhang & Yang & Ren & Shih, 2010" authorityName="Zhang & Yang & Ren & Shih" authorityYear="2010" box="[430,576,1881,1902]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B95FFB5D5830FF8FFE159270FD8A5008" box="[430,576,1881,1902]" italics="true" pageId="2" pageNumber="565">Uranorhagio</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C46265830FF8FFDF19270FD7B5008" box="[586,689,1881,1902]" pageId="2" pageNumber="565" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B94274F5830FF8FFC9D9023FB3B5259" blockId="2.[806,1421,1290,1343]" pageId="2" pageNumber="565">
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<emphasis id="B95FFB5D5830FF8FFC9D9023FC5A5279" box="[806,912,1290,1311]" italics="true" pageId="2" pageNumber="565">
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<typeStatus id="549099ED5830FF8FFC9D9023FC405279" box="[806,906,1290,1311]" pageId="2" pageNumber="565" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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CNU-DIB-NN2007016, an almost complete adult body with a wing in lateral view.
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</paragraph>
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<paragraph id="8B94274F5830FF8FFC9D9076FB3B52F5" blockId="2.[806,1421,1374,1427]" pageId="2" pageNumber="565">
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<emphasis id="B95FFB5D5830FF8FFC9D9076FC595212" box="[806,915,1375,1396]" italics="true" pageId="2" pageNumber="565">
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<typeStatus id="549099ED5830FF8FFC9D9076FC465212" box="[806,908,1375,1396]" pageId="2" pageNumber="565" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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CNU-DIB-NN2007017, an almost complete adult body with a wing in lateral view.
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</paragraph>
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<paragraph id="8B94274F5830FF8FFC9D909AFBDF5142" blockId="2.[806,1421,1458,1572]" pageId="2" pageNumber="565">
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<emphasis id="B95FFB5D5830FF8FFC9D909AFB8652A1" box="[806,1100,1458,1480]" italics="true" pageId="2" pageNumber="565">Type locality and horizon:</emphasis>
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Daohugou Village, Shantou Township, Ningcheng County,
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<collectingRegion id="49EFE9AD5830FF8FFB3990F8FAFF5281" box="[1154,1333,1489,1511]" country="China" name="Nei Mongol" pageId="2" pageNumber="565">Inner Mongolia</collectingRegion>
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,
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<collectingCountry id="F33C67DF5830FF8FFAFA90F8FA435281" box="[1345,1417,1489,1511]" name="China" pageId="2" pageNumber="565">China</collectingCountry>
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; Jiulongshan Formation, the Middle Jurassic (Aalenian–Bajocian).
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</paragraph>
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<paragraph id="8B94274F5830FF8FFC9D936DFB3C5111" blockId="2.[806,1422,1603,1900]" pageId="2" pageNumber="565">
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<emphasis id="B95FFB5D5830FF8FFC9D936DFC64513F" box="[806,942,1604,1625]" italics="true" pageId="2" pageNumber="565">Description:</emphasis>
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Body length, 18.0–
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<quantity id="4CD38AAA5830FF8FFB36936AFB38513E" box="[1165,1266,1603,1624]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="2" pageNumber="565" unit="mm" value="21.7">21.7 mm</quantity>
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; wing length, 13.5–18.0 mm; wing width,
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<quantity id="4CD38AAA5830FF8FFBD19348FB385111" box="[1130,1266,1633,1655]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.7" metricValueMin="4.2" pageId="2" pageNumber="565" unit="mm" value="4.95" valueMax="5.7" valueMin="4.2">4.2–5.7 mm</quantity>
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.
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</paragraph>
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<paragraph id="8B94274F5830FF8FFC8493A9FAB051D3" blockId="2.[806,1422,1603,1900]" pageId="2" pageNumber="565">Body stout. Head round, slightly narrower than thorax. Eyes large, bare; ommatidia visible clearly.</paragraph>
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<paragraph id="8B94274F5830FF8FFC849397FC33500A" blockId="2.[806,1422,1603,1900]" pageId="2" pageNumber="565">Mesonotum slightly convex. Legs pubescent. Femora and tibiae of fore and mid-legs slender; fore tibia with a spur; hind femur robust, strongly swollen at apical part, hind tibia with two spurs; first tarsomere distinctly longer than following tarsomeres, empodium pulvilliform.</paragraph>
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<caption id="DF5477C75831FF8EFF1F9616FE805416" ID-DOI="http://doi.org/10.5281/zenodo.5438126" ID-Zenodo-Dep="5438126" httpUri="https://zenodo.org/record/5438126/files/figure.png" pageId="3" pageNumber="566" startId="3.[164,243,831,850]" targetBox="[164,1442,197,800]" targetPageId="3">
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<paragraph id="8B94274F5831FF8EFF1F9616FE805416" blockId="3.[164,1441,831,880]" pageId="3" pageNumber="566">
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<emphasis id="B95FFB5D5831FF8EFF1F9616FD095435" bold="true" box="[164,707,831,851]" pageId="3" pageNumber="566">
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Figure 2.
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<taxonomicName id="4C2B5CCC5831FF8EFEA29616FDAC5434" authority="Zhang & Yang & Ren & Shih, 2010" authorityName="Zhang & Yang & Ren & Shih" authorityYear="2010" box="[281,614,831,850]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="daohugouensis" status="sp. nov.">
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<emphasis id="B95FFB5D5831FF8EFEA29616FDAC5434" bold="true" box="[281,614,831,850]" italics="true" pageId="3" pageNumber="566">Uranorhagio daohugouensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C46265831FF8EFDD69669FD095435" box="[621,707,832,851]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Paratype, body with a wing in lateral view. A, photograph. B, camera lucida drawing.
