312 lines
38 KiB
XML
312 lines
38 KiB
XML
<document id="D7DB8701CC7F8CAB9455058ED6EB24A3" ID-CLB-Dataset="5429" ID-DOI="10.11646/zootaxa.4981.3.3" ID-GBIF-Dataset="bd4313a1-f17a-4745-ba2d-655124108edc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921137" ID-ZooBank="CB36C384-5538-494A-8802-4F821C7E25FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623280791010" checkinUser="plazi" docAuthor="Chernyshev, Alexei V. & Goedert, James L." docDate="2021" docId="038287AEC141174EFF7F5E610B3F0EC6" docLanguage="en" docName="zootaxa.4981.3.3.pdf" docOrigin="Zootaxa 4981 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orbitestella kieli Chernyshev & Goedert 2021, sp. nov." docType="treatment" docUuid="B792FEA1-5E06-4365-AC4C-639558F1DFB5" docUuidSource="ZooBank" docVersion="8" lastPageNumber="474" masterDocId="FFBBFFD6C142174BFFE85A1D090F0F6B" masterDocTitle="New species of latest Eocene / earliest Oligocene microgastropods (Heterobranchia Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA" masterLastPageNumber="480" masterPageNumber="469" pageNumber="472" updateTime="1699009098020" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="1747AE171A0B49B6F5361A9860CCD39C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DF8767A3590C90FD757EEBD9C6208E29">
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<mods:title id="2977CBE3E093CF8FA6ABBAFF25B5BC3A">New species of latest Eocene / earliest Oligocene microgastropods (Heterobranchia Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA</mods:title>
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</mods:titleInfo>
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<mods:name id="59626698093702995EFDE93E14AD74E2" type="personal">
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<mods:role id="B96B6CD578EEC5D339DD975C7C96D8C3">
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<mods:roleTerm id="A04F5782AAF629430BF4A8F9246592EB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="22EE59BBFDD1EA039967444996AB9581">Chernyshev, Alexei V.</mods:namePart>
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<mods:affiliation id="B820C4B722D7EA8BA458FA162025E5CC">A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch, Russian Academy of Sciences, Palchevskogo Street 17, Vladivostok 690041, Russia.</mods:affiliation>
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<mods:name id="DD9F2153E4440765451A9C390F334FB0" type="personal">
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<mods:roleTerm id="BF8C719B7DA6662B3E2B2B32CD9ABF02">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="38E8D1FE9ABD2A320BECAC8FF07E5D51">Goedert, James L.</mods:namePart>
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<mods:affiliation id="640593365B1DA36BA6DC3AE60A37102E">Burke Museum of Natural History and Culture, University of Washington, Seattle, WA 98195, USA.</mods:affiliation>
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<mods:typeOfResource id="DDAF9AB724823946D195002C2EA5DE61">text</mods:typeOfResource>
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<mods:title id="8A920DBC0B2ECB23038BAB1C2A93525F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="329B9D86042F52B4E75241DEC8885EB2">
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<mods:date id="99D445FB93E21E70728ED9EBE675F701">2021</mods:date>
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<mods:detail id="457696EE1ECA7392A5F2A304CE18AF51" type="pubDate">
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<mods:number id="211484050CEBBF213EB731031C5900EA">2021-06-08</mods:number>
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<mods:detail id="16359B5A3E5A6BEDAC5F23CF3C3EEE93" type="volume">
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<mods:number id="8D612208157682B8F139D7DEA8AE9D9C">4981</mods:number>
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</mods:detail>
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<mods:detail id="740C4C754663B502F7D5AF23443CBB88" type="issue">
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<mods:number id="8A41FE5234A1335D42C3DC8A036F0C32">3</mods:number>
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<mods:start id="31EBC1BF1231812C2F573E7BA2F38B0F">469</mods:start>
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<mods:end id="5ADCF3E6D7DA0635F0A6206B3CFBE0B9">480</mods:end>
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<mods:classification id="09BFD044788F2447E58DA1CFE94C3E84">journal article</mods:classification>
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<mods:identifier id="DB2840CD573F11AAA30DD13CD6E9019E" type="CLB-Dataset">5429</mods:identifier>
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<mods:identifier id="3FE4B49D1D25E8AE56518BE969395982" type="DOI">10.11646/zootaxa.4981.3.3</mods:identifier>
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<mods:identifier id="BFBAC3238AF8CA74C54FA3D4EA2D752B" type="GBIF-Dataset">bd4313a1-f17a-4745-ba2d-655124108edc</mods:identifier>
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<mods:identifier id="DF2D18236C029CD86A154A7E8FB1114D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5457E4B19F41AE72CB603EB3B02A11D8" type="Zenodo-Dep">4921137</mods:identifier>
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<mods:identifier id="EE1D97C4D3E01C13206DE5F3FEFB2532" type="ZooBank">CB36C384-5538-494A-8802-4F821C7E25FE</mods:identifier>
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</mods:mods>
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<treatment id="038287AEC141174EFF7F5E610B3F0EC6" ID-DOI="http://doi.org/10.5281/zenodo.5109763" ID-GBIF-Taxon="183548938" ID-Zenodo-Dep="5109763" LSID="urn:lsid:zoobank.org:act:B792FEA1-5E06-4365-AC4C-639558F1DFB5" httpUri="http://treatment.plazi.org/id/038287AEC141174EFF7F5E610B3F0EC6" lastPageId="5" lastPageNumber="474" pageId="3" pageNumber="472">
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<subSubSection id="C3316533C1411748FF7F5E6108CB0BFC" box="[151,452,1148,1175]" pageId="3" pageNumber="472" type="nomenclature">
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<paragraph id="8B9436B8C1411748FF7F5E6108CB0BFC" blockId="3.[151,452,1148,1211]" box="[151,452,1148,1175]" pageId="3" pageNumber="472">
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<heading id="D0DC81D4C1411748FF7F5E6108CB0BFC" bold="true" box="[151,452,1148,1175]" fontSize="11" level="1" pageId="3" pageNumber="472" reason="1">
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||
<emphasis id="B95FEAAAC1411748FF7F5E6108CB0BFC" bold="true" box="[151,452,1148,1175]" pageId="3" pageNumber="472">
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||
<taxonomicName id="4C2B4D3BC1411748FF7F5E6108500BFC" authority="Chernyshev & Goedert, 2021" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[151,351,1148,1175]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="3" pageNumber="472" phylum="Mollusca" rank="species" species="kieli" status="sp. nov.">
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<emphasis id="B95FEAAAC1411748FF7F5E6108500BFC" bold="true" box="[151,351,1148,1175]" italics="true" pageId="3" pageNumber="472">Orbitestella kieli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C57D1C1411748FE8E5E6008CB0BFC" box="[358,452,1149,1175]" pageId="3" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3316533C1411748FF7F5EBC0D290B96" pageId="3" pageNumber="472" type="description">
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||
<paragraph id="8B9436B8C1411748FF7F5EBC08280BD0" blockId="3.[151,452,1148,1211]" box="[151,295,1185,1211]" pageId="3" pageNumber="472">
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||
<figureCitation id="13102A3DC1411748FF7F5EBC08280BD0" box="[151,295,1185,1211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1932,1956]" captionTargetBox="[151,1435,597,1909]" captionTargetId="figure-245@4.[151,1435,597,1909]" captionTargetPageId="4" captionText="FIGURE 2. Orbitestella kieli sp. nov. (SEM). A, B, F – holotype MIMB 41134, C, D, E—paratypes MIMB 41135–1 (C, E) and MIMB 41135–2 (D); A, apical view; B, apertural view; C, umbilical view; D, protoconch, apical view; E, protoconch, umbilical view; F, periphery, wrinkled sculpture formed by spiral threads, apical view. Scales: A-C, 100 µm; D-F, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4921143" httpUri="https://zenodo.org/record/4921143/files/figure.png" pageId="3" pageNumber="472">Figure 2A–F</figureCitation>
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</paragraph>
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<paragraph id="8B9436B8C1411748FF7F5EF80D290B96" blockId="3.[151,1062,1253,1277]" box="[151,1062,1253,1277]" pageId="3" pageNumber="472">
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<emphasis id="B95FEAAAC1411748FF7F5EF8084F0B96" bold="true" box="[151,320,1253,1277]" pageId="3" pageNumber="472">Zoobank LSID.</emphasis>
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||
<uri id="FFBA3ABAC1411748FEAE5EF80D290B96" box="[326,1062,1253,1277]" pageId="3" pageNumber="472">
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||
urn:lsid:zoobank.org:act:
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<uuid id="FF8D0C6DC1411748FDA85EF80D290B96" box="[576,1062,1253,1277]" pageId="3" pageNumber="472">B792FEA1-5E06-4365-AC4C-639558F1DFB5</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3316533C1411748FF7F5F360C570AD9" pageId="3" pageNumber="472" type="materials_examined">
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||
<paragraph id="8B9436B8C1411748FF7F5F360C6F0A02" blockId="3.[151,1437,1323,2034]" pageId="3" pageNumber="472">
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||
<emphasis id="B95FEAAAC1411748FF7F5F3608090A2E" bold="true" box="[151,262,1323,1349]" pageId="3" pageNumber="472">Material.</emphasis>
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||
<materialsCitation id="3B433CE5C1411748FEE65F310B250A2D" ID-GBIF-Occurrence="3323273311" box="[270,554,1324,1350]" collectionCode="MIMB" pageId="3" pageNumber="472" specimenCode="MIMB 41134" specimenCount="1" typeStatus="holotype">
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<emphasis id="B95FEAAAC1411748FEE65F31088B0A2D" bold="true" box="[270,388,1324,1350]" pageId="3" pageNumber="472">
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<typeStatus id="5490881AC1411748FEE65F3108700A2D" box="[270,383,1324,1350]" pageId="3" pageNumber="472" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCode id="DB8D9EC3C1411748FE645F310B250A2D" box="[396,554,1324,1350]" collectionCode="MIMB" country="Russia" httpUri="http://grbio.org/cool/063z-3mpp" name="Museum of the Institute of Marine Biology" pageId="3" pageNumber="472">MIMB 41134</specimenCode>
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||
</materialsCitation>
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||
;
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<materialsCitation id="3B433CE5C1411748FDDD5F310D020A2D" ID-GBIF-Occurrence="3323273313" box="[565,1037,1323,1350]" collectionCode="MIMB" pageId="3" pageNumber="472" specimenCode="MIMB 41135" specimenCount="11" typeStatus="paratype">
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||
<emphasis id="B95FEAAAC1411748FDDD5F310BB80A2D" bold="true" box="[565,695,1324,1350]" pageId="3" pageNumber="472">
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||
<typeStatus id="5490881AC1411748FDDD5F310BBE0A2D" box="[565,689,1324,1350]" pageId="3" pageNumber="472" type="paratype">Paratypes</typeStatus>
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||
:
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</emphasis>
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||
<specimenCode id="DB8D9EC3C1411748FD565F310A560A2D" box="[702,857,1323,1350]" collectionCode="MIMB" country="Russia" httpUri="http://grbio.org/cool/063z-3mpp" name="Museum of the Institute of Marine Biology" pageId="3" pageNumber="472">MIMB 41135</specimenCode>
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||
(
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<specimenCount id="9D2DFD31C1411748FC815F310D0A0A2D" box="[873,1029,1323,1350]" pageId="3" pageNumber="472" type="generic">11 specimens</specimenCount>
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||
)
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||
</materialsCitation>
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||
, NRM PAL Mo 193050–193053 (
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<specimenCount id="9D2DFD31C1411748FA655F31081F0A01" pageId="3" pageNumber="472" type="generic">4 specimens</specimenCount>
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||
),
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||
<materialsCitation id="3B433CE5C1411748FECB5F4D0B720A01" ID-GBIF-Occurrence="3323273319" box="[291,637,1359,1386]" collectionCode="SBMNH" pageId="3" pageNumber="472" specimenCode="SBMNH 467093" specimenCount="1" typeStatus="paratype">
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||
<specimenCode id="DB8D9EC3C1411748FECB5F4D08ED0A02" box="[291,482,1359,1386]" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="3" pageNumber="472" type="Museum">SBMNH 467093</specimenCode>
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||
(
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||
<specimenCount id="9D2DFD31C1411748FE1C5F4D0B7A0A01" box="[500,629,1359,1386]" pageId="3" pageNumber="472" type="generic">1 specimen</specimenCount>
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||
)
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</materialsCitation>
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||
,
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<materialsCitation id="3B433CE5C1411748FD605F4D0D6C0A01" ID-GBIF-Occurrence="3323273323" accessionNumber="IP 114130" box="[648,1123,1359,1386]" collectionCode="UWBM" pageId="3" pageNumber="472" specimenCode="IP 114130-114135" specimenCount="6" typeStatus="paratype">
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<collectionCode id="ED3AAE7DC1411748FD605F4D0BED0A01" box="[648,738,1360,1386]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="3" pageNumber="472" type="Museum">UWBM</collectionCode>
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<specimenCode id="DB8D9EC3C1411748FD015F4D0AB30A01" box="[745,956,1359,1386]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="3" pageNumber="472" type="Museum">IP 114130–114135</specimenCode>
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||
(
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<specimenCount id="9D2DFD31C1411748FC245F520D540A01" box="[972,1115,1359,1386]" pageId="3" pageNumber="472" type="generic">6 specimens</specimenCount>
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||
)
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||
</materialsCitation>
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||
; all from type locality.
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</paragraph>
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<paragraph id="8B9436B8C1411748FF2F5F690C570AD9" blockId="3.[151,1437,1323,2034]" pageId="3" pageNumber="472">
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<materialsCitation id="3B433CE5C1411748FF2F5F690C5B0AD9" ID-GBIF-Occurrence="3323273314" collectionCode="LACMIP" country="United States of America" county="Lewis County" location="Gries Ranch Formation" municipality="Ma" pageId="3" pageNumber="472" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
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<emphasis id="B95FEAAAC1411748FF2F5F69086A0AE6" bold="true" box="[199,357,1395,1422]" pageId="3" pageNumber="472">
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||
<typeStatus id="5490881AC1411748FF2F5F69080D0AE5" box="[199,258,1396,1422]" pageId="3" pageNumber="472">Type</typeStatus>
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locality.
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||
</emphasis>
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||
<collectionCode id="ED3AAE7DC1411748FE845F6908DA0AE5" box="[364,469,1396,1422]" pageId="3" pageNumber="472">LACMIP</collectionCode>
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locality 41621;
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||
<location id="8EF46063C1411748FD635F6E0A9A0AE6" LSID="urn:lsid:plazi:treatment:038287AEC141174EFF7F5E610B3F0EC6:8EF46063C1411748FD635F6E0A9A0AE6" box="[651,917,1395,1421]" country="United States of America" county="Lewis County" municipality="Ma" name="Gries Ranch Formation" pageId="3" pageNumber="472" stateProvince="Washington">Gries Ranch Formation</location>
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,
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<collectingCounty id="62F54E34C1411748FC775F6E0D300AE5" box="[927,1087,1395,1422]" pageId="3" pageNumber="472">Lewis County</collectingCounty>
|
||
,
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||
<collectingRegion id="49EFF85AC1411748FBA15F6E0C1F0AE5" box="[1097,1296,1395,1422]" country="United States of America" name="Washington" pageId="3" pageNumber="472">Washington State</collectingRegion>
|
||
,
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<collectingCountry id="F33C7628C1411748FAF25F690C590AE5" box="[1306,1366,1396,1422]" name="United States of America" pageId="3" pageNumber="472">USA</collectingCountry>
|
||
; latest
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<location id="8EF46063C1411748FF7F5F8509E60AD9" LSID="urn:lsid:plazi:treatment:038287AEC141174EFF7F5E610B3F0EC6:8EF46063C1411748FF7F5F8509E60AD9" box="[151,233,1432,1458]" country="United States of America" county="Lewis County" municipality="Ma" name="Eocene" pageId="3" pageNumber="472" stateProvince="Washington">Eocene</location>
|
||
or earliest
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||
<location id="8EF46063C1411748FE8F5F8A08D20ADA" LSID="urn:lsid:plazi:treatment:038287AEC141174EFF7F5E610B3F0EC6:8EF46063C1411748FE8F5F8A08D20ADA" box="[359,477,1431,1457]" country="United States of America" county="Lewis County" municipality="Ma" name="Oligocene" pageId="3" pageNumber="472" stateProvince="Washington">Oligocene</location>
|
||
, 33.5–34.7
|
||
<collectingMunicipality id="6BF0ACC2C1411748FDB75F850B8B0AD9" box="[607,644,1432,1458]" pageId="3" pageNumber="472">Ma</collectingMunicipality>
|
||
(latest Priabonian or earliest Rupelian;
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||
<bibRefCitation id="EFBA4B49C1411748FBA95F8A0C440AD9" author="Prothero, D. R. & Burns, C." box="[1089,1355,1431,1458]" pageId="3" pageNumber="472" pagination="179 - 183" refId="ref6601" refString="Prothero, D. R. & Burns, C. (2001) Paleomagnetism of the upper Eocene Gries Ranch Formation, Grays Harbor [sic] County, Washington. In: Prothero, D. R. (Ed.), Magnetic Stratigraphy of the Pacific Coast Cenozoic. Society for Sedimentary Geology (SEPM) Book 91. Society for Sedimentary Geology, Tulsa, Oklahoma, pp. 179 - 183." type="book chapter" year="2001">Prothero & Burns 2001</bibRefCitation>
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||
)
|
||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316533C1411748FF2F5FA109C80A91" pageId="3" pageNumber="472" type="etymology">
|
||
<paragraph id="8B9436B8C1411748FF2F5FA109C80A91" blockId="3.[151,1437,1323,2034]" pageId="3" pageNumber="472">
|
||
<emphasis id="B95FEAAAC1411748FF2F5FA108410ABD" bold="true" box="[199,334,1468,1494]" pageId="3" pageNumber="472">Etymology.</emphasis>
|
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Named for Steffen Kiel (NRM) in recognition of his many contributions to molluscan paleontology.
|
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</paragraph>
|
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</subSubSection>
|
||
<subSubSection id="C3316533C1411748FF2F5C1E0BF6090E" pageId="3" pageNumber="472" type="diagnosis">
|
||
<paragraph id="8B9436B8C1411748FF2F5C1E0BF6090E" blockId="3.[151,1437,1323,2034]" pageId="3" pageNumber="472">
|
||
<emphasis id="B95FEAAAC1411748FF2F5C1E084D0976" bold="true" box="[199,322,1539,1565]" pageId="3" pageNumber="472">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C2B4D3BC1411748FEA15C1E08C00976" authorityName="Iredale" authorityYear="1917" box="[329,463,1539,1565]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="3" pageNumber="472" phylum="Mollusca" rank="genus">
|
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<emphasis id="B95FEAAAC1411748FEA15C1E08C00976" box="[329,463,1539,1565]" italics="true" pageId="3" pageNumber="472">Orbitestella</emphasis>
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</taxonomicName>
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with flattened spire and sunken apex, nucleus of protoconch with net-like ornamentation, faint axial riblets between low axial ribs on apical surface, broad and smooth upper carina, faint spiral threads more pronounced on apical surface near the periphery.
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C3316533C141174EFF2F5C72086E0E0E" lastPageId="5" lastPageNumber="474" pageId="3" pageNumber="472" type="description">
|
||
<paragraph id="8B9436B8C1411748FF2F5C720A8308EE" blockId="3.[151,1437,1323,2034]" pageId="3" pageNumber="472">
|
||
<emphasis id="B95FEAAAC1411748FF2F5C72085609E2" bold="true" box="[199,345,1647,1673]" pageId="3" pageNumber="472">Description.</emphasis>
|
||
Shell small for genus, depressed, with a flat spire; maximum shell diameter
|
||
<quantity id="4CD39B5DC1411748FB205C720C7F09E1" box="[1224,1392,1647,1674]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" metricValueMax="9.3" metricValueMin="6.3" pageId="3" pageNumber="472" unit="mm" value="0.78" valueMax="0.93" valueMin="0.63">0.63–0.93 mm</quantity>
|
||
; dimensions of the
|
||
<typeStatus id="5490881AC1411748FEB95C8E08BC09C6" box="[337,435,1683,1709]" pageId="3" pageNumber="472" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CD39B5DC1411748FE535C890B1009C5" box="[443,543,1684,1710]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="472" unit="mm" value="0.33">0.33 mm</quantity>
|
||
height ×
|
||
<quantity id="4CD39B5DC1411748FD635C890BE009C5" box="[651,751,1684,1710]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="3" pageNumber="472" unit="mm" value="0.93">0.93 mm</quantity>
|
||
diameter. Protoconch approximately 1.5 whorls,
|
||
<quantity id="4CD39B5DC1411748FAF35C890C7009C5" box="[1307,1407,1684,1710]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="472" unit="mm" value="0.08">0.08 mm</quantity>
|
||
in maximal diameter, nucleus with net-like ornamentation, visible in both apical and basal view. Teleoconch approximately 1.5–1.8 whorls, with radial ribs prominent only on the inner portion of each whorl, becoming obsolete on periphery so that the distinct cord on the basal, outer part of each whorl is smooth, lacking nodes; subapical margin of each whorl slightly wider that the subbasal margin, also delineated by a strong cord without nodes, radial ribs present but not prominent. Umbilicus broad and wide, second whorl of teleoconch partly covers first whorl. Apical surface of whorls smooth low arch to shoulder. Aperture polygonal.
|
||
</paragraph>
|
||
<paragraph id="8B9436B8C141174FFF2F5D920D100EBA" blockId="3.[151,1437,1323,2034]" lastBlockId="4.[151,1437,151,573]" lastPageId="4" lastPageNumber="473" pageId="3" pageNumber="472">
|
||
<emphasis id="B95FEAAAC1411748FF2F5D92086208C2" bold="true" box="[199,365,1935,1961]" pageId="3" pageNumber="472">Comparisons.</emphasis>
|
||
<taxonomicName id="4C2B4D3BC1411748FE9F5D920B3808C2" authority="Chernyshev & Goedert, 2021" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[375,567,1935,1961]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="3" pageNumber="472" phylum="Mollusca" rank="species" species="kieli" status="sp. nov.">
|
||
<emphasis id="B95FEAAAC1411748FE9F5D920B3808C2" box="[375,567,1935,1961]" italics="true" pageId="3" pageNumber="472">Orbitestella kieli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C57D1C1411748FDA95D8D0B9408C1" box="[577,667,1936,1962]" pageId="3" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C2B4D3BC1411748FCD05D8D0AF008C2" authorityName="Squires & Goedert" authorityYear="1996" box="[824,1023,1935,1962]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="3" pageNumber="472" phylum="Mollusca" rank="species" species="palaiopacifica">
|
||
<emphasis id="B95FEAAAC1411748FCD05D8D0AF008C2" box="[824,1023,1935,1962]" italics="true" pageId="3" pageNumber="472">O. palaiopacifica</emphasis>
|
||
</taxonomicName>
|
||
with the nucleus of the protoconch clearly visible in both the basal and, to a lesser extent, apical view.
|
||
<taxonomicName id="4C2B4D3BC1411748FC9F5DAE0D3D08A6" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[887,1074,1971,1997]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="3" pageNumber="472" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC1411748FC9F5DAE0D3D08A6" box="[887,1074,1971,1997]" italics="true" pageId="3" pageNumber="472">Orbitestella kieli</emphasis>
|
||
</taxonomicName>
|
||
differs in lacking a distinct keellike border to the apical periphery of the whorls and lacks nodes on the basal periphery of the whorls.
|
||
<taxonomicName id="4C2B4D3BC141174FFAFE5DCA09C80FDA" authorityName="Chernyshev & Goedert" authorityYear="2021" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" lastPageId="4" lastPageNumber="473" order="Heterostropha" pageId="3" pageNumber="472" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC141174FFAFE5DCA09C80FDA" italics="true" lastPageId="4" lastPageNumber="473" pageId="3" pageNumber="472">Orbitestella kieli</emphasis>
|
||
</taxonomicName>
|
||
also has only fine spiral lirae, except near the periphery in apical view where faint spiral threads produce a wrinkled texture on the last whorl of mature specimens (
|
||
<figureCitation id="13102A3DC146174FFCF45AA60AAE0FBE" box="[796,929,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1932,1956]" captionTargetBox="[151,1435,597,1909]" captionTargetId="figure-245@4.[151,1435,597,1909]" captionTargetPageId="4" captionText="FIGURE 2. Orbitestella kieli sp. nov. (SEM). A, B, F – holotype MIMB 41134, C, D, E—paratypes MIMB 41135–1 (C, E) and MIMB 41135–2 (D); A, apical view; B, apertural view; C, umbilical view; D, protoconch, apical view; E, protoconch, umbilical view; F, periphery, wrinkled sculpture formed by spiral threads, apical view. Scales: A-C, 100 µm; D-F, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4921143" httpUri="https://zenodo.org/record/4921143/files/figure.png" pageId="4" pageNumber="473">Figs. 2A, F</figureCitation>
|
||
). More pronounced spiral threads are visible on the shell of
|
||
<taxonomicName id="4C2B4D3BC146174FFEA15AC20B1F0F92" authorityName="Squires & Goedert" authorityYear="1996" box="[329,528,223,249]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="palaiopacifica">
|
||
<emphasis id="B95FEAAAC146174FFEA15AC20B1F0F92" box="[329,528,223,249]" italics="true" pageId="4" pageNumber="473">O. palaiopacifica</emphasis>
|
||
</taxonomicName>
|
||
, including on the keel-like upper carina as well as the basal carina (e.g.,
|
||
<figureCitation id="13102A3DC146174FFA8D5AC209D40E76" captionStart="FIGURE 1" captionStartId="3.[151,250,1017,1041]" captionTargetBox="[151,1435,179,991]" captionTargetId="figure-19@3.[151,1435,179,991]" captionTargetPageId="3" captionText="FIGURE 1. Orbitestella palaiopacifica Squires & Goedert (SEM). A, specimen MIMB 41133–1, apical view; B, specimen MIMB 41133–2, umbilical view; C, specimen MIMB 41133–3, apertural view. Scales: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4921141" httpUri="https://zenodo.org/record/4921141/files/figure.png" pageId="4" pageNumber="473">Figs. 1B–C</figureCitation>
|
||
), and
|
||
<taxonomicName id="4C2B4D3BC146174FFEC95B1E08E90E76" authorityName="Squires & Goedert" authorityYear="1996" box="[289,486,259,285]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="palaiopacifica">
|
||
<emphasis id="B95FEAAAC146174FFEC95B1E08E90E76" box="[289,486,259,285]" italics="true" pageId="4" pageNumber="473">O. palaiopacifica</emphasis>
|
||
</taxonomicName>
|
||
has a more domed or humped apical surface of the whorls than
|
||
<taxonomicName id="4C2B4D3BC146174FFB575B1E0C1D0E76" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[1215,1298,259,285]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC146174FFB575B1E0C1D0E76" box="[1215,1298,259,285]" italics="true" pageId="4" pageNumber="473">O. kieli</emphasis>
|
||
</taxonomicName>
|
||
. Radial ribs are prominent on both the apical and basal surface of the shell of
|
||
<taxonomicName id="4C2B4D3BC146174FFC6F5B3A0D420E2A" authorityName="Squires & Goedert" authorityYear="1996" box="[903,1101,295,321]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="palaiopacifica">
|
||
<emphasis id="B95FEAAAC146174FFC6F5B3A0D420E2A" box="[903,1101,295,321]" italics="true" pageId="4" pageNumber="473">O. palaiopacifica</emphasis>
|
||
</taxonomicName>
|
||
and in most specimens, even extend to and are present on the apical carina, whereas in
|
||
<taxonomicName id="4C2B4D3BC146174FFCC95B560A7C0E0E" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[801,883,331,357]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC146174FFCC95B560A7C0E0E" box="[801,883,331,357]" italics="true" pageId="4" pageNumber="473">O. kieli</emphasis>
|
||
</taxonomicName>
|
||
they fade toward the periphery, especially on the basal surface, to smooth completely as they reach the periphery, and those of
|
||
<taxonomicName id="4C2B4D3BC146174FFC145B720D410EE2" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[1020,1102,367,393]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC146174FFC145B720D410EE2" box="[1020,1102,367,393]" italics="true" pageId="4" pageNumber="473">O. kieli</emphasis>
|
||
</taxonomicName>
|
||
are not prominent on the apical surface, although they are clearly present. Compared with
|
||
<taxonomicName id="4C2B4D3BC146174FFCBF5B8E0D120EC6" authorityName="Squires & Goedert" authorityYear="1996" box="[855,1053,403,429]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="palaiopacifica">
|
||
<emphasis id="B95FEAAAC146174FFCBF5B8E0A640EC6" box="[855,875,403,429]" italics="true" pageId="4" pageNumber="473">O</emphasis>
|
||
.
|
||
<emphasis id="B95FEAAAC146174FFC935B8E0D120EC6" box="[891,1053,403,429]" italics="true" pageId="4" pageNumber="473">palaiopacifica</emphasis>
|
||
</taxonomicName>
|
||
, the flattened basal surface of the whorls is slightly wider in
|
||
<taxonomicName id="4C2B4D3BC146174FFE2B5BAA0B1A0EBA" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[451,533,439,465]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC146174FFE2B5BAA0B1A0EBA" box="[451,533,439,465]" italics="true" pageId="4" pageNumber="473">O. kieli</emphasis>
|
||
</taxonomicName>
|
||
, with a distinct cord marking the outer border.
|
||
</paragraph>
|
||
<paragraph id="8B9436B8C146174FFF2F5BC60A5E0D56" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="473">
|
||
The only other fossil orbitestellid from western North America,
|
||
<taxonomicName id="4C2B4D3BC146174FFC4E5BC60C1A0E9E" authority="Hybertsen & Kiel, 2018" authorityName="Hybertsen & Kiel" authorityYear="2018" box="[934,1301,475,501]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="dioi">
|
||
<emphasis id="B95FEAAAC146174FFC4E5BC60AF90E9E" box="[934,1014,475,501]" italics="true" pageId="4" pageNumber="473">O. dioi</emphasis>
|
||
<bibRefCitation id="EFBA4B49C146174FFC175BC60C1A0E9E" author="Hybertsen, F. & Kiel, S." box="[1023,1301,475,501]" pageId="4" pageNumber="473" pagination="751 - 768" refId="ref6146" refString="Hybertsen, F. & Kiel, S. (2018) A middle Eocene seep deposit with silicified fauna from the Humptulips Formation in western Washington State, USA. Acta Palaeontologica Polonica, 63, 751 - 768. https: // doi. org / 10.4202 / app. 00525.2018" type="journal article" year="2018">Hybertsen & Kiel, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
, from a hydrocarbon seep deposit in the Eocene deep water Humptulips Formation, differs in having more prominent spiral ornamentation, a rounded whorl profile, and an elevated spire.
|
||
</paragraph>
|
||
<caption id="DF546630C146174FFF7F5D910DF10886" ID-DOI="http://doi.org/10.5281/zenodo.4921143" ID-Zenodo-Dep="4921143" httpUri="https://zenodo.org/record/4921143/files/figure.png" pageId="4" pageNumber="473" startId="4.[151,250,1932,1956]" targetBox="[151,1435,597,1909]" targetPageId="4">
|
||
<paragraph id="8B9436B8C146174FFF7F5D910DF10886" blockId="4.[151,1437,1932,2029]" pageId="4" pageNumber="473">
|
||
<emphasis id="B95FEAAAC146174FFF7F5D91081D08CF" bold="true" box="[151,274,1932,1956]" pageId="4" pageNumber="473">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C2B4D3BC146174FFEF05D9108CC08CF" authority="Chernyshev & Goedert, 2021" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[280,451,1932,1956]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="473" phylum="Mollusca" rank="species" species="kieli" status="sp. nov.">
|
||
<emphasis id="B95FEAAAC146174FFEF05D9108CC08CF" box="[280,451,1932,1956]" italics="true" pageId="4" pageNumber="473">Orbitestella kieli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C57D1C146174FFE215D900B1508CF" box="[457,538,1933,1957]" pageId="4" pageNumber="473" rank="species">
|
||
<emphasis id="B95FEAAAC146174FFE215D900B1C08CE" bold="true" box="[457,531,1933,1957]" pageId="4" pageNumber="473">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(SEM). A, B, F – holotype MIMB 41134, C, D, E—paratypes MIMB 41135–1 (C, E) and MIMB 41135–2 (D); A, apical view; B, apertural view; C, umbilical view; D, protoconch, apical view; E, protoconch, umbilical view; F, periphery, wrinkled sculpture formed by spiral threads, apical view. Scales: A-C, 100 µm; D-F, 20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9436B8C147174EFF2F5A8A086E0E0E" blockId="5.[151,1437,151,429]" pageId="5" pageNumber="474">
|
||
A single living species,
|
||
<taxonomicName id="4C2B4D3BC147174EFE305A8A0A100FDA" authority="(Bartsch, 1907)" baseAuthorityName="Bartsch" baseAuthorityYear="1907" box="[472,799,151,177]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="diegensis">
|
||
<emphasis id="B95FEAAAC147174EFE305A8A0B6B0FDA" box="[472,612,151,177]" italics="true" pageId="5" pageNumber="474">O. diegensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBA4B49C147174EFD9C5A8A0A180FDA" author="Bartsch, P." box="[628,791,151,177]" pageId="5" pageNumber="474" pagination="167 - 176" refId="ref5143" refString="Bartsch, P. (1907) New mollusks of the family Vitrinellidae from the West Coast of America. Proceedings of the United States National Museum, 32, 167 - 176. https: // doi. org / 10.5479 / si. 00963801.32 - 1520.167" type="journal article" year="1907">Bartsch, 1907</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, inhabits the Pacific
|
||
<collectingRegion id="49EFF85AC147174EFBE65A8A0D410FDA" box="[1038,1102,151,177]" country="New Zealand" name="West Coast" pageId="5" pageNumber="474">Coast</collectingRegion>
|
||
of North America southward from central
|
||
<collectingRegion id="49EFF85AC147174EFECF5AA608930FBE" box="[295,412,187,213]" country="United States of America" name="California" pageId="5" pageNumber="474">California</collectingRegion>
|
||
.
|
||
<taxonomicName id="4C2B4D3BC147174EFE4F5AA60B940FBE" baseAuthorityName="Bartsch" baseAuthorityYear="1907" box="[423,667,187,213]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="diegensis">
|
||
<emphasis id="B95FEAAAC147174EFE4F5AA60B940FBE" box="[423,667,187,213]" italics="true" pageId="5" pageNumber="474">Orbitestella diegensis</emphasis>
|
||
</taxonomicName>
|
||
differs greatly from all the fossil species from
|
||
<collectingRegion id="49EFF85AC147174EFB405AA60C220FBE" box="[1192,1325,187,213]" country="United States of America" name="Washington" pageId="5" pageNumber="474">Washington</collectingRegion>
|
||
in having whorls with a strong peripheral keel and differs from both
|
||
<taxonomicName id="4C2B4D3BC147174EFCD85AC20AFA0F92" authorityName="Squires & Goedert" authorityYear="1996" box="[816,1013,223,249]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="palaiopacifica">
|
||
<emphasis id="B95FEAAAC147174EFCD85AC20AFA0F92" box="[816,1013,223,249]" italics="true" pageId="5" pageNumber="474">O. palaiopacifica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D3BC147174EFBC55AC20D8F0F92" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[1069,1152,223,249]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC147174EFBC55AC20D8F0F92" box="[1069,1152,223,249]" italics="true" pageId="5" pageNumber="474">O. kieli</emphasis>
|
||
</taxonomicName>
|
||
in having a more domed apical surface of the whorls, with deeply incised apical suture. According to
|
||
<bibRefCitation id="EFBA4B49C147174EFBEF5B1E0DEC0E76" author="Bartsch, P." box="[1031,1251,259,285]" pageId="5" pageNumber="474" pagination="167 - 176" refId="ref5143" refString="Bartsch, P. (1907) New mollusks of the family Vitrinellidae from the West Coast of America. Proceedings of the United States National Museum, 32, 167 - 176. https: // doi. org / 10.5479 / si. 00963801.32 - 1520.167" type="journal article" year="1907">Bartsch (1907: 172</bibRefCitation>
|
||
, figs. 7a–b) and
|
||
<bibRefCitation id="EFBA4B49C147174EFF7F5B3A08230E2A" author="Myers, B. W." box="[151,300,295,321]" pageId="5" pageNumber="474" pagination="64 - 66" refId="ref6379" refString="Myers, B. W. (1979) An historical review of the systematics of Orbitestella diegensis (Bartsch, 1907). The Festivus, 11, 64 - 66." type="journal article" year="1979">Myers (1979</bibRefCitation>
|
||
: fig.1) the base of the whorls of
|
||
<taxonomicName id="4C2B4D3BC147174EFD485B3A0A240E2A" baseAuthorityName="Bartsch" baseAuthorityYear="1907" box="[672,811,295,321]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="diegensis">
|
||
<emphasis id="B95FEAAAC147174EFD485B3A0A240E2A" box="[672,811,295,321]" italics="true" pageId="5" pageNumber="474">O. diegensis</emphasis>
|
||
</taxonomicName>
|
||
are rounded, whereas in
|
||
<taxonomicName id="4C2B4D3BC147174EFBA35B3A0C1F0E2A" authorityName="Squires & Goedert" authorityYear="1996" box="[1099,1296,295,321]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="palaiopacifica">
|
||
<emphasis id="B95FEAAAC147174EFBA35B3A0C1F0E2A" box="[1099,1296,295,321]" italics="true" pageId="5" pageNumber="474">O. palaiopacifica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D3BC147174EFAA15B3A0C930E2A" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[1353,1436,295,321]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC147174EFAA15B3A0C930E2A" box="[1353,1436,295,321]" italics="true" pageId="5" pageNumber="474">O. kieli</emphasis>
|
||
</taxonomicName>
|
||
they are flattened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316533C147174EFF2F5B720B3F0EC6" pageId="5" pageNumber="474" type="discussion">
|
||
<paragraph id="8B9436B8C147174EFF2F5B720B3F0EC6" blockId="5.[151,1437,151,429]" pageId="5" pageNumber="474">
|
||
<emphasis id="B95FEAAAC147174EFF2F5B7208340EE2" bold="true" box="[199,315,367,393]" pageId="5" pageNumber="474">Remarks.</emphasis>
|
||
<taxonomicName id="4C2B4D3BC147174EFEAB5B720B0D0EE2" authorityName="Chernyshev & Goedert" authorityYear="2021" box="[323,514,367,393]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="species" species="kieli">
|
||
<emphasis id="B95FEAAAC147174EFEAB5B720B0D0EE2" box="[323,514,367,393]" italics="true" pageId="5" pageNumber="474">Orbitestella kieli</emphasis>
|
||
</taxonomicName>
|
||
is the third fossil species of the genus
|
||
<taxonomicName id="4C2B4D3BC147174EFC575B720D4A0EE2" authorityName="Iredale" authorityYear="1917" box="[959,1093,367,393]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="5" pageNumber="474" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B95FEAAAC147174EFC575B720D4A0EE2" box="[959,1093,367,393]" italics="true" pageId="5" pageNumber="474">Orbitestella</emphasis>
|
||
</taxonomicName>
|
||
and family
|
||
<taxonomicName id="4C2B4D3BC147174EFB385B720C7A0EE2" authorityName="Iredale" authorityYear="1917" box="[1232,1397,367,393]" class="Gastropoda" family="Orbitestellidae" kingdom="Animalia" pageId="5" pageNumber="474" phylum="Mollusca" rank="family">Orbitestellidae</taxonomicName>
|
||
reported from western North America.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |