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<document id="4D1D480DF06AF6E7C7370725EB2F074A" ID-DOI="10.11646/zootaxa.4098.3.5" ID-GBIF-Dataset="ea61a19c-f9c5-4991-a486-9aa027b14e81" ID-ISSN="1175-5326" ID-Zenodo-Dep="264482" ID-ZooBank="C2DF33F9-A216-43A8-B2AD-ECC243DC9D2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460012126931" checkinUser="plazi" docAuthor="Iwatsuki, Yukio, Al-Mamry, Juma M. &amp; Heemstra, Phillip C." docDate="2016" docId="038287D99910FFA2CF8FFF79E22B6A0A" docLanguage="en" docName="zootaxa.4098.3.5.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lutjanus bengalensis Bloch 1790" docType="treatment" docVersion="9" lastPageNumber="523" masterDocId="FFBBFFA1991AFFAECF18FFEEE73D6C02" masterDocTitle="Validity of a blue stripe snapper, Lutjanus octolineatus (Cuvier 1828) and a related species, L. bengalensis (Bloch 1790) with a new species (Pisces; Lutjanidae) from the Arabian Sea" masterLastPageNumber="528" masterPageNumber="511" pageNumber="521" updateTime="1698233468564" updateUser="plazi">
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<mods:title id="ECAA2B329B9A87D7A194E5E69401BDFA">Validity of a blue stripe snapper, Lutjanus octolineatus (Cuvier 1828) and a related species, L. bengalensis (Bloch 1790) with a new species (Pisces; Lutjanidae) from the Arabian Sea</mods:title>
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<mods:namePart id="75CD17075F61C262F27F655DB0D24415">Iwatsuki, Yukio</mods:namePart>
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<mods:namePart id="CF3541E87B78F6EB2F85AA54F4AB1387">Al-Mamry, Juma M.</mods:namePart>
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<mods:namePart id="7F9AF536C240B54D490F587A06536701">Heemstra, Phillip C.</mods:namePart>
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<treatment id="038287D99910FFA2CF8FFF79E22B6A0A" ID-DOI="http://doi.org/10.5281/zenodo.6087590" ID-GBIF-Taxon="119381517" ID-Zenodo-Dep="6087590" LSID="urn:lsid:plazi:treatment:038287D99910FFA2CF8FFF79E22B6A0A" httpUri="http://treatment.plazi.org/id/038287D99910FFA2CF8FFF79E22B6A0A" lastPageId="12" lastPageNumber="523" pageId="10" pageNumber="521">
<subSubSection id="C33165449910FFA4CF8FFF79E5326CF5" pageId="10" pageNumber="521" type="nomenclature">
<paragraph id="8B9436CF9910FFA4CF8FFF79E5006CB0" blockId="10.[151,573,151,247]" box="[151,573,151,178]" pageId="10" pageNumber="521">
<heading id="D0DC81A39910FFA4CF8FFF79E5006CB0" bold="true" box="[151,573,151,178]" fontSize="11" level="1" pageId="10" pageNumber="521" reason="1">
<taxonomicName id="4C2B4D4C9910FFA4CF8FFF79E5006CB0" ID-CoL="3WG7Z" authority="Bloch 1790" authorityName="Bloch" authorityYear="1790" box="[151,573,151,178]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9910FFA4CF8FFF79E5006CB0" bold="true" box="[151,573,151,178]" pageId="10" pageNumber="521">
<emphasis id="B95FEADD9910FFA4CF8FFF79E6A46CB3" bold="true" box="[151,409,151,177]" italics="true" pageId="10" pageNumber="521">Lutjanus bengalensis</emphasis>
(
<bibRefCitation id="EFBA4B3E9910FFA4CEB0FF76E5086CB0" author="Bloch" box="[424,565,152,178]" pageId="10" pageNumber="521" refString="Bloch, M. E. (1790) Naturgeschichte der auslandischen Fische. Vol. 4. Icthyologie, ou Histoire naturelle des Poissons, Berlin, xii + 128 pp., pls. 217 - 252." type="book chapter" year="1790">Bloch 1790</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9436CF9910FFA4CF8FFF54E6D16CD0" blockId="10.[151,573,151,247]" box="[151,492,186,211]" pageId="10" pageNumber="521">English name: Bengal snapper</paragraph>
<paragraph id="8B9436CF9910FFA4CF8FFF30E5326CF5" blockId="10.[151,573,151,247]" box="[151,527,222,247]" pageId="10" pageNumber="521">
<figureCitation id="13102A4A9910FFA4CF8FFF30E7C06CF5" box="[151,253,222,247]" captionStart="FIGURE 1" captionStartId="6.[151,250,1902,1924]" captionTargetBox="[188,1398,216,1878]" captionTargetId="figure@6.[188,1398,195,1878]" captionTargetPageId="6" captionText="FIGURE 1. Type specimens of Lutjanus octolineatus (A, B) and L. bengalensis (C): A. MNHN 0000 - A 5446 (no label attached with a blue glass plate), 122 mm SL, lectotype of Diacope octolineata, Mauritius, Mascarenes; B. MNHN 0000 - 7046 (holotype of Diacope octovittata), 137 mm SL, Mauritius, Mascarenes; C. ZMB 303, lectotype of Holocentrus bengalensis), 94 mm SL, Bengal, Bay of Bengal, eastern Indian Ocean." httpUri="https://zenodo.org/record/264483/files/figure.png" pageId="10" pageNumber="521">Figures 1</figureCitation>
C, 2B, &amp; 3BC; Table 1
</paragraph>
</subSubSection>
<subSubSection id="C33165449910FFA4CF8FFEC8E3236E5C" pageId="10" pageNumber="521" type="reference_group">
<paragraph id="8B9436CF9910FFA4CF8FFEC8E5F36D78" blockId="10.[151,1437,294,500]" pageId="10" pageNumber="521">
<treatmentCitationGroup id="AB3B11E19910FFA4CF8FFEC8E5F36D78" pageId="10" pageNumber="521">
<taxonomicName id="4C2B4D4C9910FFA4CF8FFEC8E57E6D3E" authority="Bloch 1790: 102" authorityName="Bloch" authorityPageNumber="102" authorityYear="1790" box="[151,579,294,316]" class="Actinopterygii" family="Holocentridae" genus="Holocentrus" kingdom="Animalia" order="Beryciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9910FFA4CF8FFEC8E6AF6D3E" box="[151,402,294,316]" italics="true" pageId="10" pageNumber="521">Holocentrus bengalensis</emphasis>
<treatmentCitation id="0A8A10DE9910FFA4CE80FEC8E57E6D3E" author="Bloch" box="[408,579,294,316]" page="102" pageId="10" pageNumber="521" year="1790">
<bibRefCitation id="EFBA4B3E9910FFA4CE80FEC8E52C6D3E" author="Bloch" box="[408,529,294,316]" pageId="10" pageNumber="521" refString="Bloch, M. E. (1790) Naturgeschichte der auslandischen Fische. Vol. 4. Icthyologie, ou Histoire naturelle des Poissons, Berlin, xii + 128 pp., pls. 217 - 252." type="book chapter" year="1790">Bloch 1790</bibRefCitation>
: 102
</treatmentCitation>
</taxonomicName>
, pl. 246, fig. 2 (
<typeStatus id="5490886D9910FFA4CDFBFEC8E42A6D3E" box="[739,791,294,316]" pageId="10" pageNumber="521">Type</typeStatus>
locality: Bay of Bengal, eastern Indian Ocean).
<taxonomicName id="4C2B4D4C9910FFA4CF8FFEA8E56E6D59" authority="Bleeker 1855: 407" authorityName="Bleeker" authorityPageNumber="407" authorityYear="1855" box="[151,595,325,347]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="pomacanthus">
<emphasis id="B95FEADD9910FFA4CF8FFEA8E6AF6D59" box="[151,402,325,347]" italics="true" pageId="10" pageNumber="521">Mesoprion pomacanthus</emphasis>
<treatmentCitation id="0A8A10DE9910FFA4CE80FEABE56E6D59" author="Bleeker" box="[408,595,325,347]" page="407" pageId="10" pageNumber="521" year="1855">
<bibRefCitation id="EFBA4B3E9910FFA4CE80FEABE51E6D59" author="Bleeker" box="[408,547,325,347]" pageId="10" pageNumber="521" refString="Bleeker, P. (1855) Zesde bijdrage tot de kennis der ichthyologische fauna van Amboina. Natuurkundig Tijdschrift voor Nederlandsch Indie, 8, 391 - 434." type="journal article" year="1855">Bleeker 1855</bibRefCitation>
: 407
</treatmentCitation>
</taxonomicName>
(in part,
<typeStatus id="5490886D9910FFA4CDA8FEA8E5DF6D59" box="[688,738,326,347]" pageId="10" pageNumber="521">Type</typeStatus>
locality:
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Island, Molucca Islands,
<collectingCountry id="F33C765F9910FFA4CB89FEABE3C56D59" box="[1169,1272,325,347]" name="Indonesia" pageId="10" pageNumber="521">Indonesia</collectingCountry>
).?
<taxonomicName id="4C2B4D4C9910FFA4CFBAFE8AE6B26D78" box="[162,399,356,378]" family="Lutjanidae" genus="Genyoroge" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9910FFA4CFBAFE8AE6B26D78" box="[162,399,356,378]" italics="true" pageId="10" pageNumber="521">Genyoroge bengalensis</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A8A10DE9910FFA4CE85FE8AE5616D78" author="Gunther" box="[413,604,356,378]" page="178" pageId="10" pageNumber="521" year="1859">
<bibRefCitation id="EFBA4B3E9910FFA4CE85FE8AE5116D78" author="Gunther" box="[413,556,356,378]" pageId="10" pageNumber="521" refString="Gunther, A. (1859) Catalogue of the acanthopterygian fishes in the collection of the British Museum. Gasterosteidae, Berycidae, Percidae, Aphredoderidae, Pristipomatidae, Mullidae, Sparidae. Catalogue of the fishes in the British Museum, 1, i - xxxi + 1 - 524." type="journal article" year="1859">Günther 1859</bibRefCitation>
: 178
</treatmentCitation>
(Red Sea).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9436CF9910FFA4CF8FFE6DE4D36DF6" blockId="10.[151,1437,294,500]" pageId="10" pageNumber="521">
<taxonomicName id="4C2B4D4C9910FFA4CF8FFE6DE6516D9A" ID-CoL="3WG7Z" box="[151,364,386,408]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9910FFA4CF8FFE6DE6516D9A" box="[151,364,386,408]" italics="true" pageId="10" pageNumber="521">Lutjanus bengalensis</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFBA4B3E9910FFA4CE62FE6CE5796D9A" author="Kyushin" box="[378,580,386,408]" pageId="10" pageNumber="521" refString="Kyushin, K., Amaoka, K., Nakaya, K. &amp; Ida, H. (1977) Fishes of Indian Ocean. Japan Marine Fishery Resource Research Center, Tokyo, 392 pp., 179 pls. [in Japanese and English]" type="book" year="1977">
Kyushin
<emphasis id="B95FEADD9910FFA4CECEFE6DE53F6D9A" box="[470,514,386,408]" italics="true" pageId="10" pageNumber="521">et al</emphasis>
. 1977
</bibRefCitation>
: 68, unnumbered photograph (in part, Andaman Sea);
<bibRefCitation id="EFBA4B3E9910FFA4CB71FE6CE2036D9A" author="Allen" box="[1129,1342,386,408]" pageId="10" pageNumber="521" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot 1985</bibRefCitation>
: 21, pl. I, EI, F. (in part, as
<taxonomicName id="4C2B4D4C9910FFA4CE8DFE4CE50D6DB5" ID-CoL="3WGBT" box="[405,560,417,439]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="octolineatus">
<emphasis id="B95FEADD9910FFA4CE8DFE4CE50D6DB5" box="[405,560,417,439]" italics="true" pageId="10" pageNumber="521">L. octolineatus</emphasis>
</taxonomicName>
in plate, Andaman Sea and
<collectingCountry id="F33C765F9910FFA4CC7BFE4FE4F76DB5" box="[867,970,417,439]" name="Indonesia" pageId="10" pageNumber="521">Indonesia</collectingCountry>
); Allen 1985: 61, pl. XII, 45 (in part, as
<taxonomicName id="4C2B4D4C9910FFA4CA9FFE4CE67E6DD4" ID-CoL="3WGBT" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="octolineatus">
<emphasis id="B95FEADD9910FFA4CA9FFE4CE67E6DD4" italics="true" pageId="10" pageNumber="521">L. octolineatus</emphasis>
</taxonomicName>
in plate, Andaman Sea and
<collectingCountry id="F33C765F9910FFA4CD78FE2EE5FA6DD4" box="[608,711,448,470]" name="Indonesia" pageId="10" pageNumber="521">Indonesia</collectingCountry>
);
<bibRefCitation id="EFBA4B3E9910FFA4CDC1FE2EE49B6DD4" author="Iwatsuki" box="[729,934,448,470]" pageId="10" pageNumber="521" refString="Iwatsuki, Y., Yoshino, T. &amp; Shimada, K. (1999) Comparison of Lutjanus bengalensis from the western Pacific with a related species, L. kasmira, and variations in both species (Perciformes: Lutjanidae). Ichthyological Research, 46 (3), 314 - 317. http: // dx. doi. org / 10.1007 / BF 02678519" type="journal article" year="1999">
Iwatsuki
<emphasis id="B95FEADD9910FFA4CC2EFE2FE45E6DD4" box="[822,867,448,470]" italics="true" pageId="10" pageNumber="521">et al</emphasis>
. 1999
</bibRefCitation>
: 316, fig. 1A (Iye Isl., Okinawa,
<collectingCountry id="F33C765F9910FFA4CBEDFE2FE2096DD4" box="[1269,1332,449,470]" name="Japan" pageId="10" pageNumber="521">Japan</collectingCountry>
):
<bibRefCitation id="EFBA4B3E9910FFA4CA5DFE2EE6AB6DF6" author="Iwatsuki" pageId="10" pageNumber="521" refString="Iwatsuki, Y. &amp; Satapoomin, U. (2009) Lutjanidae, Snappers. In: Kimura, S., Satapoomin, U. &amp; Matsuura, K. (Eds.), Fishes of Andaman Sea: West coast of southern Thailand. National Museum of Nature and Science, Tokyo, pp. 135 - 143." type="book chapter" year="2009">Iwatsuki &amp; Satapoomin 2009</bibRefCitation>
: 136, unnumbered underwater photograph (Andaman Sea).
</paragraph>
<paragraph id="8B9436CF9910FFA4CF8FFDABE3236E5C" blockId="10.[151,1436,581,1974]" box="[151,1054,581,606]" pageId="10" pageNumber="521">
<emphasis id="B95FEADD9910FFA4CF8FFDABE62B6E5C" bold="true" box="[151,278,581,606]" pageId="10" pageNumber="521">
<typeStatus id="5490886D9910FFA4CF8FFDABE62C6E5C" box="[151,273,581,606]" pageId="10" pageNumber="521" type="lectotype">Lectotype</typeStatus>
:
</emphasis>
ZMB 303, 94 mm SL, Bengal, Bay of Bengal, eastern Indian Ocean.
</paragraph>
</subSubSection>
<subSubSection id="C33165449910FFA4CFDFFD87E5876FC7" pageId="10" pageNumber="521" type="materials_examined">
<paragraph id="8B9436CF9910FFA4CFDFFD87E41A6F7C" blockId="10.[151,1436,581,1974]" pageId="10" pageNumber="521">
<emphasis id="B95FEADD9910FFA4CFDFFD87E68A6E80" bold="true" box="[199,439,617,642]" pageId="10" pageNumber="521">Non-type specimens</emphasis>
(
<emphasis id="B95FEADD9910FFA4CED2FD84E6E56E83" box="[458,472,618,641]" italics="true" pageId="10" pageNumber="521">n</emphasis>
=17):
<collectionCode id="ED3AAE0A9910FFA4CD05FD84E5546E80" box="[541,617,618,642]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="10" pageNumber="521">MUFS</collectionCode>
10452104568,
<specimenCount id="9D2DFD469910FFA4CC33FD84E4846E80" box="[811,953,617,642]" pageId="10" pageNumber="521" type="generic">7 specimens</specimenCount>
,
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SL, Nago, Okinawa Island, Ryukyu Islands,
<collectingCountry id="F33C765F9910FFA4CE41FD60E6A16EA7" box="[345,412,654,677]" name="Japan" pageId="10" pageNumber="521">Japan</collectingCountry>
;
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<quantity id="4CD39B2A9910FFA4CD19FD63E5856EA7" box="[513,696,653,678]" metricMagnitude="4" metricUnit="m" metricValue="1.2267139" pageId="10" pageNumber="521" unit="mm" value="1.2267139E7">12267, 139 mm</quantity>
SL,
<collectingCountry id="F33C765F9910FFA4CDECFD63E4636EA4" box="[756,862,653,678]" name="Malaysia" pageId="10" pageNumber="521">Malaysia</collectingCountry>
;
<collectionCode id="ED3AAE0A9910FFA4CC74FD60E4846EA4" box="[876,953,654,678]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="10" pageNumber="521">MUFS</collectionCode>
<quantity id="4CD39B2A9910FFA4CCDAFD63E3476EA7" box="[962,1146,653,678]" metricMagnitude="4" metricUnit="m" metricValue="1.8621128999999996" pageId="10" pageNumber="521" unit="mm" value="1.8621129E7">18621, 129 mm</quantity>
SL and
<collectionCode id="ED3AAE0A9910FFA4CBF9FD60E2136EA4" box="[1249,1326,654,678]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="10" pageNumber="521">MUFS</collectionCode>
18886,
<specimenCount id="9D2DFD469910FFA4CA96FD60E6326EC8" pageId="10" pageNumber="521" type="generic">2 specimens</specimenCount>
,
<quantity id="4CD39B2A9910FFA4CE05FD5CE6886ECB" box="[285,437,690,714]" metricMagnitude="-1" metricUnit="m" metricValue="1.305" metricValueMax="1.3199999999999998" metricValueMin="1.29" pageId="10" pageNumber="521" unit="mm" value="130.5" valueMax="132.0" valueMin="129.0">129132 mm</quantity>
SL, Makassar (Ujung Pangdang), South Sulawesi,
<collectingCountry id="F33C765F9910FFA4CB15FD5FE3436EC8" box="[1037,1150,689,714]" name="Indonesia" pageId="10" pageNumber="521">Indonesia</collectingCountry>
;
<collectionCode id="ED3AAE0A9910FFA4CB94FD5CE3E56EC8" box="[1164,1240,690,714]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="10" pageNumber="521">MUFS</collectionCode>
4288442885,
<specimenCount id="9D2DFD469910FFA4CA96FD5CE6326EEC" pageId="10" pageNumber="521" type="generic">2 specimens</specimenCount>
,
<quantity id="4CD39B2A9910FFA4CE02FD3BE6AE6EEF" box="[282,403,725,750]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.4" metricValueMin="5.1" pageId="10" pageNumber="521" unit="mm" value="52.5" valueMax="54.0" valueMin="51.0">5154 mm</quantity>
SL, Indian Ocean (purchased by aquarium shop but presumably
<collectingCountry id="F33C765F9910FFA4CB74FD3BE3D46EEC" box="[1132,1257,725,750]" name="Seychelles" pageId="10" pageNumber="521">Seychelles</collectingCountry>
);
<collectionCode id="ED3AAE0A9910FFA4CBE4FD38E2756EEC" box="[1276,1352,726,750]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="10" pageNumber="521">MUFS</collectionCode>
<quantity id="4CD39B2A9910FFA4CA57FD38E7CA6F13" metricMagnitude="4" metricUnit="m" metricValue="4.0234112" pageId="10" pageNumber="521" unit="mm" value="4.0234112E7">40234, 112 mm</quantity>
SL, Meitsu, Miyazaki,
<collectingCountry id="F33C765F9910FFA4CD16FD14E56C6F13" box="[526,593,762,785]" name="Japan" pageId="10" pageNumber="521">Japan</collectingCountry>
;
<collectionCode id="ED3AAE0A9910FFA4CD78FD14E5896F13" LSID="urn:lsid:biocol.org:col:34992" box="[608,692,762,785]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="521">RMNH</collectionCode>
<collectionCode id="ED3AAE0A9910FFA4CDA7FD14E5C36F10" box="[703,766,762,786]" pageId="10" pageNumber="521">PISC</collectionCode>
. 38327 (
<typeStatus id="5490886D9910FFA4CC7CFD17E4F06F10" box="[868,973,761,786]" pageId="10" pageNumber="521" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C2B4D4C9910FFA4CCE1FD14E22F6F13" box="[1017,1298,761,785]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="pomacanthus">
<emphasis id="B95FEADD9910FFA4CCE1FD14E22F6F13" box="[1017,1298,761,785]" italics="true" pageId="10" pageNumber="521">Mesoprion pomacanthus</emphasis>
</taxonomicName>
, designated herein),
<quantity id="4CD39B2A9910FFA4CFECFCF0E67F6F37" box="[244,322,798,822]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="521" unit="mm" value="30.0">30 mm</quantity>
SL (37+ mm TL),
<collectingCountry id="F33C765F9910FFA4CD0EFCF3E5576F34" box="[534,618,797,822]" name="Indonesia" pageId="10" pageNumber="521">Ambon</collectingCountry>
Island, Molucca Islands,
<collectingCountry id="F33C765F9910FFA4CC90FCF3E4C56F34" box="[904,1016,797,822]" name="Indonesia" pageId="10" pageNumber="521">Indonesia</collectingCountry>
, removed from
<collectionCode id="ED3AAE0A9910FFA4CBBFFCF0E3C66F37" LSID="urn:lsid:biocol.org:col:34992" box="[1191,1275,798,821]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="521">RMNH</collectionCode>
5521;
<collectionCode id="ED3AAE0A9910FFA4CA50FCF0E2A16F37" LSID="urn:lsid:biocol.org:col:34992" box="[1352,1436,798,821]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="521">RMNH</collectionCode>
<collectionCode id="ED3AAE0A9910FFA4CF8FFCACE7EB6F58" box="[151,214,834,858]" pageId="10" pageNumber="521">PISC</collectionCode>
. 38328 (
<typeStatus id="5490886D9910FFA4CE2CFCAFE6F06F58" box="[308,461,833,858]" pageId="10" pageNumber="521" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C2B4D4C9910FFA4CEEBFCACE5906F5B" box="[499,685,833,857]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="pomacanthus">
<emphasis id="B95FEADD9910FFA4CEEBFCACE5906F5B" box="[499,685,833,857]" italics="true" pageId="10" pageNumber="521">M. pomacanthus</emphasis>
</taxonomicName>
), same locality as the
<typeStatus id="5490886D9910FFA4CCB0FCAFE3326F58" box="[936,1039,833,858]" pageId="10" pageNumber="521" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C2B4D4C9910FFA4CB2CFCACE3D36F5B" box="[1076,1262,833,857]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="pomacanthus">
<emphasis id="B95FEADD9910FFA4CB2CFCACE3D36F5B" box="[1076,1262,833,857]" italics="true" pageId="10" pageNumber="521">M. pomacanthus</emphasis>
</taxonomicName>
, removed from
<collectionCode id="ED3AAE0A9910FFA4CF8FFC88E7D66F7F" LSID="urn:lsid:biocol.org:col:34992" box="[151,235,870,893]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="521">RMNH</collectionCode>
5521; SU 26900 at
<collectionCode id="ED3AAE0A9910FFA4CED4FC88E5386F7C" LSID="urn:lsid:biocol.org:col:34700" box="[460,517,870,894]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="10" pageNumber="521">CAS</collectionCode>
,
<quantity id="4CD39B2A9910FFA4CD08FC88E5516F7F" box="[528,620,870,894]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="10" pageNumber="521" unit="mm" value="112.0">112 mm</quantity>
SL,
<collectingCountry id="F33C765F9910FFA4CDB9FC8BE41E6F7C" box="[673,803,869,894]" name="Philippines" pageId="10" pageNumber="521">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8B9436CF9910FFA4CFDFFC67E5876FC7" blockId="10.[151,1436,581,1974]" pageId="10" pageNumber="521">
<emphasis id="B95FEADD9910FFA4CFDFFC67E55A6FA0" bold="true" box="[199,615,905,930]" pageId="10" pageNumber="521">Photographic voucher specimens.</emphasis>
KAUM-I. 1226612267,
<specimenCount id="9D2DFD469910FFA4CCBCFC64E3056FA0" box="[932,1080,905,930]" pageId="10" pageNumber="521" type="generic">2 specimens</specimenCount>
,
<quantity id="4CD39B2A9910FFA4CB54FC64E3D46FA3" box="[1100,1257,906,930]" metricMagnitude="-1" metricUnit="m" metricValue="1.3599999999999999" metricValueMax="1.42" metricValueMin="1.3" pageId="10" pageNumber="521" unit="mm" value="136.0" valueMax="142.0" valueMin="130.0">130142 mm</quantity>
SL,
<collectingCountry id="F33C765F9910FFA4CA36FC67E2A56FA0" box="[1326,1432,905,930]" name="Malaysia" pageId="10" pageNumber="521">Malaysia</collectingCountry>
; KAUM-I. 54316, 96 mm SL, Kagoshima,
<collectingCountry id="F33C765F9910FFA4CD6BFC40E58A6FC7" box="[627,695,942,965]" name="Japan" pageId="10" pageNumber="521">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33165449910FFA4CFDFFC3FE7C66908" pageId="10" pageNumber="521" type="diagnosis">
<paragraph id="8B9436CF9910FFA4CFDFFC3FE7C66908" blockId="10.[151,1436,581,1974]" pageId="10" pageNumber="521">
<emphasis id="B95FEADD9910FFA4CFDFFC3FE67C6FE8" bold="true" box="[199,321,977,1002]" pageId="10" pageNumber="521">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C2B4D4C9910FFA4CEF6FC3CE56D6FEB" box="[494,592,978,1001]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="genus">
<emphasis id="B95FEADD9910FFA4CEF6FC3CE56D6FEB" box="[494,592,978,1001]" italics="true" pageId="10" pageNumber="521">Lutjanus</emphasis>
</taxonomicName>
with the following combination of characters: Dorsal fin XI, 13 or 14 (usually 13); anal fin III, 8; pectoral-fin 16 or 17; lateral-line scales 4750; horizontal scale rows above lateral line 8 or 9; scale rows above lateral line rising obliquely toward dorsal profile; predorsal scales extending forward nearly to level of just above anterior edge of orbit; scale rows on cheek 811; no subocular extension of cheek scales; gill rakers on first arch 7 or 8 + 1517 = 2225; tongue smooth, without teeth; preopercular notch developed just above angle; interopercular knob inconspicuous; the first blue stripe running from above anterior edge of eye ending below the 79th dorsal-fin spine base; the third stripe usually running from middle of upper part of opercle or anterior edge of upper opercle to below posterior fourth of soft portion of dorsal fin, reaching the last soft dorsalray base.
</paragraph>
</subSubSection>
<subSubSection id="C33165449910FFA5CFDFFAFBE2616901" lastPageId="11" lastPageNumber="522" pageId="10" pageNumber="521" type="description">
<paragraph id="8B9436CF9910FFA4CFDFFAFBE4286974" blockId="10.[151,1436,581,1974]" pageId="10" pageNumber="521">
<emphasis id="B95FEADD9910FFA4CFDFFAFBE664692C" bold="true" box="[199,345,1301,1326]" pageId="10" pageNumber="521">Description.</emphasis>
Meristic counts and proportional measurements as percentages of standard length (SL), plus two diagnostic characters in coloration and squamation of subocular extension for the
<typeStatus id="5490886D9910FFA4CB92FAD7E3CD6950" box="[1162,1264,1337,1362]" pageId="10" pageNumber="521" type="lectotype">lectotype</typeStatus>
and non-type specimens of
<taxonomicName id="4C2B4D4C9910FFA4CE29FAB0E5226977" box="[305,543,1373,1397]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9910FFA4CE29FAB0E5226977" box="[305,543,1373,1397]" italics="true" pageId="10" pageNumber="521">Lutjanus bengalensis</emphasis>
</taxonomicName>
are shown in Table 1.
</paragraph>
<paragraph id="8B9436CF9910FFA4CFDFFA6FE5D26BB4" blockId="10.[151,1436,581,1974]" pageId="10" pageNumber="521">
Body oblong to moderately deep, greatest depth 2.4
<emphasis id="B95FEADD9910FFA4CC16FA6CE421699B" box="[782,796,1410,1433]" italics="true" pageId="10" pageNumber="521"></emphasis>
2.8, head length
<quantity id="4CD39B2A9910FFA4CCC0FA6FE3746998" box="[984,1097,1409,1434]" metricMagnitude="-2" metricUnit="m" metricValue="6.731" metricValueMax="7.111999999999999" metricValueMin="6.35" pageId="10" pageNumber="521" unit="in" value="2.65" valueMax="2.8" valueMin="2.5">2.52.8 in</quantity>
SL; snout length 2.63.4, eye diameter 3.04.2, interorbita1 width 4.55.4, maxilla length 2.22.9 and preorbital depth 5.17.3, in head length; head relatively small with pointed snout; snout-forehead profile weakly rounded; pair of small nostrils rounded anteriorly and somewhat ovate posteriorly, in anterior and posterior openings on each side of snout, with very low flap on outer edge of anterior nostril; body scales rising obliquely above lateral line; body, opercle and preopercle with finely ctenoid scales; parietal, frontal, interorbital and rear edge of preopercle scaly; preopercular margin finely serrate (serrae increasing in size ventrally); sharp, flattened, flexible projection at middle of opercular margin, with blunt bony spine (mostly covered by scales) immediately above, its base; margins of opercle, interopercle and subopercle smooth; preopercular notch and interopercular knob well developed; vomerine tooth patch crescent-shaped without a posterior extension (similar small-sized lateral canine teeth present in 1 row on both right and left branches); tongue smooth without granular teeth; upper jaw with dagger-like very short canine tooth, on each side anteriorly, with smaller canines between it and symphysis; remaining lateral portion of upper jaw with a series of ~10 visible smaller embedded canines, lower jaw with ~10 small embedded canine teeth on each side of jaws, most not visible when mouth closed; dorsal-fin outline very weakly incised, fourth or fifth spines longest, remaining spines gradually decreasing in length; profile of soft dorsal-fin relatively low and weakly rounded; pectoral fins pointed, caudal fin emarginate.
</paragraph>
<caption id="DF5466479911FFA5CF8FFCABE3AE6F59" box="[151,1171,837,859]" httpUri="https://zenodo.org/record/264486/files/figure.png" pageId="11" pageNumber="522" targetBox="[194,1391,193,815]" targetPageId="11">
<paragraph id="8B9436CF9911FFA5CF8FFCABE3AE6F59" blockId="11.[151,1171,837,859]" box="[151,1171,837,859]" pageId="11" pageNumber="522">
<emphasis id="B95FEADD9911FFA5CF8FFCABE62F6F58" bold="true" box="[151,274,837,859]" pageId="11" pageNumber="522">FIGURE 4.</emphasis>
<taxonomicName id="4C2B4D4C9911FFA5CE01FCA8E51C6F59" box="[281,545,837,859]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="522" phylum="Chordata" rank="species" species="sapphirolineatus" status="sp. nov.">
<emphasis id="B95FEADD9911FFA5CE01FCA8E51C6F59" box="[281,545,837,859]" italics="true" pageId="11" pageNumber="522">Lutjanus sapphirolineatus</emphasis>
</taxonomicName>
<emphasis id="B95FEADD9911FFA5CD3FFCABE5556F58" bold="true" box="[551,616,837,858]" pageId="11" pageNumber="522">
<taxonomicNameLabel id="A26C57A69911FFA5CD3FFCABE55E6F58" box="[551,611,837,858]" pageId="11" pageNumber="522" rank="species">n. sp.</taxonomicNameLabel>
:
</emphasis>
MUFS 33724, holotype, 153 mm SL, Muttrah, Oman.
</paragraph>
</caption>
<paragraph id="8B9436CF9911FFA5CFDFFC6CE5BF68DD" blockId="11.[151,1437,898,2003]" pageId="11" pageNumber="522">
<emphasis id="B95FEADD9911FFA5CFDFFC6CE65D6F98" box="[199,352,898,922]" italics="true" pageId="11" pageNumber="522">Color in life,</emphasis>
based on SAIABs Indian Ocean specimens (
<specimenCount id="9D2DFD469911FFA5CC98FC6DE32C6F99" box="[896,1041,898,923]" pageId="11" pageNumber="522" type="generic">2 specimens</specimenCount>
, SAIAB 82483 and uncatalogued 1815#6, 94 and
<quantity id="4CD39B2A9911FFA5CE72FC48E6EC6FBC" box="[362,465,934,959]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="11" pageNumber="522" unit="mm" value="105.0">105 mm</quantity>
SL, respectively, Sodwana Bay,
<collectingCountry id="F33C765F9911FFA5CC6CFC48E3326FBD" box="[884,1039,934,959]" name="South Africa" pageId="11" pageNumber="522">South Africa</collectingCountry>
) and Pacific specimens (MUFS 1045210458,
<specimenCount id="9D2DFD469911FFA5CE27FC25E6F66FE1" box="[319,459,970,995]" pageId="11" pageNumber="522" type="generic">7 specimens</specimenCount>
,
<quantity id="4CD39B2A9911FFA5CECEFC24E5566FE0" box="[470,619,970,995]" metricMagnitude="-1" metricUnit="m" metricValue="1.775" metricValueMax="1.9" metricValueMin="1.65" pageId="11" pageNumber="522" unit="mm" value="177.5" valueMax="190.0" valueMin="165.0">165190 mm</quantity>
SL and uncatalogued photographs from
<collectingCountry id="F33C765F9911FFA5CB2AFC25E3456FE1" box="[1074,1144,971,995]" name="Japan" pageId="11" pageNumber="522">Japan</collectingCountry>
): head and body generally bright yellow on upper half and white ventrally, sometimes with reddish tinge, often after death; upper side with four dark-edged blue stripes, uppermost stripe extending from interorbital to middle of spinous dorsal fin, the second from upper rear edge of eye to anterior part of soft dorsal-fin base, third stripe from posterodorsal margin of preopercle or middle of upper part of opercle to area posteriorly on body a short distance anterior to base of last soft dorsal-fin rays, and lowest stripe from just above of middle of maxilla across lower edge of eye, extending through middle preopercle and opercle to anterior third of caudal peduncle just below lateral line; median fins yellow; pectoral and pelvic fins whitish; iris yellow.
</paragraph>
<paragraph id="8B9436CF9911FFA5CFDFFB04E2616901" blockId="11.[151,1437,898,2003]" box="[199,1372,1258,1283]" pageId="11" pageNumber="522">
<emphasis id="B95FEADD9911FFA5CFDFFB04E6B46900" box="[199,393,1258,1282]" italics="true" pageId="11" pageNumber="522">Color in alcohol:</emphasis>
generally yellowish tan (including fins), with 4 brown or bluish stripes as noted above.
</paragraph>
</subSubSection>
<subSubSection id="C33165449911FFA2CFDFFAE0E3856D1F" lastPageId="12" lastPageNumber="523" pageId="11" pageNumber="522" type="distribution">
<paragraph id="8B9436CF9911FFA5CFDFFAE0E4746A8D" blockId="11.[151,1437,898,2003]" pageId="11" pageNumber="522">
<emphasis id="B95FEADD9911FFA5CFDFFAE0E65D6925" bold="true" box="[199,352,1294,1319]" pageId="11" pageNumber="522">Distribution.</emphasis>
<taxonomicName id="4C2B4D4C9911FFA5CE71FAE1E56A6924" box="[361,599,1294,1318]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="522" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9911FFA5CE71FAE1E56A6924" box="[361,599,1294,1318]" italics="true" pageId="11" pageNumber="522">Lutjanus bengalensis</emphasis>
</taxonomicName>
is widely distributed in the Indo-West Pacific (
<bibRefCitation id="EFBA4B3E9911FFA5CB6CFAE0E7E86949" author="Allen" pageId="11" pageNumber="522" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot 1985; this study</bibRefCitation>
) and is currently known from
<collectingCountry id="F33C765F9911FFA5CD2BFADCE5FA6949" box="[563,711,1330,1355]" name="South Africa" pageId="11" pageNumber="522">South Africa</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CDCCFADCE4726949" box="[724,847,1330,1355]" name="Seychelles" pageId="11" pageNumber="522">Seychelles</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CC44FADCE4F16949" box="[860,972,1330,1355]" name="Indonesia" pageId="11" pageNumber="522">Indonesia</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CCC0FADCE37C6949" box="[984,1089,1330,1355]" name="Malaysia" pageId="11" pageNumber="522">Malaysia</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CB56FADCE3F16949" box="[1102,1228,1330,1355]" name="Philippines" pageId="11" pageNumber="522">Philippines</collectingCountry>
and
<collectingCountry id="F33C765F9911FFA5CA1EFADDE2746948" box="[1286,1353,1331,1354]" name="Japan" pageId="11" pageNumber="522">Japan</collectingCountry>
, on the basis of specimens examined. The species is also confirmed from the following areas by photographic vouchers and references (
<bibRefCitation id="EFBA4B3E9911FFA5CE48FA94E5036991" author="Kyushin" box="[336,574,1402,1427]" pageId="11" pageNumber="522" refString="Kyushin, K., Amaoka, K., Nakaya, K. &amp; Ida, H. (1977) Fishes of Indian Ocean. Japan Marine Fishery Resource Research Center, Tokyo, 392 pp., 179 pls. [in Japanese and English]" type="book" year="1977">
Kyushin
<emphasis id="B95FEADD9911FFA5CEA4FA95E6CC6990" box="[444,497,1402,1426]" italics="true" pageId="11" pageNumber="522">et al</emphasis>
. 1977
</bibRefCitation>
;
<bibRefCitation id="EFBA4B3E9911FFA5CD56FA94E4706991" author="Allen" box="[590,845,1402,1427]" pageId="11" pageNumber="522" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot, 1985</bibRefCitation>
;
<bibRefCitation id="EFBA4B3E9911FFA5CC45FA94E4E66991" author="Shen" box="[861,987,1402,1427]" pageId="11" pageNumber="522" refString="Shen, S. C. (Ed.) (1993) Fishes of Taiwan. Department of Zoology, National Taiwan University, Taipei, 960 pp." type="book" year="1993">Shen 1993</bibRefCitation>
;
<bibRefCitation id="EFBA4B3E9911FFA5CCF3FA94E3516991" author="Chen" box="[1003,1132,1402,1427]" pageId="11" pageNumber="522" refString="Chen, C. - H. (1996) Checklist of the fishes of Penghu. FRI Special Publication, 4, 1 - 175." type="journal article" year="1996">Chen 1996</bibRefCitation>
;
<bibRefCitation id="EFBA4B3E9911FFA5CB64FA94E2496991" author="Randall" box="[1148,1396,1402,1427]" pageId="11" pageNumber="522" refString="Randall, J. E. &amp; Lim, K. K. P. (Eds.) (2000) A checklist of the fishes of the South China Sea. Raffles Bulletin of Zoology, 8 (Supplement), 569 - 667." type="journal article" year="2000">Randall &amp; Lim 2000</bibRefCitation>
; E. Heemstra pers. comm.; U. Satapoomin pers. comm.):
<collectingCountry id="F33C765F9911FFA5CC17FA70E49B69B5" box="[783,934,1438,1463]" name="Mozambique" pageId="11" pageNumber="522">Mozambique</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CCACFA70E30069B5" box="[948,1085,1438,1463]" name="Madagascar" pageId="11" pageNumber="522">Madagascar</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CB54FA71E3A569B5" box="[1100,1176,1439,1463]" name="Kenya" pageId="11" pageNumber="522">Kenya</collectingCountry>
, Saya de Malha Bank (southwestern area off the Chagos Archipelago), Andaman Sea (Similan Island),
<collectingCountry id="F33C765F9911FFA5CB35FA2CE34269D9" box="[1069,1151,1474,1499]" name="Taiwan" pageId="11" pageNumber="522">Taiwan</collectingCountry>
and
<collectingCountry id="F33C765F9911FFA5CBAFFA2CE3C369D9" box="[1207,1278,1474,1499]" name="China" pageId="11" pageNumber="522">China</collectingCountry>
. Furthermore, photographic vouchers (E. Heemstra, pers. comm.) show the species from the following localities: southern and northern
<collectingCountry id="F33C765F9911FFA5CFE4F9E4E6AE6A21" box="[252,403,1546,1571]" name="Mozambique" pageId="11" pageNumber="522">Mozambique</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CE85F9E4E5196A21" box="[413,548,1546,1571]" name="Madagascar" pageId="11" pageNumber="522">Madagascar</collectingCountry>
and
<collectingCountry id="F33C765F9911FFA5CD42F9E4E5D46A21" box="[602,745,1546,1571]" name="South Africa" pageId="11" pageNumber="522">South Africa</collectingCountry>
(Sodwana Bay and Cape Vidal), plus a photograph that might be the species from
<collectingCountry id="F33C765F9911FFA5CE98F9C0E6DB6A45" box="[384,486,1582,1607]" name="Tanzania" pageId="11" pageNumber="522">Zanzibar</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CEEBF9C0E5616A45" box="[499,604,1582,1607]" name="Tanzania" pageId="11" pageNumber="522">Tanzania</collectingCountry>
. However, records of the distribution of
<taxonomicName id="4C2B4D4C9911FFA5CB2AF9C1E3EB6A44" box="[1074,1238,1582,1606]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="522" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9911FFA5CB2AF9C1E3EB6A44" box="[1074,1238,1582,1606]" italics="true" pageId="11" pageNumber="522">L. bengalensis</emphasis>
</taxonomicName>
from the Middle East, including the Red Sea, may have been erroneous (
<bibRefCitation id="EFBA4B3E9911FFA5CC18F9BCE4D66A69" author="Allen" box="[768,1003,1618,1643]" pageId="11" pageNumber="522" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot 1985</bibRefCitation>
), because
<taxonomicName id="4C2B4D4C9911FFA5CB45F9BDE3C36A68" box="[1117,1278,1618,1642]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="522" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9911FFA5CB45F9BDE3C36A68" box="[1117,1278,1618,1642]" italics="true" pageId="11" pageNumber="522">L. bengalensis</emphasis>
</taxonomicName>
was confused with
<taxonomicName id="4C2B4D4C9911FFA5CFD7F999E6956A8C" box="[207,424,1654,1678]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="522" phylum="Chordata" rank="species" species="sapphirolineatus" status="sp. nov.">
<emphasis id="B95FEADD9911FFA5CFD7F999E6956A8C" box="[207,424,1654,1678]" italics="true" pageId="11" pageNumber="522">L. sapphirolineatus</emphasis>
</taxonomicName>
<emphasis id="B95FEADD9911FFA5CEB7F998E6D36A8D" bold="true" box="[431,494,1654,1679]" pageId="11" pageNumber="522">
<taxonomicNameLabel id="A26C57A69911FFA5CEB7F998E6D36A8D" box="[431,494,1654,1679]" pageId="11" pageNumber="522" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, described below in this study.
</paragraph>
<paragraph id="8B9436CF9911FFA5CFDFF975E38D6B41" blockId="11.[151,1437,898,2003]" pageId="11" pageNumber="522">
In occurrence notes on
<taxonomicName id="4C2B4D4C9911FFA5CEC5F975E5BE6AB0" box="[477,643,1690,1714]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="522" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9911FFA5CEC5F975E5BE6AB0" box="[477,643,1690,1714]" italics="true" pageId="11" pageNumber="522">L. bengalensis</emphasis>
</taxonomicName>
from the western Pacific,
<bibRefCitation id="EFBA4B3E9911FFA5CCD9F974E3F56AB1" author="Allen" box="[961,1224,1690,1715]" pageId="11" pageNumber="522" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot (1985)</bibRefCitation>
and Allen (1985) reported one record of this species from
<collectingCountry id="F33C765F9911FFA5CD42F950E58C6AD5" box="[602,689,1726,1751]" name="Indonesia" pageId="11" pageNumber="522">Ambon</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CDA3F950E4116AD5" box="[699,812,1726,1751]" name="Indonesia" pageId="11" pageNumber="522">Indonesia</collectingCountry>
. An underwater photograph clearly showing a school of snappers (
<bibRefCitation id="EFBA4B3E9911FFA5CE14F90CE51B6AF9" author="Allen" box="[268,550,1762,1787]" pageId="11" pageNumber="522" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef Fishes of the East Indies. Vol. II. Tropical Reef Research, Perth, 856 pp." type="book" year="2012">Allen &amp; Erdmann 2012</bibRefCitation>
), as
<taxonomicName id="4C2B4D4C9911FFA5CD79F90DE5E06AF8" box="[609,733,1762,1786]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="522" phylum="Chordata" rank="species" species="kasmira">
<emphasis id="B95FEADD9911FFA5CD79F90DE5E06AF8" box="[609,733,1762,1786]" italics="true" pageId="11" pageNumber="522">L. kasmira</emphasis>
</taxonomicName>
, was taken at Mios Kon, Raja Ampat Islands, West
<collectingCountry id="F33C765F9911FFA5CA48F90DE2A56AF8" box="[1360,1432,1763,1786]" name="Papua New Guinea" pageId="11" pageNumber="522">Papua</collectingCountry>
,
<collectingCountry id="F33C765F9911FFA5CF8FF8E8E6396B1D" box="[151,260,1798,1823]" name="Indonesia" pageId="11" pageNumber="522">Indonesia</collectingCountry>
(by M. Allen), and could be the easternmost record of
<taxonomicName id="4C2B4D4C9911FFA5CC6CF8E9E32A6B1C" box="[884,1047,1798,1822]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="522" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9911FFA5CC6CF8E9E32A6B1C" box="[884,1047,1798,1822]" italics="true" pageId="11" pageNumber="522">L. bengalensis</emphasis>
</taxonomicName>
(A. Ryanskiy, pers. comm.); there are no gray (or narrow reddish) stripes on the abdomen, and no dusky upper pectoral-fin rays.
</paragraph>
<paragraph id="8B9436CF9911FFA2CFDFF8A1E3856D1F" blockId="11.[151,1437,898,2003]" lastBlockId="12.[151,1437,151,1544]" lastPageId="12" lastPageNumber="523" pageId="11" pageNumber="522">
In the northwestern Pacific, the species was firstly reported from Okinawa Island,
<collectingCountry id="F33C765F9911FFA5CB46F8A1E3A36B64" box="[1118,1182,1871,1894]" name="Japan" pageId="11" pageNumber="522">Japan</collectingCountry>
(
<bibRefCitation id="EFBA4B3E9911FFA5CBB4F8A0E2AD6B65" author="Iwatsuki" box="[1196,1424,1870,1895]" pageId="11" pageNumber="522" refString="Iwatsuki, Y., Yoshino, T. &amp; Shimada, K. (1999) Comparison of Lutjanus bengalensis from the western Pacific with a related species, L. kasmira, and variations in both species (Perciformes: Lutjanidae). Ichthyological Research, 46 (3), 314 - 317. http: // dx. doi. org / 10.1007 / BF 02678519" type="journal article" year="1999">
Iwatsuki
<emphasis id="B95FEADD9911FFA5CA0DF8A1E27B6B64" box="[1301,1350,1870,1894]" italics="true" pageId="11" pageNumber="522">et al</emphasis>
. 1999
</bibRefCitation>
). Currently this species is very common from southern
<collectingCountry id="F33C765F9911FFA5CC18F89DE47D6B88" box="[768,832,1907,1930]" name="Japan" pageId="11" pageNumber="522">Japan</collectingCountry>
(Ryukyu Islands, and ranging as far north as the Kii Peninsula (
<bibRefCitation id="EFBA4B3E9911FFA5CE0AF878E6D76BAD" author="Kimura" box="[274,490,1942,1967]" pageId="11" pageNumber="522" refString="Kimura, S., Sakakibara, H., Tsumoto, K. &amp; Iwatsuki, Y. (2008) A lutjanid fish, Lutjanus bengalensis (Actinopterygii: Perciformes), from Owase: the first record from Mie Prefecture and northernmost records for the species. Bulletin of Faculty of Mie University, 35, 65 - 69." type="journal article" year="2008">
Kimura
<emphasis id="B95FEADD9911FFA5CE68F879E69C6BAC" box="[368,417,1942,1966]" italics="true" pageId="11" pageNumber="522">et al</emphasis>
. 2008
</bibRefCitation>
), Tokai District, and Kanto District near Tokyo, observed by the first author), as well as
<collectingCountry id="F33C765F9911FFA5CFAFF854E6346BD1" box="[183,265,1978,2003]" name="Taiwan" pageId="11" pageNumber="522">Taiwan</collectingCountry>
and southern
<collectingCountry id="F33C765F9911FFA5CEB3F854E6CF6BD1" box="[427,498,1978,2003]" name="China" pageId="11" pageNumber="522">China</collectingCountry>
. However, during the period
<date id="FF95100F9911FFA5CC27F854E4866BD1" box="[831,955,1978,2003]" pageId="11" pageNumber="522" value="1983" valueMax="1995">1983-1995</date>
, the first author visited southwestern Asia (
<collectingCountry id="F33C765F9916FFA2CF86FF79E61C6CB2" box="[158,289,151,176]" name="Philippines" pageId="12" pageNumber="523">Philippines</collectingCountry>
,
<collectingCountry id="F33C765F9916FFA2CE34FF79E6A16CB2" box="[300,412,151,176]" name="Indonesia" pageId="12" pageNumber="523">Indonesia</collectingCountry>
,
<collectingCountry id="F33C765F9916FFA2CEBFFF79E5246CB2" box="[423,537,151,176]" name="Singapore" pageId="12" pageNumber="523">Singapore</collectingCountry>
and
<collectingCountry id="F33C765F9916FFA2CD48FF79E5996CB2" box="[592,676,151,176]" name="Taiwan" pageId="12" pageNumber="523">Taiwan</collectingCountry>
) several times, but identified only one specimen (ca.
<quantity id="4CD39B2A9916FFA2CBE3FF79E27C6CAD" box="[1275,1345,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="523" unit="cm" value="15.0">15 cm</quantity>
TL, not kept) of this species at the Chinese fish market in
<collectingCountry id="F33C765F9916FFA2CDD0FF52E4006CD7" box="[712,829,188,213]" name="Singapore" pageId="12" pageNumber="523">Singapore</collectingCountry>
; it seemed a rare species from the Pacific at that time. The distributional range of
<taxonomicName id="4C2B4D4C9916FFA2CED6FF0FE54D6CFA" box="[462,624,224,248]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CED6FF0FE54D6CFA" box="[462,624,224,248]" italics="true" pageId="12" pageNumber="523">L. bengalensis</emphasis>
</taxonomicName>
as well as
<taxonomicName id="4C2B4D4C9916FFA2CDE8FF0FE49D6CFA" box="[752,928,224,248]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="xanthopinnis">
<emphasis id="B95FEADD9916FFA2CDE8FF0FE49D6CFA" box="[752,928,224,248]" italics="true" pageId="12" pageNumber="523">L. xanthopinnis</emphasis>
</taxonomicName>
is likely to extend northwards, southwestern
<collectingCountry id="F33C765F9916FFA2CF8FFEEBE7EA6D1E" box="[151,215,261,284]" name="Japan" pageId="12" pageNumber="523">Japan</collectingCountry>
from the northwestern Pacific presumably due to global warming (
<bibRefCitation id="EFBA4B3E9916FFA2CCDFFEEAE3916D1F" author="Iwatsuki" box="[967,1196,260,285]" pageId="12" pageNumber="523" refString="Iwatsuki, Y., Tanaka, F. &amp; Allen, G. R. (2015) Lutjanus xanthopinnis, a new species of snapper (Pisces: Lutjanidae) from the Indo-west Pacific, with a redescription of Lutjanus madras (Valenciennes 1831). Journal of the Ocean Science Foundation, 17, 22 - 42." type="journal article" year="2015">
Iwatsuki
<emphasis id="B95FEADD9916FFA2CB28FEEBE35D6D1E" box="[1072,1120,260,284]" italics="true" pageId="12" pageNumber="523">et al</emphasis>
. 2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33165449916FFA2CFDFFEC9E22B6A0A" pageId="12" pageNumber="523" type="discussion">
<paragraph id="8B9436CF9916FFA2CFDFFEC9E6856DAF" blockId="12.[151,1437,151,1544]" pageId="12" pageNumber="523">
<emphasis id="B95FEADD9916FFA2CFDFFEC9E6066D42" bold="true" box="[199,315,295,320]" pageId="12" pageNumber="523">Remarks.</emphasis>
A
<typeStatus id="5490886D9916FFA2CE7CFEC9E6F06D42" box="[356,461,295,320]" pageId="12" pageNumber="523" type="lectotype">lectotype</typeStatus>
, ZMB 303, was designated by
<bibRefCitation id="EFBA4B3E9916FFA2CC26FEC9E4D96D42" author="Paepke" box="[830,996,295,320]" pageId="12" pageNumber="523" refString="Paepke, H. - J. (1999) Bloch's fish collection in the Museum fur Naturkunde der Humboldt Universitat zu Berlin: an illustrated catalog and historical account. Ruggell (Liechtenstein). Theses Zoologicae, 32, 1 - 216, pls. 1 - 32 (unpaginated)." type="journal article" year="1999">Paepke (1999)</bibRefCitation>
and we confirm this specimen as the
<typeStatus id="5490886D9916FFA2CF8FFEA2E7C36D67" box="[151,254,332,357]" pageId="12" pageNumber="523" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="13102A4A9916FFA2CE0EFEA2E6626D66" box="[278,351,332,357]" captionStart="FIGURE 1" captionStartId="6.[151,250,1902,1924]" captionTargetBox="[188,1398,216,1878]" captionTargetId="figure@6.[188,1398,195,1878]" captionTargetPageId="6" captionText="FIGURE 1. Type specimens of Lutjanus octolineatus (A, B) and L. bengalensis (C): A. MNHN 0000 - A 5446 (no label attached with a blue glass plate), 122 mm SL, lectotype of Diacope octolineata, Mauritius, Mascarenes; B. MNHN 0000 - 7046 (holotype of Diacope octovittata), 137 mm SL, Mauritius, Mascarenes; C. ZMB 303, lectotype of Holocentrus bengalensis), 94 mm SL, Bengal, Bay of Bengal, eastern Indian Ocean." httpUri="https://zenodo.org/record/264483/files/figure.png" pageId="12" pageNumber="523">Fig. 1</figureCitation>
C,
<quantity id="4CD39B2A9916FFA2CE94FEA2E6D96D66" box="[396,484,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="12" pageNumber="523" unit="mm" value="94.0">94 mm</quantity>
SL, Bengal, Bay of Bengal,
<collectingCountry id="F33C765F9916FFA2CC47FEA2E4A76D67" box="[863,922,332,357]" name="India" pageId="12" pageNumber="523">India</collectingCountry>
) of
<taxonomicName id="4C2B4D4C9916FFA2CCC2FEA2E2A46D66" authority="Bloch 1790" authorityName="Bloch" authorityYear="1790" box="[986,1433,332,357]" class="Actinopterygii" family="Holocentridae" genus="Holocentrus" kingdom="Animalia" order="Beryciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CCC2FEA2E3C46D66" box="[986,1273,332,356]" italics="true" pageId="12" pageNumber="523">Holocentrus bengalensis</emphasis>
<bibRefCitation id="EFBA4B3E9916FFA2CA12FEA2E2A46D66" author="Bloch" box="[1290,1433,332,357]" pageId="12" pageNumber="523" refString="Bloch, M. E. (1790) Naturgeschichte der auslandischen Fische. Vol. 4. Icthyologie, ou Histoire naturelle des Poissons, Berlin, xii + 128 pp., pls. 217 - 252." type="book chapter" year="1790">Bloch 1790</bibRefCitation>
</taxonomicName>
. Unfortunately, we did not have the opportunity to examine the
<typeStatus id="5490886D9916FFA2CC71FE81E3336D8A" box="[873,1038,367,392]" pageId="12" pageNumber="523" type="paralectotype">paralectotypes</typeStatus>
, ZMB 300 (
<specimenCount id="9D2DFD469916FFA2CB86FE9EE2226D8A" box="[1182,1311,367,392]" pageId="12" pageNumber="523" type="generic">1 specimen</specimenCount>
) and ZMB 33057 (1, right side skin).
</paragraph>
<paragraph id="8B9436CF9916FFA2CFDFFE56E3D96E3E" blockId="12.[151,1437,151,1544]" pageId="12" pageNumber="523">
<taxonomicName id="4C2B4D4C9916FFA2CFDFFE56E6896DD2" box="[199,436,440,464]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CFDFFE56E6896DD2" box="[199,436,440,464]" italics="true" pageId="12" pageNumber="523">Lutjanus bengalensis</emphasis>
</taxonomicName>
has long been confused with
<taxonomicName id="4C2B4D4C9916FFA2CC1DFE57E4916DD2" box="[773,940,440,464]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="octolineatus">
<emphasis id="B95FEADD9916FFA2CC1DFE57E4916DD2" box="[773,940,440,464]" italics="true" pageId="12" pageNumber="523">L. octolineatus</emphasis>
</taxonomicName>
, but careful examination of the
<typeStatus id="5490886D9916FFA2CA0EFE59E2416DD2" box="[1302,1404,439,464]" pageId="12" pageNumber="523" type="lectotype">lectotype</typeStatus>
of this species shows a blue stripe on the upper part of opercle, 11 dorsal-fin spines and no suborbital extension of cheek scales (vs. no stripe on upper part of opercle, 12 dorsal-fin spines and the presence of a suborbital extension of cheek scales in
<taxonomicName id="4C2B4D4C9916FFA2CE7CFDCBE5376E3E" box="[356,522,548,572]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="octolineatus">
<emphasis id="B95FEADD9916FFA2CE7CFDCBE5376E3E" box="[356,522,548,572]" italics="true" pageId="12" pageNumber="523">L. octolineatus</emphasis>
</taxonomicName>
;
<figureCitation id="13102A4A9916FFA2CD01FDCAE5BE6E3E" box="[537,643,548,573]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="6.[151,250,1902,1924]" captionStartId-1="7.[151,250,1620,1642]" captionStartId-2="9.[151,250,1944,1966]" captionTargetBox-0="[188,1398,216,1878]" captionTargetBox-1="[491,1095,193,1598]" captionTargetBox-2="[262,1323,193,1883]" captionTargetId-0="figure@6.[188,1398,195,1878]" captionTargetId-1="figure@7.[491,1096,193,1598]" captionTargetId-2="figure@9.[262,1323,193,1917]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionText-0="FIGURE 1. Type specimens of Lutjanus octolineatus (A, B) and L. bengalensis (C): A. MNHN 0000 - A 5446 (no label attached with a blue glass plate), 122 mm SL, lectotype of Diacope octolineata, Mauritius, Mascarenes; B. MNHN 0000 - 7046 (holotype of Diacope octovittata), 137 mm SL, Mauritius, Mascarenes; C. ZMB 303, lectotype of Holocentrus bengalensis), 94 mm SL, Bengal, Bay of Bengal, eastern Indian Ocean." captionText-1="FIGURE 2. Squamation of subocular extension of cheek scales: A. Lutjanus octolineatus MNHN 0000 - 7046, holotype of Diacope octovittata, 137 mm SL; B. L. bengalensis, ZMB 303 (lectotype of Holocentrus bengalensis), 94 mm SL; C. L. sapphirolineatus n. sp., MUFS 33724, holotype, 153 mm SL." captionText-2="FIGURE 3. Photographs of fresh specimens of Lutjanus octolineatus (A), and L. bengalensis (B, C) A. uncatalogued specimen photographed by J. E. Randall, 108 mm SL, Mauritius; B. uncatalogued SAIAB specimen, 105 mm SL, South Africa; C. KAUM 54316, 96 mm SL, Japan." httpUri-0="https://zenodo.org/record/264483/files/figure.png" httpUri-1="https://zenodo.org/record/264484/files/figure.png" httpUri-2="https://zenodo.org/record/264485/files/figure.png" pageId="12" pageNumber="523">Figs. 13</figureCitation>
), discussed above under
<emphasis id="B95FEADD9916FFA2CC84FDCAE3346E3F" bold="true" box="[924,1033,548,573]" pageId="12" pageNumber="523">Remarks</emphasis>
for
<taxonomicName id="4C2B4D4C9916FFA2CB2FFDCBE3E36E3E" box="[1079,1246,548,572]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="octolineatus">
<emphasis id="B95FEADD9916FFA2CB2FFDCBE3E36E3E" box="[1079,1246,548,572]" italics="true" pageId="12" pageNumber="523">L. octolineatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9436CF9916FFA2CFDFFDA9E5B468EA" blockId="12.[151,1437,151,1544]" pageId="12" pageNumber="523">
<bibRefCitation id="EFBA4B3E9916FFA2CFDFFDA9E64D6E62" author="Bleeker" box="[199,368,583,608]" pageId="12" pageNumber="523" refString="Bleeker, P. (1855) Zesde bijdrage tot de kennis der ichthyologische fauna van Amboina. Natuurkundig Tijdschrift voor Nederlandsch Indie, 8, 391 - 434." type="journal article" year="1855">Bleeker (1855)</bibRefCitation>
described
<taxonomicName id="4C2B4D4C9916FFA2CEF4FDA6E43E6E62" box="[492,771,584,608]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="pomacanthus">
<emphasis id="B95FEADD9916FFA2CEF4FDA6E43E6E62" box="[492,771,584,608]" italics="true" pageId="12" pageNumber="523">Mesoprion pomacanthus</emphasis>
</taxonomicName>
on the basis of five specimens (
<quantity id="4CD39B2A9916FFA2CB6BFDA6E3D26E5D" box="[1139,1263,584,608]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" metricValueMax="4.8" metricValueMin="3.8" pageId="12" pageNumber="523" unit="mm" value="43.0" valueMax="48.0" valueMin="38.0">3848 mm</quantity>
in total length, Bleekers measurement, from snout tip to tip of caudal fin). However,
<bibRefCitation id="EFBA4B3E9916FFA2CCEDFD82E3A06E86" author="Bleeker" box="[1013,1181,620,645]" pageId="12" pageNumber="523" refString="Bleeker, P. (1874) Poissons de Madagascar et de l'ile de La Reunion. In: Bleeker, P. &amp; Pollen, F. P. L. (Eds.), Recherches sur la faune de Madagascar et de ses dependances, d'apres les decouvertes de Francois P. L. Pollen et D. C. van Dam. Part 4. Brill, Leiden, 104 + 2 unnumbered pp., 21 pls." type="journal article" year="1874">Bleeker (1874</bibRefCitation>
; Bleeker
<date id="FF95100F9916FFA2CA0DFD82E2A96E87" box="[1301,1428,620,645]" pageId="12" pageNumber="523" value="1876" valueMax="1877">18761877</date>
) subsequently recognized
<taxonomicName id="4C2B4D4C9916FFA2CEAFFD7FE54E6EAA" box="[439,627,656,680]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="pomacanthus">
<emphasis id="B95FEADD9916FFA2CEAFFD7FE54E6EAA" box="[439,627,656,680]" italics="true" pageId="12" pageNumber="523">M. pomacanthus</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C2B4D4C9916FFA2CC89FD7EE2196EAA" authority="Bloch 1790" authorityName="Bloch" authorityYear="1790" box="[913,1316,655,680]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CC89FD7EE3BD6EAA" box="[913,1152,656,680]" italics="true" pageId="12" pageNumber="523">Lutjanus bengalensis</emphasis>
(
<bibRefCitation id="EFBA4B3E9916FFA2CB8AFD61E2266EAA" author="Bloch" box="[1170,1307,655,680]" pageId="12" pageNumber="523" refString="Bloch, M. E. (1790) Naturgeschichte der auslandischen Fische. Vol. 4. Icthyologie, ou Histoire naturelle des Poissons, Berlin, xii + 128 pp., pls. 217 - 252." type="book chapter" year="1790">Bloch 1790</bibRefCitation>
)
</taxonomicName>
. Bleekers (1873) description of
<taxonomicName id="4C2B4D4C9916FFA2CE91FD5BE5166ECE" box="[393,555,692,716]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CE91FD5BE5166ECE" box="[393,555,692,716]" italics="true" pageId="12" pageNumber="523">L. bengalensis</emphasis>
</taxonomicName>
was based on
<specimenCount id="9D2DFD469916FFA2CDD3FD5AE45B6ECF" box="[715,870,692,717]" pageId="12" pageNumber="523" type="generic">20 specimens</specimenCount>
,
<quantity id="4CD39B2A9916FFA2CC68FD5AE4CA6ECE" box="[880,1015,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.54" metricValueMax="2.65" metricValueMin="0.43" pageId="12" pageNumber="523" unit="mm" value="154.0" valueMax="265.0" valueMin="43.0">43265 mm</quantity>
TL. The smallest of the five
<typeStatus id="5490886D9916FFA2CA22FD5BE2A16ECF" box="[1338,1436,693,717]" pageId="12" pageNumber="523" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C2B4D4C9916FFA2CFAFFD37E64E6EF2" box="[183,371,728,752]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="pomacanthus">
<emphasis id="B95FEADD9916FFA2CFAFFD37E64E6EF2" box="[183,371,728,752]" italics="true" pageId="12" pageNumber="523">M. pomacanthus</emphasis>
</taxonomicName>
(
<quantity id="4CD39B2A9916FFA2CE9CFD36E6EB6EED" box="[388,470,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="523" unit="mm" value="38.0">38 mm</quantity>
TL) was apparently already lost in 1873. According to M. van Oijen (pers. comm.), there are presently 28 of the
<specimenCount id="9D2DFD469916FFA2CEC5FD12E54B6F17" box="[477,630,764,789]" pageId="12" pageNumber="523" type="generic">29 specimens</specimenCount>
of
<taxonomicName id="4C2B4D4C9916FFA2CD85FD13E47D6F16" box="[669,832,764,788]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CD85FD13E47D6F16" box="[669,832,764,788]" italics="true" pageId="12" pageNumber="523">L. bengalensis</emphasis>
</taxonomicName>
listed/recorded in Bleeker (1873) in the collection of Naturalis Biodiversity Center (RMNH). The original Bleeker jar bought at the auction of the Bleeker collection (in
<date id="FF95100F9916FFA2CF8FFCAAE6736F5E" box="[151,334,836,861]" pageId="12" pageNumber="523" value="1879-12">December 1879</date>
) now contains
<specimenCount id="9D2DFD469916FFA2CD1BFCAAE59F6F5F" box="[515,674,836,861]" pageId="12" pageNumber="523" type="generic">24 specimens</specimenCount>
. At the time of the auction, it contained
<specimenCount id="9D2DFD469916FFA2CB9DFCAAE21D6F5F" box="[1157,1312,836,861]" pageId="12" pageNumber="523" type="generic">25 specimens</specimenCount>
(
<bibRefCitation id="EFBA4B3E9916FFA2CA2AFCAAE7EE6F82" author="Hubrecht" pageId="12" pageNumber="523" refString="Hubrecht, A. A. W. (1879) Catalogue des collections formees et laissees par M. - P. Bleeker. De Breuk &amp; Smits, Leiden, v + 71 pp." type="book" year="1879">Hubrecht 1879</bibRefCitation>
). Later, three specimens from the Series BE must have been added as
<specimenCount id="9D2DFD469916FFA2CB04FC86E38A6F82" box="[1052,1207,871,896]" pageId="12" pageNumber="523" type="generic">28 specimens</specimenCount>
are recorded in the RMNH register. Two specimens identified by G.R. Allen (WAM) as
<taxonomicName id="4C2B4D4C9916FFA2CCAEFC63E3D26FA7" authority="Forsskal 1775" authorityName="Forsskal" authorityYear="1775" box="[950,1263,908,933]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="kasmira">
<emphasis id="B95FEADD9916FFA2CCAEFC63E30F6FA6" box="[950,1074,908,932]" italics="true" pageId="12" pageNumber="523">L. kasmira</emphasis>
(
<bibRefCitation id="EFBA4B3E9916FFA2CB5BFC62E3DA6FA7" author="Forsskal" box="[1091,1255,908,933]" pageId="12" pageNumber="523" refString="Forsskal, P. S. (1775) Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit. Post mortem auctoris edidit Carsten Niebuhr. Molleri, Copenhagen, xxxiv + 164 pp." type="book" year="1775">Forsskål 1775</bibRefCitation>
)
</taxonomicName>
were removed from RMNH
<quantity id="4CD39B2A9916FFA2CE28FC41E6B86FCA" box="[304,389,943,968]" metricMagnitude="2" metricUnit="m" metricValue="1.402334" pageId="12" pageNumber="523" unit="in" value="5521.0">5521 in</quantity>
1978 and registered as RMNH
<quantity id="4CD39B2A9916FFA2CDF2FC41E4706FCA" box="[746,845,943,968]" metricMagnitude="2" metricUnit="m" metricValue="7.06755" pageId="12" pageNumber="523" unit="in" value="27825.0">27825 in</quantity>
this report. One of these two specimens is Bleekers largest specimen of
<taxonomicName id="4C2B4D4C9916FFA2CE62FC3BE5206FEE" box="[378,541,980,1004]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CE62FC3BE5206FEE" box="[378,541,980,1004]" italics="true" pageId="12" pageNumber="523">L. bengalensis</emphasis>
</taxonomicName>
from which Bleeker removed the pharyngeal jaws for the description (Bleeker 1873). Moreover, RMNH 5521 contained four
<typeStatus id="5490886D9916FFA2CDBDFC16E43A6812" box="[677,775,1016,1040]" pageId="12" pageNumber="523" type="syntype">syntypes</typeStatus>
of
<emphasis id="B95FEADD9916FFA2CC34FC17E4D26812" box="[812,1007,1016,1040]" italics="true" pageId="12" pageNumber="523">
<taxonomicName id="4C2B4D4C9916FFA2CC34FC17E4D56812" box="[812,1000,1016,1040]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="pomacanthus">M. pomacanthus</taxonomicName>
;
</emphasis>
two specimens 35 (42+) and 37 (44+) mm, identified by
<bibRefCitation id="EFBA4B3E9916FFA2CE60FBF2E5B86837" author="Allen" box="[376,645,1052,1077]" pageId="12" pageNumber="523" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot (1985)</bibRefCitation>
as
<taxonomicName id="4C2B4D4C9916FFA2CDAFFBF3E40B6836" box="[695,822,1052,1076]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="kasmira">
<emphasis id="B95FEADD9916FFA2CDAFFBF3E40B6836" box="[695,822,1052,1076]" italics="true" pageId="12" pageNumber="523">L. kasmira</emphasis>
</taxonomicName>
(now registered as RMNH PISC. 38326), and two specimens 30 (37+) and 36 (43+) mm, identified by
<bibRefCitation id="EFBA4B3E9916FFA2CDECFBD1E4C9685A" author="Allen" box="[756,1012,1087,1112]" pageId="12" pageNumber="523" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot (1985)</bibRefCitation>
as
<taxonomicName id="4C2B4D4C9916FFA2CB06FBAFE3FC685A" box="[1054,1217,1088,1112]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CB06FBAFE3FC685A" box="[1054,1217,1088,1112]" italics="true" pageId="12" pageNumber="523">L. bengalensis</emphasis>
</taxonomicName>
(now registered as RMNH PISC.38327 and RMNH PISC.38328, respectively).
<bibRefCitation id="EFBA4B3E9916FFA2CC41FB8AE35C687F" author="Allen" box="[857,1121,1124,1149]" pageId="12" pageNumber="523" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot (1985)</bibRefCitation>
identified these specimens solely on the basis of the number of dorsal-fin spines. In order to avoid further confusion, we here designate the specimen of
<quantity id="4CD39B2A9916FFA2CE3FFB42E64A68C6" box="[295,375,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="523" unit="mm" value="30.0">30 mm</quantity>
SL (37+ mm TL), RMNH PISC.38327 as the
<typeStatus id="5490886D9916FFA2CC9EFB42E4D068C7" box="[902,1005,1196,1221]" pageId="12" pageNumber="523" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C2B4D4C9916FFA2CB0CFB43E21768C6" box="[1044,1322,1196,1220]" family="Lutjanidae" genus="Mesoprion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="pomacanthus">
<emphasis id="B95FEADD9916FFA2CB0CFB43E21768C6" box="[1044,1322,1196,1220]" italics="true" pageId="12" pageNumber="523">Mesoprion pomacanthus</emphasis>
</taxonomicName>
; the three other
<typeStatus id="5490886D9916FFA2CFCFFB3FE60468EB" box="[215,313,1233,1257]" pageId="12" pageNumber="523" type="syntype">syntypes</typeStatus>
becoming the
<typeStatus id="5490886D9916FFA2CEC7FB21E5B968EA" box="[479,644,1231,1256]" pageId="12" pageNumber="523" type="paralectotype">paralectotypes</typeStatus>
.
</paragraph>
<paragraph id="8B9436CF9916FFA2CFDFFB1BE22B6A0A" blockId="12.[151,1437,151,1544]" pageId="12" pageNumber="523">
In addition,
<bibRefCitation id="EFBA4B3E9916FFA2CE56FB1AE6D5690E" author="Fricke" box="[334,488,1268,1293]" pageId="12" pageNumber="523" refString="Fricke, R. (1999) Fishes of the Mascarene Islands (Reunion, Mauritius, Rodriguez). An annotated checklist with descriptions of new species. Koeltz Scientific Books, Koenigstein, viii + 759 pp." type="book" year="1999">Fricke (1999)</bibRefCitation>
placed
<taxonomicName id="4C2B4D4C9916FFA2CD26FB1AE4F5690E" authority="Lacepede 1801" authorityName="Lacepede" authorityYear="1801" box="[574,968,1268,1293]" class="Actinopterygii" family="Labridae" genus="Labrus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="octovittatus">
<emphasis id="B95FEADD9916FFA2CD26FB1AE42A690E" box="[574,791,1268,1292]" italics="true" pageId="12" pageNumber="523">Labrus octovittatus</emphasis>
<bibRefCitation id="EFBA4B3E9916FFA2CC06FB1AE4F5690E" author="Lacepede" box="[798,968,1268,1293]" pageId="12" pageNumber="523" refString="Lacepede, B. G. E. (1801) Histoire naturelle des poissons (Lacepede). Histoire naturelle des poisons, 3, 1 - 558." type="journal article" year="1801">Lacepéde 1801</bibRefCitation>
</taxonomicName>
(p. 430, 477, pl., 22, fig. 2;
<typeStatus id="5490886D9916FFA2CA19FB1BE204690F" box="[1281,1337,1269,1293]" pageId="12" pageNumber="523">Type</typeStatus>
locality: Grand Ocean) in synonymy of
<taxonomicName id="4C2B4D4C9916FFA2CEE4FAF7E5A26932" box="[508,671,1304,1328]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B95FEADD9916FFA2CEE4FAF7E5A26932" box="[508,671,1304,1328]" italics="true" pageId="12" pageNumber="523">L. bengalensis</emphasis>
</taxonomicName>
although
<bibRefCitation id="EFBA4B3E9916FFA2CC0BFAF9E32E6932" author="Allen" box="[787,1043,1303,1328]" pageId="12" pageNumber="523" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot (1985)</bibRefCitation>
did not note this nominal species. The figure clearly shows presumably?X, probably
<quantity id="4CD39B2A9916FFA2CDCBFAD2E4366957" box="[723,779,1340,1365]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="12" pageNumber="523" unit="in" value="14.0">14 in</quantity>
dorsal-fin rays (as counted as IX, but the first short dorsalfin spine may have been missed or overlooked and not drawn) and presence of the third stripe, beginning from the middle of upper part of opercle. Such characters strongly suggest
<taxonomicName id="4C2B4D4C9916FFA2CCBAFA6BE3DD699F" authority="Forsskal 1775" authorityName="Forsskal" authorityYear="1775" box="[930,1248,1412,1437]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="kasmira">
<emphasis id="B95FEADD9916FFA2CCBAFA6BE322699E" box="[930,1055,1412,1436]" italics="true" pageId="12" pageNumber="523">L. kasmira</emphasis>
(
<bibRefCitation id="EFBA4B3E9916FFA2CB29FA6AE3E5699F" author="Forsskal" box="[1073,1240,1412,1437]" pageId="12" pageNumber="523" refString="Forsskal, P. S. (1775) Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit. Post mortem auctoris edidit Carsten Niebuhr. Molleri, Copenhagen, xxxiv + 164 pp." type="book" year="1775">Forsskål 1775</bibRefCitation>
)
</taxonomicName>
, (see diagnostic characters of
<bibRefCitation id="EFBA4B3E9916FFA2CE36FA49E52169C2" author="Allen" box="[302,540,1447,1472]" pageId="12" pageNumber="523" refString="Allen, G. R. &amp; Talbot, J. H. (1985) Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo-Pacific, with the description of a new species. Indo-Pacific Fish, 11, 1 - 87, pls. 1 - 10." type="journal article" year="1985">Allen &amp; Talbot 1985</bibRefCitation>
). The two characters could not be applied to
<taxonomicName id="4C2B4D4C9916FFA2CB0FFA47E38069C2" box="[1047,1213,1448,1472]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="octolineatus">
<emphasis id="B95FEADD9916FFA2CB0FFA47E38069C2" box="[1047,1213,1448,1472]" italics="true" pageId="12" pageNumber="523">L. octolineatus</emphasis>
</taxonomicName>
(with XII, 12 or 13 dorsal-fin rays) and
<taxonomicName id="4C2B4D4C9916FFA2CE98FA23E56669E6" box="[384,603,1484,1508]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="sapphirolineatus" status="sp. nov.">
<emphasis id="B95FEADD9916FFA2CE98FA23E56669E6" box="[384,603,1484,1508]" italics="true" pageId="12" pageNumber="523">L. sapphirolineatus</emphasis>
</taxonomicName>
<emphasis id="B95FEADD9916FFA2CD7DFA22E59B69E7" bold="true" box="[613,678,1484,1509]" pageId="12" pageNumber="523">
<taxonomicNameLabel id="A26C57A69916FFA2CD7DFA22E59B69E7" box="[613,678,1484,1509]" pageId="12" pageNumber="523" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(with the third stripe beginning just from upper rear edge of eye, through upper part of opercle, to upper part of caudal peduncle; see each diagnosis of the two species).
</paragraph>
</subSubSection>
</treatment>
</document>