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</paragraph>
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</caption>
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<paragraph id="8B94274F5831FF8EFF079686FD13514F" blockId="3.[164,779,943,1638]" pageId="3" pageNumber="566">
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Wing hyaline, slender relative to body; veins thick and strong. Vein Sc ending slightly beyond middle of wing. Veins R
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<subScript id="17AF250A5831FF8EFEFE96DEFE875365" attach="left" box="[325,333,1015,1027]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
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and Rs
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<subScript id="17AF250A5831FF8EFE1296DEFE7B5365" attach="left" box="[425,433,1015,1027]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
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long; vein R
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<subScript id="17AF250A5831FF8EFDF296DEFDAB5365" attach="left" box="[585,609,1015,1027]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
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distinctly and strongly bent upwards at base; veins R
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<subScript id="17AF250A5831FF8EFDD0913CFD495347" attach="left" box="[619,643,1045,1057]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
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, R
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<subScript id="17AF250A5831FF8EFD1E913CFD675347" attach="left" box="[677,685,1045,1057]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
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, and R
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<subScript id="17AF250A5831FF8EFCB9913CFCC05347" attach="left" box="[770,778,1045,1057]" fontSize="5" pageId="3" pageNumber="566">5</subScript>
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straight and parallel with each other (except for bases of veins R
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<subScript id="17AF250A5831FF8EFEAE917BFEE75338" attach="left" box="[277,301,1106,1118]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
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and R
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<subScript id="17AF250A5831FF8EFEC1917BFE485338" attach="left" box="[378,386,1106,1118]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
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); beginning of vein R
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<subScript id="17AF250A5831FF8EFDCC917BFDB55338" attach="left" box="[631,639,1106,1118]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
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proximal to apex of cell d; vein R
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<subScript id="17AF250A5831FF8EFE289158FE51531B" attach="left" box="[403,411,1137,1149]" fontSize="5" pageId="3" pageNumber="566">5</subScript>
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ending at wing apex. Crossvein r–m at basal 3/7 of cell d, distal to beginning of vein R
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<subScript id="17AF250A5831FF8EFF0D9187FF0453DC" attach="left" box="[182,206,1198,1210]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
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; relative length of veins Rs
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<subScript id="17AF250A5831FF8EFDAD9187FDD453DC" attach="left" box="[534,542,1198,1210]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
|
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, Rs
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<subScript id="17AF250A5831FF8EFDF79187FD9E53DC" attach="left" box="[588,596,1198,1210]" fontSize="5" pageId="3" pageNumber="566">2</subScript>
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, and Rs
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<subScript id="17AF250A5831FF8EFD039187FD0A53DC" attach="left" box="[696,704,1198,1210]" fontSize="5" pageId="3" pageNumber="566">3</subScript>
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about 3.5: 1: 2.5 to 4.5: 1: 4. Veins M
|
||
<subScript id="17AF250A5831FF8EFDA591E4FDEC53BF" attach="left" box="[542,550,1229,1241]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
|
||
and M
|
||
<subScript id="17AF250A5831FF8EFDC091E4FD4953BF" attach="left" box="[635,643,1229,1241]" fontSize="5" pageId="3" pageNumber="566">2</subScript>
|
||
bifurcating distal of crossvein m–m, with base of vein M
|
||
<subScript id="17AF250A5831FF8EFD1691C2FD7F5391" attach="left" box="[685,693,1259,1271]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
|
||
curved upwards; vein M
|
||
<subScript id="17AF250A5831FF8EFED29023FEBB5270" attach="left" box="[361,369,1290,1302]" fontSize="5" pageId="3" pageNumber="566">3</subScript>
|
||
short. Vein CuA
|
||
<subScript id="17AF250A5831FF8EFD839023FD8A5270" attach="left" box="[568,576,1290,1302]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
|
||
arising from cell bm. Anal lobe with several veins (or just some creases of wing). Mouths of cells sc and r
|
||
<subScript id="17AF250A5831FF8EFD82906EFD8B5235" attach="left" box="[569,577,1351,1363]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
|
||
wide; mouths of cells r
|
||
<subScript id="17AF250A5831FF8EFF4A904FFEC35214" attach="left" box="[241,265,1382,1394]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
|
||
and r
|
||
<subScript id="17AF250A5831FF8EFEE4904FFEAD5214" attach="left" box="[351,359,1382,1394]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
|
||
distinctly narrower; cell r
|
||
<subScript id="17AF250A5831FF8EFD0F904FFD765214" attach="left" box="[692,700,1382,1394]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
|
||
much narrower. Cell bm slightly wider than cell br; cell d large, nearby wing margin, curved quadrangle. Five posterior cells present and wide open; mouths of cells m
|
||
<subScript id="17AF250A5831FF8EFF0190C9FF08528A" attach="left" box="[186,194,1504,1516]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
|
||
and m
|
||
<subScript id="17AF250A5831FF8EFE9990C9FEE0528A" attach="left" box="[290,298,1504,1516]" fontSize="5" pageId="3" pageNumber="566">2</subScript>
|
||
subequal in length; mouth of cell m
|
||
<subScript id="17AF250A5831FF8EFCB990C9FCC0528A" attach="left" box="[770,778,1504,1516]" fontSize="5" pageId="3" pageNumber="566">3</subScript>
|
||
narrow, about 1/5–1/2 as wide as that of cell cua
|
||
<subScript id="17AF250A5831FF8EFD7C90D6FD05516D" attach="left" box="[711,719,1535,1547]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
|
||
. Cell cup distinctly open; anal lobe small and narrow.
|
||
</paragraph>
|
||
<paragraph id="8B94274F5831FF8EFF07931BFE325100" blockId="3.[164,779,943,1638]" pageId="3" pageNumber="566">Abdomen robust, pubescent. Nine segments visible; segment I evidently shortest.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |