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<document id="7327E0E4E870147D4F42157954E5CF6B" ID-DOI="10.5852/ejt.2020.720.1109" ID-GBIF-Dataset="545aa4ea-1874-4f74-a4c4-a2aeb97ed657" ID-ISSN="2118-9773" ID-Zenodo-Dep="4068355" ID-ZooBank="0CF83E89-2532-407B-921F-C05F9A29C68D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1601993099685" checkinUser="valdenar" docAuthor="Bergsten, Johannes, Ranarilalatiana, Tolotra &amp; Biström, Olof" docDate="2020" docId="03828798D37F6464FD98FEB5A856FDBC" docLanguage="en" docName="ejt-720_begsten_ranarilalatinana_bistroem.pdf.imf" docOrigin="European Journal of Taxonomy 720" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Bidessus nero Gschwendtner 1933" docType="treatment" docVersion="11" lastPageNumber="13" masterDocId="FFBBFFE0D3756468FFAEFFBDAD21FFBB" masterDocTitle="A new species of Bidessus from Anjozorobe-Angavo and a review of Malagasy Bidessus (Coleoptera, Dytiscidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="11" updateTime="1701999645753" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="C4181F3DE5B7FC1E203CE70C1CD05B27" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B9455D38E38B6C16EF7E291186397D40">
<mods:title id="63F9AF1D2E929B2D9B5DA4096896B19A">A new species of Bidessus from Anjozorobe-Angavo and a review of Malagasy Bidessus (Coleoptera, Dytiscidae)</mods:title>
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<mods:name id="1416946E5AEADE33F4BFAF85B686A129" type="personal">
<mods:role id="EC4C7930A1BAF761C26F0E94E28F580E">
<mods:roleTerm id="D900641A9C3549B37FE15C2A1598AB9D">Author</mods:roleTerm>
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<mods:namePart id="519F17F52B56AE9EC0B23EE670F0B4DF">Bergsten, Johannes</mods:namePart>
<mods:nameIdentifier id="C952C2843AE53AF9338173B6291A7680" type="LSID">64120AE6-1D4B-4C29-AA37-46DA7C7A26A8</mods:nameIdentifier>
<mods:affiliation id="99516FBE9584561A37FE5F3E01C8FF44">Department of Zoology, Swedish Museum of Natural History, Box 50007, SE- 10405 Stockholm, Sweden.</mods:affiliation>
<mods:nameIdentifier id="7C53DC402EDE2455DD403E0096A59C5F" type="email">johannes.bergsten@nrm.se</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="77B95046D6FE2F5332CC755967EE5455">Ranarilalatiana, Tolotra</mods:namePart>
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<mods:affiliation id="D727B7EE7E68201B2D9587689079788C">Department of Entomology, Faculty of Sciences, Box 906, University of Antananarivo, 101 Antananarivo, Madagascar.</mods:affiliation>
<mods:nameIdentifier id="45F6488503D57246D1CA5DDBAD6E7E22" type="email">t.ranarilalatiana@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="CF05721F76E65FBADA621B4FE0798CA2">Author</mods:roleTerm>
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<mods:namePart id="D7E8A92C131294C110EEEC1D6A3DFB82">Biström, Olof</mods:namePart>
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<mods:affiliation id="0AC811C6C50D5B564715A22AA99F569B">Finnish Museum of Natural History, Zoology Unit, P. O. Box 17, FI- 00014 University of Helsinki, Finland.</mods:affiliation>
<mods:nameIdentifier id="9F2E96FA4B65CBF9EC232ECC71A9CA59" type="email">olof.bistrom@helsinki.fi</mods:nameIdentifier>
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<mods:title id="10540722C6DB4716AD077C3992094E94">European Journal of Taxonomy</mods:title>
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<mods:date id="336FB0762300918C3B380E5C8DDC92A1">2020</mods:date>
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<mods:number id="886552F002DF92934C5D68132619DCBC">2020-10-05</mods:number>
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<mods:number id="EA558ADD5809EB6E7513BA6E20BACDDB">720</mods:number>
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<treatment id="03828798D37F6464FD98FEB5A856FDBC" ID-DOI="http://doi.org/10.5281/zenodo.4329741" ID-GBIF-Taxon="168237708" ID-Zenodo-Dep="4329741" LSID="urn:lsid:plazi:treatment:03828798D37F6464FD98FEB5A856FDBC" httpUri="http://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="C3316505D37F6462FD98FEB5AEDFFE98" box="[566,1022,264,291]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8B94368ED37F6462FD98FEB5AEDFFE98" blockId="10.[566,1022,264,326]" box="[566,1022,264,291]" pageId="10" pageNumber="11">
<heading id="D0DC81E2D37F6462FD98FEB5AEDFFE98" box="[566,1022,264,291]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="4C2B4D0DD37F6462FD98FEB5AEDFFE98" ID-CoL="LSRD" authority="Gschwendtner, 1933" authorityName="Gschwendtner" authorityYear="1933" box="[566,1022,264,291]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nero">
<emphasis id="B95FEA9CD37F6462FD98FEB5AF81FE98" bold="true" box="[566,672,264,291]" italics="true" pageId="10" pageNumber="11">Bidessus</emphasis>
cf.
<emphasis id="B95FEA9CD37F6462FD63FEB4AE25FE98" bold="true" box="[717,772,265,291]" italics="true" pageId="10" pageNumber="11">nero</emphasis>
<bibRefCitation id="EFBA4B7FD37F6462FCA2FEB4AEDFFE98" author="Gschwendtner L." box="[780,1022,265,291]" pageId="10" pageNumber="11" pagination="84 - 95" refId="ref7416" refString="Gschwendtner L. 1933. Neue Schwimmkafer aus Belgisch Kongo. IV. Teil. Revue de Zoologie et botanique africaines 23: 84 - 95." type="journal article" year="1933">Gschwendtner, 1933</bibRefCitation>
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<subSubSection id="C3316505D37F6462FD7CFE91AE43FEFD" box="[722,866,300,326]" pageId="10" pageNumber="11" type="description">
<paragraph id="8B94368ED37F6462FD7CFE91AE43FEFD" blockId="10.[566,1022,264,326]" box="[722,866,300,326]" pageId="10" pageNumber="11">
<figureCitation id="13102A0BD37F6462FD7CFE91AE10FEFD" box="[722,817,300,326]" captionStart="Fig" captionStartId="3.[189,232,1887,1913]" captionTargetBox="[233,1355,265,1851]" captionTargetId="figure-11@3.[233,1355,265,1851]" captionTargetPageId="3" captionText="Fig. 1. Habitus photographs of Malagasy species of Bidessus Sharp, 1882. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986, ♀, paratype, http://id.luomus.fi/GL.8408 (FMNH). B. B. cf. ceratus Guignot, 1941, http://id.luomus.fi/GC.2001 (FMNH). C. B. anjozorobe sp. nov., paratype (NHRS-JLKB000025034). D. B. cf. nero Gschwendtner, 1933 (NHRS-JLKB000025032)." figureDoi="http://doi.org/10.5281/zenodo.4068357" httpUri="https://zenodo.org/record/4068357/files/figure.png" pageId="10" pageNumber="11">Figs 1D</figureCitation>
,
<figureCitation id="13102A0BD37F6462FC90FE91AE43FEFD" box="[830,866,300,326]" captionStart="Fig" captionStartId="9.[189,232,1888,1914]" captionTargetBox="[366,1209,282,1833]" captionTargetId="figure-11@9.[308,1280,265,1849]" captionTargetPageId="9" captionText="Fig. 3. Male genitalia of the Malagasy species of Bidessus Sharp, 1882. For each subpanel from left to right penis in dorsal view, penis in lateral view and paramere in lateral view. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986. B. B. cf. ceratus Guignot, 1941. C. B. anjozorobe sp. nov. D. B. cf. nero Gschwendtner, 1933. E. B. nesioticus Guignot, 1956. F. B. perexiguus Kolbe, 1883. G. B. longistriga Régimbart, 1895." figureDoi="http://doi.org/10.5281/zenodo.4068361" httpUri="https://zenodo.org/record/4068361/files/figure.png" pageId="10" pageNumber="11">3D</figureCitation>
</paragraph>
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<subSubSection id="C3316505D37F6462FF13FED3AE74FDB0" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8B94368ED37F6462FF13FED3AFACFE3C" blockId="10.[189,653,365,392]" box="[189,653,365,392]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB3B11A0D37F6462FF13FED3AFACFE3C" box="[189,653,365,392]" pageId="10" pageNumber="11">
<treatmentCitation id="0A8A109FD37F6462FF13FED3AFA8FE3C" author="Gschwendtner L." box="[189,649,365,392]" page="86" pageId="10" pageNumber="11" year="1933">
<taxonomicName id="4C2B4D0DD37F6462FF13FED3AFA8FE3C" ID-CoL="LSRD" authority="Gschwendtner, 1933: 86" authorityName="Gschwendtner" authorityPageNumber="86" authorityYear="1933" box="[189,649,365,392]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nero">
<emphasis id="B95FEA9CD37F6462FF13FED3AC41FE3C" box="[189,352,366,392]" italics="true" pageId="10" pageNumber="11">Bidessus nero</emphasis>
<bibRefCitation id="EFBA4B7FD37F6462FEC9FED3AFA8FE3C" author="Gschwendtner L." box="[359,649,365,392]" pageId="10" pageNumber="11" pagination="84 - 95" refId="ref7416" refString="Gschwendtner L. 1933. Neue Schwimmkafer aus Belgisch Kongo. IV. Teil. Revue de Zoologie et botanique africaines 23: 84 - 95." type="journal article" year="1933">Gschwendtner, 1933: 86</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8B94368ED37F6462FF13FE12A9F0FE72" blockId="10.[189,1233,431,458]" box="[189,1233,431,458]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB3B11A0D37F6462FF13FE12A9F0FE72" box="[189,1233,431,458]" pageId="10" pageNumber="11">
<taxonomicName id="4C2B4D0DD37F6462FF13FE12AC41FE72" ID-CoL="LSRD" authorityName="Gschwendtner" authorityYear="1933" box="[189,352,431,457]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nero">
<emphasis id="B95FEA9CD37F6462FF13FE12AC41FE72" box="[189,352,431,457]" italics="true" pageId="10" pageNumber="11">Bidessus nero</emphasis>
</taxonomicName>
– Bistr̂m 1985: 18. —
<treatmentCitation id="0A8A109FD37F6462FDD0FE12AE76FE71" author="Nilsson A. N." box="[638,855,431,458]" page="115" pageId="10" pageNumber="11" year="2001">
<bibRefCitation id="EFBA4B7FD37F6462FDD0FE12AE76FE71" author="Nilsson A. N." box="[638,855,431,458]" pageId="10" pageNumber="11" pagination="1 - 395" refId="ref7602" refString="Nilsson A. N. 2001. Dytiscidae (Coleoptera). World Catalogue of Insects 3: 1 - 395." type="journal article" year="2001">Nilsson 2001: 115</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0A8A109FD37F6462FC29FE12A9ECFE72" author="Nilsson A. N. &amp; Hajek J." box="[903,1229,431,457]" page="104" pageId="10" pageNumber="11" year="2020">
<bibRefCitation id="EFBA4B7FD37F6462FC29FE12A9ECFE72" author="Nilsson A. N. &amp; Hajek J." box="[903,1229,431,457]" pageId="10" pageNumber="11" pagination="1 - 395" refId="ref7624" refString="Nilsson A. N. &amp; Hajek J. 2020. A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga) Version 1. I. 2020. Electronic and updated version of Nilsson A. N. (2001). Dytiscidae. In: World Catalogue of Insects: 1 - 395. Available from www. waterbeetles. eu [accessed 5 Sep. 2020]." type="book chapter" year="2020">Nilsson &amp; Hajek 2020: 104</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B94368ED37F6462FF13FE4DAE74FDB0" blockId="10.[189,853,496,523]" box="[189,853,496,523]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB3B11A0D37F6462FF13FE4DAE74FDB0" box="[189,853,496,523]" pageId="10" pageNumber="11">
<taxonomicName id="4C2B4D0DD37F6462FF13FE4DAF73FDB0" authority="(Guignot, 1946)" baseAuthorityName="Guignot" baseAuthorityYear="1946" box="[189,594,496,523]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B95FEA9CD37F6462FF13FE4DACAFFDB1" box="[189,398,496,522]" italics="true" pageId="10" pageNumber="11">Bidessus unicolor</emphasis>
(Guignot, 1946)
</taxonomicName>
–
<treatmentCitation id="0A8A109FD37F6462FDDEFE4DAE73FDB0" author="Guignot F." box="[624,850,496,523]" page="255" pageId="10" pageNumber="11" year="1959">
<bibRefCitation id="EFBA4B7FD37F6462FDDEFE4DAE73FDB0" author="Guignot F." box="[624,850,496,523]" pageId="10" pageNumber="11" pagination="1 - 313" refId="ref7502" refString="Guignot F. 1959. Revision des Hydrocanthares d'Afrique (Coleoptera: Dytiscoidea). Premiere Partie. Annales de Musee royal du Congo Belge, Tervuren: Sciences zoologiques 70: 1 - 313." type="journal article" year="1959">Guignot 1959: 255</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3316505D37F6462FF13FD8FAF2FFC6D" pageId="10" pageNumber="11" type="description">
<paragraph id="8B94368ED37F6462FF13FD8FACEDFDF7" blockId="10.[189,460,562,588]" box="[189,460,562,588]" pageId="10" pageNumber="11">
<heading id="D0DC81E2D37F6462FF13FD8FACEDFDF7" bold="true" box="[189,460,562,588]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B95FEA9CD37F6462FF13FD8FACEDFDF7" bold="true" box="[189,460,562,588]" pageId="10" pageNumber="11">Diagnostic characters</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED37F6462FF13FDDDA94FFC2F" blockId="10.[189,1398,608,917]" pageId="10" pageNumber="11">
Head frontally not margined; provided with a slight depression slightly posterior to anterior edge of head (between eyes); frontal depression is medially broadly vague and indistinct. Pronotum on disc with a broad impunctate area. Pronotal striae at base not deeply impressed, distinctly angled and not smoothly curved. Basal striae of elytra medium long and distinct. Anteriorly sutural lines of elytra disappear abruptly a short distance before reaching frontal edge; posteriorly lines fade away long before reaching apex. Elytra with punctures on each side of basal striae of equal size and density (rather fine, sparse and slightly irregular). Apical ventrite with rather fine, slightly sparse punctuation. Female with outline of elytra posteriorly smooth (minor process absent). Penis in lateral view quite slender and evenly curved; apex distinct (also slightly curved). Penis apex not distinctly extended (
<figureCitation id="13102A0BD37F6462FBACFCC7A940FC2F" box="[1026,1121,890,916]" captionStart="Fig" captionStartId="9.[189,232,1888,1914]" captionTargetBox="[366,1209,282,1833]" captionTargetId="figure-11@9.[308,1280,265,1849]" captionTargetPageId="9" captionText="Fig. 3. Male genitalia of the Malagasy species of Bidessus Sharp, 1882. For each subpanel from left to right penis in dorsal view, penis in lateral view and paramere in lateral view. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986. B. B. cf. ceratus Guignot, 1941. C. B. anjozorobe sp. nov. D. B. cf. nero Gschwendtner, 1933. E. B. nesioticus Guignot, 1956. F. B. perexiguus Kolbe, 1883. G. B. longistriga Régimbart, 1895." figureDoi="http://doi.org/10.5281/zenodo.4068361" httpUri="https://zenodo.org/record/4068361/files/figure.png" pageId="10" pageNumber="11">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="8B94368ED37F6462FF13FC06AF2FFC6D" blockId="10.[189,526,955,982]" box="[189,526,955,982]" pageId="10" pageNumber="11">
Length of body
<quantity id="4CD39B6BD37F6462FED7FC06AF2BFC6D" box="[377,522,955,982]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="10" pageNumber="11" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.5–1.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3316505D37F6462FF13FC40AE20F8BD" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8B94368ED37F6462FF13FC40AC8BFBAC" blockId="10.[189,1399,1021,1798]" box="[189,426,1021,1047]" pageId="10" pageNumber="11">
<heading id="D0DC81E2D37F6462FF13FC40AC8BFBAC" bold="true" box="[189,426,1021,1047]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B95FEA9CD37F6462FF13FC40AC8BFBAC" bold="true" box="[189,426,1021,1047]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED37F6462FF13FB91AE20F8BD" blockId="10.[189,1399,1021,1798]" pageId="10" pageNumber="11">
<materialsCitation id="3B433CD3D37F6462FF13FB91AE0AFB0B" ID-GBIF-Occurrence="2871635309" collectingDate="2014-11-01" collectionCode="NHRS-JLKB" collectorName="J. Bergsten &amp; T. Ranarilalatiana &amp; S. Holmgren" country="Madagascar" elevation="1700" latitude="-20.7722" location="Amoron'i Mania region" longLatPrecision="7" longitude="47.1809" pageId="10" pageNumber="11" specimenCode="NHRS-JLKB000025032" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa former province">
<collectingCountry id="F33C761ED37F6462FF13FB91ACAFFBFD" box="[189,398,1068,1094]" name="Madagascar" pageId="10" pageNumber="11">MADAGASCAR</collectingCountry>
•
<specimenCount id="9D2DFD07D37F6462FE18FB91ACCBFBFE" box="[438,490,1065,1094]" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49EFF86CD37F6462FDAFFB96AE4FFBFE" box="[513,878,1067,1093]" pageId="10" pageNumber="11">Fianarantsoa former province</collectingRegion>
,
<location id="8EF46055D37F6462FC2EFB96A98BFBFE" LSID="urn:lsid:plazi:treatment:03828798D37F6464FD98FEB5A856FDBC:8EF46055D37F6462FC2EFB96A98BFBFE" box="[896,1194,1067,1093]" country="Madagascar" latitude="-20.7722" longLatPrecision="7" longitude="47.1809" name="Amoron'i Mania region" pageId="10" pageNumber="11" stateProvince="Fianarantsoa former province">Amoron’i Mania region</location>
,
<locationDeviation id="9A262FACD37F6462FB13FB91AF11FBD2" pageId="10" pageNumber="11">
<quantity id="4CD39B6BD37F6462FB13FB91A821FBFE" box="[1213,1280,1067,1094]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="km" value="3.0">3 km</quantity>
south of Ambalamanakana next to RN7
</locationDeviation>
,
<location id="8EF46055D37F6462FD92FBF2AE61FBD2" LSID="urn:lsid:plazi:treatment:03828798D37F6464FD98FEB5A856FDBC:8EF46055D37F6462FD92FBF2AE61FBD2" box="[572,832,1103,1129]" country="Madagascar" latitude="-20.7722" longLatPrecision="7" longitude="47.1809" name="Ankazomivady forest" pageId="10" pageNumber="11" stateProvince="Fianarantsoa former province">Ankazomivady forest</location>
;
<geoCoordinate id="EE1F5049D37F6462FCE3FBF2AEF5FBD2" box="[845,980,1103,1129]" degrees="20.7722" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="-20.7722">20.7722° S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D37F6462FC4DFBF2A94DFBD2" box="[995,1132,1103,1129]" degrees="47.1809" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="47.1809">47.1809° E</geoCoordinate>
;
<quantity id="4CD39B6BD37F6462FBD4FBF2A9F4FBD2" box="[1146,1237,1103,1129]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="11" unit="m" value="1700.0">
<elevation id="0006D1BDD37F6462FBD4FBF2A9F4FBD2" box="[1146,1237,1103,1129]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="11" unit="m" value="1700.0">1700 m</elevation>
</quantity>
a.s.l.;
<date id="FF95104ED37F6462FA89FBF2ADDAFB37" pageId="10" pageNumber="11" value="2014-11-01">
<collectingDate id="EFD1E9A6D37F6462FA89FBF2ADDAFB37" pageId="10" pageNumber="11" value="2014-11-01">1 Nov. 2014</collectingDate>
</date>
;
<collectorName id="26DE5358D37F6462FEAAFBCFACA7FB37" box="[260,390,1138,1164]" pageId="10" pageNumber="11">J. Bergsten</collectorName>
,
<collectorName id="26DE5358D37F6462FE23FBCFAF7AFB37" box="[397,603,1138,1164]" pageId="10" pageNumber="11">T. Ranarilalatiana</collectorName>
and
<collectorName id="26DE5358D37F6462FD23FBCFAE3EFB37" box="[653,799,1138,1164]" pageId="10" pageNumber="11">S.Holmgren</collectorName>
leg.; collecting event MAD14-02; hygropetric rocks and marsh with vegetation;
<specimenCode id="DB8D9EF5D37F6462FDA7FB28AE0AFB0B" box="[521,811,1173,1200]" collectionCode="NHRS-JLKB" pageId="10" pageNumber="11">NHRS-JLKB000025032</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D37F6462FCE6FB28AFFEFB68" ID-GBIF-Occurrence="2871635302" collectingDate="2014-11-01" collectionCode="NHRS-JLKB" collectorName="J. Bergsten &amp; T. Ranarilalatiana &amp; S. Holmgren" country="Madagascar" elevation="1700" latitude="-20.7722" location="Amoron'i Mania region" longLatPrecision="7" longitude="47.1809" pageId="10" pageNumber="11" specimenCode="NHRS-JLKB000025024" specimenCount="5" stateProvince="Fianarantsoa former province">
<specimenCount id="9D2DFD07D37F6462FCE6FB28AEBEFB0B" box="[840,927,1173,1200]" pageId="10" pageNumber="11">5 specs</specimenCount>
(preserved in ethanol); same collection data as for preceding;
<specimenCode id="DB8D9EF5D37F6462FE13FB05AFFEFB68" box="[445,735,1208,1235]" collectionCode="NHRS-JLKB" pageId="10" pageNumber="11">NHRS-JLKB000025024</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D37F6462FD58FB05A847FB68" ID-GBIF-Occurrence="2871635303" box="[758,1382,1208,1235]" collectingDate="2014-11-01" collectionCode="FMNH, PBZT, MBC" collectorName="J. Bergsten &amp; T. Ranarilalatiana &amp; S. Holmgren" country="Madagascar" elevation="1700" latitude="-20.7722" location="Amoron'i Mania region" longLatPrecision="7" longitude="47.1809" pageId="10" pageNumber="11" specimenCount="2" stateProvince="Fianarantsoa former province">
<specimenCount id="9D2DFD07D37F6462FD58FB05AE6BFB68" box="[758,842,1208,1235]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED3AAE4BD37F6462FABFFB04A847FB68" box="[1297,1382,1209,1235]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="11" type="Museum">FMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D37F6462FF13FB61AE46FB4D" ID-GBIF-Occurrence="2871635305" box="[189,871,1243,1270]" collectingDate="2014-11-01" collectionCode="FMNH, PBZT, MBC" collectorName="J. Bergsten &amp; T. Ranarilalatiana &amp; S. Holmgren" country="Madagascar" elevation="1700" latitude="-20.7722" location="Amoron'i Mania region" longLatPrecision="7" longitude="47.1809" pageId="10" pageNumber="11" specimenCount="2" stateProvince="Fianarantsoa former province">
<specimenCount id="9D2DFD07D37F6462FF13FB61AC30FB4D" box="[189,273,1244,1270]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED3AAE4BD37F6462FD78FB61AE3FFB4D" box="[726,798,1244,1270]" collectionName="Madagascar, Antananarivo, Parc Botanique et Zoologique de Tsimbazaza" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="10" pageNumber="11">PBZT</collectionCode>
/
<collectionCode id="ED3AAE4BD37F6462FC89FB61AE46FB4D" box="[807,871,1244,1270]" country="USA" name="Montgomery Botanical Center" pageId="10" pageNumber="11">MBC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D37F6462FCD0FB61ACA6FA38" ID-GBIF-Occurrence="2871635301" collectingDate="2014-11-03" collectionCode="NHRS" collectorName="J. Bergsten &amp; T. Ranarilalatiana &amp; S. Holmgren" elevation="1110" latitude="-21.2397" location="Matsiara Ambony region" longLatPrecision="7" longitude="47.3774" pageId="10" pageNumber="11" specimenCode="NHRS- JLKB000025025" specimenCount="1" stateProvince="Fianarantsoa former province">
<specimenCount id="9D2DFD07D37F6462FCD0FB61AEEFFB4D" box="[894,974,1244,1270]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
(preserved in ethanol);
<collectingRegion id="49EFF86CD37F6462FB4FFB66AC5FFAA2" pageId="10" pageNumber="11">Fianarantsoa former province</collectingRegion>
,
<location id="8EF46055D37F6462FE26FB42AF91FAA2" LSID="urn:lsid:plazi:treatment:03828798D37F6464FD98FEB5A856FDBC:8EF46055D37F6462FE26FB42AF91FAA2" box="[392,688,1279,1305]" latitude="-21.2397" longLatPrecision="7" longitude="47.3774" name="Matsiara Ambony region" pageId="10" pageNumber="11" stateProvince="Fianarantsoa former province">Matsiara Ambony region</location>
,
<location id="8EF46055D37F6462FD15FB42AE6EFAA2" LSID="urn:lsid:plazi:treatment:03828798D37F6464FD98FEB5A856FDBC:8EF46055D37F6462FD15FB42AE6EFAA2" box="[699,847,1279,1305]" latitude="-21.2397" longLatPrecision="7" longitude="47.3774" name="Ranomafana" pageId="10" pageNumber="11" stateProvince="Fianarantsoa former province">Ranomafana</location>
NP,
<locationDeviation id="9A262FACD37F6462FC28FB43A850FAA2" box="[902,1393,1278,1305]" pageId="10" pageNumber="11">
<quantity id="4CD39B6BD37F6462FC28FB43AEF1FAA2" box="[902,976,1278,1305]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="m" value="500.0">500 m</quantity>
from the road at Vohiparara village
</locationDeviation>
;
<geoCoordinate id="EE1F5049D37F6462FF13FA9FAC62FA87" box="[189,323,1314,1340]" degrees="21.2397" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="-21.2397">21.2397° S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D37F6462FEFDFA9FACFDFA87" box="[339,476,1314,1340]" degrees="47.3774" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="47.3774">47.3774° E</geoCoordinate>
;
<quantity id="4CD39B6BD37F6462FE45FA9FAF65FA87" box="[491,580,1314,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="10" pageNumber="11" unit="m" value="1110.0">
<elevation id="0006D1BDD37F6462FE45FA9FAF65FA87" box="[491,580,1314,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="10" pageNumber="11" unit="m" value="1110.0">1110 m</elevation>
</quantity>
a.s.l.;
<date id="FF95104ED37F6462FD3BFA9FAE0CFA87" box="[661,813,1314,1340]" pageId="10" pageNumber="11" value="2014-11-03">
<collectingDate id="EFD1E9A6D37F6462FD3BFA9FAE0CFA87" box="[661,813,1314,1340]" pageId="10" pageNumber="11" value="2014-11-03">3 Nov. 2014</collectingDate>
</date>
;
<collectorName id="26DE5358D37F6462FC95FA9FAEE3FA87" box="[827,962,1314,1340]" pageId="10" pageNumber="11">J. Bergsten</collectorName>
,
<collectorName id="26DE5358D37F6462FC61FA9FA980FA87" box="[975,1185,1314,1340]" pageId="10" pageNumber="11">T. Ranarilalatiana</collectorName>
and
<collectorName id="26DE5358D37F6462FB70FA9FA857FA87" box="[1246,1398,1314,1340]" pageId="10" pageNumber="11">S. Holmgren</collectorName>
leg.; collecting event MAD14-10; small forest lake with floating margins of vegetation;
<specimenCode id="DB8D9EF5D37F6462FAB0FAF8ACA6FA38" pageId="10" pageNumber="11">NHRS- JLKB000025025</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D37F6462FE35FAD5AF11FA56" ID-GBIF-Occurrence="2871635308" collectingDate="2014-11-24" collectionCode="NHRS-JLKB" collectorName="J. Bergsten &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" elevation="900" latitude="-18.9247" location="Alaotra Mangoro region" longLatPrecision="7" longitude="48.1828" pageId="10" pageNumber="11" specimenCode="NHRS-JLKB000025026" specimenCount="1" stateProvince="Toamasina former province">
<specimenCount id="9D2DFD07D37F6462FE35FAD5ACCFFA38" box="[411,494,1384,1411]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="49EFF86CD37F6462FE59FAD5AE1BFA39" box="[503,826,1384,1410]" pageId="10" pageNumber="11">Toamasina former province</collectingRegion>
,
<location id="8EF46055D37F6462FCEFFAD5A97CFA39" LSID="urn:lsid:plazi:treatment:03828798D37F6464FD98FEB5A856FDBC:8EF46055D37F6462FCEFFAD5A97CFA39" box="[833,1117,1384,1411]" latitude="-18.9247" longLatPrecision="7" longitude="48.1828" name="Alaotra Mangoro region" pageId="10" pageNumber="11" stateProvince="Toamasina former province">Alaotra Mangoro region</location>
,
<locationDeviation id="9A262FACD37F6462FBC8FAD5AE35FA1E" pageId="10" pageNumber="11">
<quantity id="4CD39B6BD37F6462FBC8FAD5A9BEFA39" box="[1126,1183,1384,1410]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="11" unit="km" value="6.0">6 km</quantity>
W of Moramanga, Betsabora river by RN2 near Antsampanana village
</locationDeviation>
;
<geoCoordinate id="EE1F5049D37F6462FCB1FA31AE83FA1D" box="[799,930,1420,1446]" degrees="18.9247" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="-18.9247">18.9247° S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D37F6462FC00FA31A915FA1D" box="[942,1076,1420,1446]" degrees="48.1828" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="48.1828">48.1828° E</geoCoordinate>
;
<quantity id="4CD39B6BD37F6462FB91FA31A9A9FA1D" box="[1087,1160,1420,1446]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="m" value="900.0">
<elevation id="0006D1BDD37F6462FB91FA31A9A9FA1D" box="[1087,1160,1420,1446]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="m" value="900.0">900 m</elevation>
</quantity>
a.s.l.;
<date id="FF95104ED37F6462FB7CFA31A853FA1D" box="[1234,1394,1420,1446]" pageId="10" pageNumber="11" value="2014-11-24">
<collectingDate id="EFD1E9A6D37F6462FB7CFA31A853FA1D" box="[1234,1394,1420,1446]" pageId="10" pageNumber="11" value="2014-11-24">24 Nov. 2014</collectingDate>
</date>
;
<collectorName id="26DE5358D37F6462FF13FA12AC60FA72" box="[189,321,1455,1481]" pageId="10" pageNumber="11">J. Bergsten</collectorName>
,
<collectorName id="26DE5358D37F6462FEE2FA12AF0EFA72" box="[332,559,1455,1481]" pageId="10" pageNumber="11">J.H. Randriamihaja</collectorName>
and
<collectorName id="26DE5358D37F6462FDC8FA12AE17FA72" box="[614,822,1455,1481]" pageId="10" pageNumber="11">T. Ranarilalatiana</collectorName>
leg.; collecting event MAD14-81; river with side pools;
<specimenCode id="DB8D9EF5D37F6462FEA0FA6FAF11FA56" box="[270,560,1490,1517]" collectionCode="NHRS-JLKB" pageId="10" pageNumber="11">NHRS-JLKB000025026</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D37F6462FDE2FA6FAFEAF989" ID-GBIF-Occurrence="2871635304" collectedFrom="wide and calm portion of small river" collectingDate="2016-05-18" collectionCode="CMM" elevation="1531" latitude="-20.666666" location="Ramahandrison and Manuel" longLatPrecision="15" longitude="47.128193" pageId="10" pageNumber="11" specimenCount="2" stateProvince="Fianarantsoa former province">
<specimenCount id="9D2DFD07D37F6462FDE2FA6FAF86FA57" box="[588,679,1490,1516]" pageId="10" pageNumber="11">2 specs</specimenCount>
;
<collectingRegion id="49EFF86CD37F6462FD1BFA6FA936FA57" box="[693,1047,1490,1516]" pageId="10" pageNumber="11">Fianarantsoa former province</collectingRegion>
,
<locationDeviation id="9A262FACD37F6462FB8AFA6FA857FA57" box="[1060,1398,1490,1516]" pageId="10" pageNumber="11">
ca
<quantity id="4CD39B6BD37F6462FBE9FA6FA9A2FA57" box="[1095,1155,1490,1516]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="km" value="9.0">9 km</quantity>
SW Ivato, Bemanta
</locationDeviation>
(inselberg);
<geoCoordinate id="EE1F5049D37F6462FEE5FA4EACFAF9B4" box="[331,475,1523,1551]" degrees="20" direction="south" minutes="40" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="00" value="-20.666666">20°40′00″ S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D37F6462FE46FA4EAFB0F9AB" box="[488,657,1523,1552]" degrees="47" direction="east" minutes="07" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="41.5" value="47.128193">47°07′41.5″ E</geoCoordinate>
;
<quantity id="4CD39B6BD37F6462FD30FA48AFD6F9AB" box="[670,759,1525,1552]" metricMagnitude="3" metricUnit="m" metricValue="1.531" pageId="10" pageNumber="11" unit="m" value="1531.0">
<elevation id="0006D1BDD37F6462FD30FA48AFD6F9AB" box="[670,759,1525,1552]" metricMagnitude="3" metricUnit="m" metricValue="1.531" pageId="10" pageNumber="11" unit="m" value="1531.0">1531 m</elevation>
</quantity>
a.s.l.;
<date id="FF95104ED37F6462FCEAFA48AEC5F9B4" box="[836,996,1525,1552]" pageId="10" pageNumber="11" value="2016-05-18">
<collectingDate id="EFD1E9A6D37F6462FCEAFA48AEC5F9B4" box="[836,996,1525,1552]" pageId="10" pageNumber="11" value="2016-05-18">18 May 2016</collectingDate>
</date>
;
<location id="8EF46055D37F6462FC5FFA48A81DF9B4" LSID="urn:lsid:plazi:treatment:03828798D37F6464FD98FEB5A856FDBC:8EF46055D37F6462FC5FFA48A81DF9B4" box="[1009,1340,1525,1551]" latitude="-20.666666" longLatPrecision="15" longitude="47.128193" name="Ramahandrison and Manuel" pageId="10" pageNumber="11" stateProvince="Fianarantsoa former province">Ramahandrison and Manuel</location>
leg.;
<collectedFrom id="38377573D37F6462FF13F9A5AF57F989" box="[189,630,1560,1586]" pageId="10" pageNumber="11">wide and calm portion of small river</collectedFrom>
;
<collectionCode id="ED3AAE4BD37F6462FD2DF9A5AFEAF989" box="[643,715,1560,1586]" country="Brazil" httpUri="http://grbio.org/cool/e1dy-v5iv" name="Culture Collection of Phytopathogenic Fungi (Colecao de Culturas de Fungos Fitopatogenicos Prof. Maria Menezes)" pageId="10" pageNumber="11">CMM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D37F6462FD49F9A5AE59F927" ID-GBIF-Occurrence="2871635306" collectedFrom="large pond with Nymphaeaceae and surrounding peat bog with Sphagnum and Drosera" collectingDate="2016-05-19" collectionCode="CMM" collectorName="Ramahandrison &amp; Manuel" elevation="1702" latitude="-20.775833" location="Fianarantsoa former province" longLatPrecision="21" longitude="47.18139" pageId="10" pageNumber="11" specimenCount="16" stateProvince="Fianarantsoa former province">
<specimenCount id="9D2DFD07D37F6462FD49F9A5AE71F988" box="[743,848,1560,1587]" pageId="10" pageNumber="11">16 specs</specimenCount>
;
<collectingRegion id="49EFF86CD37F6462FCF0F9A5A99EF989" box="[862,1215,1560,1586]" pageId="10" pageNumber="11">Fianarantsoa former province</collectingRegion>
,
<locationDeviation id="9A262FACD37F6462FB62F9A5AF5FF9EE" pageId="10" pageNumber="11">along road N7 between Ambositra and Fianarantsoa</locationDeviation>
,
<locationDeviation id="9A262FACD37F6462FD23F981A909F9EE" box="[653,1064,1595,1622]" pageId="10" pageNumber="11">
ca
<quantity id="4CD39B6BD37F6462FD1CF986AE26F9EE" box="[690,775,1595,1622]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="11" unit="km" value="3.5">3.5 km</quantity>
S of Ambalamanakana
</locationDeviation>
;
<geoCoordinate id="EE1F5049D37F6462FB99F987A9EBF9ED" box="[1079,1226,1594,1622]" degrees="20" direction="south" minutes="46" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="33" value="-20.775833">20°46′33″ S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D37F6462FB75F987A850F9ED" box="[1243,1393,1594,1622]" degrees="47" direction="east" minutes="10" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="53" value="47.18139">47°10′53″ E</geoCoordinate>
;
<quantity id="4CD39B6BD37F6462FF13F9E2AC3BF9C2" box="[189,282,1631,1657]" metricMagnitude="3" metricUnit="m" metricValue="1.702" pageId="10" pageNumber="11" unit="m" value="1702.0">
<elevation id="0006D1BDD37F6462FF13F9E2AC3BF9C2" box="[189,282,1631,1657]" metricMagnitude="3" metricUnit="m" metricValue="1.702" pageId="10" pageNumber="11" unit="m" value="1702.0">1702 m</elevation>
</quantity>
a.s.l.;
<date id="FF95104ED37F6462FEC1F9E2AF36F9C2" box="[367,535,1631,1657]" pageId="10" pageNumber="11" value="2016-05-19">
<collectingDate id="EFD1E9A6D37F6462FEC1F9E2AF36F9C2" box="[367,535,1631,1657]" pageId="10" pageNumber="11" value="2016-05-19">19 May 2016</collectingDate>
</date>
;
<collectorName id="26DE5358D37F6462FD89F9E2AFFEF9C2" box="[551,735,1631,1657]" pageId="10" pageNumber="11">Ramahandrison</collectorName>
and
<collectorName id="26DE5358D37F6462FC8EF9E2AE5BF9C2" box="[800,890,1631,1657]" pageId="10" pageNumber="11">Manuel</collectorName>
leg.;
<collectedFrom id="38377573D37F6462FC6DF9E2AE3FF927" pageId="10" pageNumber="11">
large pond with
<taxonomicName id="4C2B4D0DD37F6462FB20F9E2A860F9C2" authorityName="R.A.Salisbury" authorityYear="1805" box="[1166,1345,1631,1657]" class="Magnoliopsida" family="Nymphaeaceae" kingdom="Plantae" order="Nymphaeales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="family">Nymphaeaceae</taxonomicName>
and surrounding peat bog with
<taxonomicName id="4C2B4D0DD37F6462FDADF93FAF5EF927" authorityName="C.Linnaeus" authorityYear="1753" box="[515,639,1666,1692]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="10" pageNumber="11" phylum="Bryophyta" rank="genus">
<emphasis id="B95FEA9CD37F6462FDADF93FAF5EF927" box="[515,639,1666,1692]" italics="true" pageId="10" pageNumber="11">Sphagnum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2B4D0DD37F6462FD13F93EAE3FF927" box="[701,798,1667,1692]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="genus">
<emphasis id="B95FEA9CD37F6462FD13F93EAE3FF927" box="[701,798,1667,1692]" italics="true" pageId="10" pageNumber="11">Drosera</emphasis>
</taxonomicName>
</collectedFrom>
;
<collectionCode id="ED3AAE4BD37F6462FC9EF93FAE59F927" box="[816,888,1666,1692]" country="Brazil" httpUri="http://grbio.org/cool/e1dy-v5iv" name="Culture Collection of Phytopathogenic Fungi (Colecao de Culturas de Fungos Fitopatogenicos Prof. Maria Menezes)" pageId="10" pageNumber="11">CMM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D37F6462FC38F93FAE20F8BD" ID-GBIF-Occurrence="2871635307" collectedFrom="small marsh, very shallow, with small Poaceae and Cyperaceae, along slowly flowing stream" collectingDate="2017-12-29" collectionCode="CMM" elevation="1134" latitude="-21.2475" location="Ramahandrison and Manuel" longLatPrecision="15" longitude="47.376247" pageId="10" pageNumber="11" specimenCount="13" stateProvince="Fianarantsoa former province">
<specimenCount id="9D2DFD07D37F6462FC38F93FA921F927" box="[918,1024,1666,1692]" pageId="10" pageNumber="11">13 specs</specimenCount>
;
<collectingRegion id="49EFF86CD37F6462FBA1F93FA853F927" box="[1039,1394,1666,1692]" pageId="10" pageNumber="11">Fianarantsoa former province</collectingRegion>
,
<locationDeviation id="9A262FACD37F6462FF13F91BAF0FF904" box="[189,558,1701,1728]" pageId="10" pageNumber="11">
ca
<quantity id="4CD39B6BD37F6462FF71F918AC3BF904" box="[223,282,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="km" value="8.0">8 km</quantity>
WNW of Ranomafana
</locationDeviation>
,
<locationDeviation id="9A262FACD37F6462FD94F91BAE56F904" box="[570,887,1701,1728]" pageId="10" pageNumber="11">
ca
<quantity id="4CD39B6BD37F6462FDF2F918AFB9F904" box="[604,664,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="km" value="1.0">1 km</quantity>
SW of Vohiparara
</locationDeviation>
;
<geoCoordinate id="EE1F5049D37F6462FC2DF91EA935F904" box="[899,1044,1699,1727]" degrees="21" direction="south" minutes="14" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="51" value="-21.2475">21°14′51″ S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D37F6462FB8FF91EA9EAF97B" box="[1057,1227,1699,1728]" degrees="47" direction="east" minutes="22" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="34.5" value="47.376247">47°22′34.5″ E</geoCoordinate>
;
<quantity id="4CD39B6BD37F6462FB79F918A811F97B" box="[1239,1328,1701,1728]" metricMagnitude="3" metricUnit="m" metricValue="1.134" pageId="10" pageNumber="11" unit="m" value="1134.0">
<elevation id="0006D1BDD37F6462FB79F918A811F97B" box="[1239,1328,1701,1728]" metricMagnitude="3" metricUnit="m" metricValue="1.134" pageId="10" pageNumber="11" unit="m" value="1134.0">1134 m</elevation>
</quantity>
a.s.l.;
<date id="FF95104ED37F6462FF13F975AC45F959" box="[189,356,1736,1763]" pageId="10" pageNumber="11" value="2017-12-29">
<collectingDate id="EFD1E9A6D37F6462FF13F975AC45F959" box="[189,356,1736,1763]" pageId="10" pageNumber="11" value="2017-12-29">29 Dec. 2017</collectingDate>
</date>
;
<location id="8EF46055D37F6462FEDDF975AFE5F959" LSID="urn:lsid:plazi:treatment:03828798D37F6464FD98FEB5A856FDBC:8EF46055D37F6462FEDDF975AFE5F959" box="[371,708,1736,1762]" latitude="-21.2475" longLatPrecision="15" longitude="47.376247" name="Ramahandrison and Manuel" pageId="10" pageNumber="11" stateProvince="Fianarantsoa former province">Ramahandrison and Manuel</location>
leg.;
<collectedFrom id="38377573D37F6462FCA2F975AF87F8BD" pageId="10" pageNumber="11">
small marsh, very shallow, with small
<taxonomicName id="4C2B4D0DD37F6462FB4CF974A863F958" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1250,1346,1737,1763]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
and
<taxonomicName id="4C2B4D0DD37F6462FF13F951AC68F8BD" authorityName="de Jussieu" authorityYear="1789" box="[189,329,1772,1798]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
, along slowly flowing stream
</collectedFrom>
;
<collectionCode id="ED3AAE4BD37F6462FD1FF951AFDCF8BD" box="[689,765,1772,1798]" country="Brazil" httpUri="http://grbio.org/cool/e1dy-v5iv" name="Culture Collection of Phytopathogenic Fungi (Colecao de Culturas de Fungos Fitopatogenicos Prof. Maria Menezes)" pageId="10" pageNumber="11">CMM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3316505D37F6462FF13F890A98EF807" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8B94368ED37F6462FF13F890AC77F8FC" blockId="10.[189,342,1837,1863]" box="[189,342,1837,1863]" pageId="10" pageNumber="11">
<heading id="D0DC81E2D37F6462FF13F890AC77F8FC" bold="true" box="[189,342,1837,1863]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B95FEA9CD37F6462FF13F890AC77F8FC" bold="true" box="[189,342,1837,1863]" pageId="10" pageNumber="11">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED37F6462FF13F8E6A98EF807" blockId="10.[189,1398,1883,1981]" pageId="10" pageNumber="11">
First record from
<collectingCountry id="F33C761ED37F6462FE37F8E6AF06F8CE" box="[409,551,1883,1909]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
and here recorded from Ranomafana NP, near Ambalamanakana and Ivato along RN7 and near Moramanga (
<figureCitation id="13102A0BD37F6462FD38F8C2AFD5F822" box="[662,756,1919,1945]" captionStart="Fig" captionStartId="11.[189,232,1888,1914]" captionTargetBox="[270,1323,299,1826]" captionTargetId="figure-12@11.[232,1355,265,1852]" captionTargetPageId="11" captionText="Fig. 4. Distribution maps of the Malagasy species of Bidessus Sharp, 1882. A. B. longistriga Régimbart, 1895. B. B. perexiguus Kolbe, 1883. C. B. cf. nero Gschwendtner, 1933 (circle), B. nesioticus Guignot, 1956 (square), B. anjozorobe sp. nov. (triangle). D. B. apicidens Biström &amp; Sanfilippo, 1986 (square), B. cf. ceratus Guignot, 1941 (circle)." figureDoi="http://doi.org/10.5281/zenodo.4068363" httpUri="https://zenodo.org/record/4068363/files/figure.png" pageId="10" pageNumber="11">Fig. 4C</figureCitation>
). As tentatively defined here, the species is distributed in central and east Africa and
<collectingCountry id="F33C761ED37F6462FDB5F81FAF88F807" box="[539,681,1954,1980]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
(see Bistr̂m 1985;
<bibRefCitation id="EFBA4B7FD37F6462FC39F81FA982F807" author="Nilsson A. N. &amp; Hajek J." box="[919,1187,1954,1980]" pageId="10" pageNumber="11" pagination="1 - 395" refId="ref7624" refString="Nilsson A. N. &amp; Hajek J. 2020. A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga) Version 1. I. 2020. Electronic and updated version of Nilsson A. N. (2001). Dytiscidae. In: World Catalogue of Insects: 1 - 395. Available from www. waterbeetles. eu [accessed 5 Sep. 2020]." type="book chapter" year="2020">Nilsson &amp; Hajek 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF546606D37E6463FF13F8DDAF4FF85F" ID-DOI="http://doi.org/10.5281/zenodo.4068363" ID-Zenodo-Dep="4068363" httpUri="https://zenodo.org/record/4068363/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1888,1914]" targetBox="[270,1323,299,1826]" targetPageId="11">
<paragraph id="8B94368ED37E6463FF13F8DDAF4FF85F" blockId="11.[189,1399,1888,2020]" pageId="11" pageNumber="12">
<emphasis id="B95FEA9CD37E6463FF13F8DDAC28F8C1" bold="true" box="[189,265,1888,1914]" pageId="11" pageNumber="12">Fig. 4.</emphasis>
Distribution maps of the Malagasy species of
<taxonomicName id="4C2B4D0DD37E6463FC8DF8DDA901F8C1" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[803,1056,1888,1914]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEA9CD37E6463FC8DF8DDAEABF8C1" box="[803,906,1888,1914]" italics="true" pageId="11" pageNumber="12">Bidessus</emphasis>
Sharp, 1882
</taxonomicName>
.
<emphasis id="B95FEA9CD37E6463FB84F8DDA91EF8C1" bold="true" box="[1066,1087,1888,1914]" pageId="11" pageNumber="12">A</emphasis>
.
<taxonomicName id="4C2B4D0DD37E6463FBE3F8DCADD9F825" authority="Regimbart, 1895" authorityName="Regimbart" authorityYear="1895" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="longistriga">
<emphasis id="B95FEA9CD37E6463FBE3F8DCA9CCF8C1" box="[1101,1261,1888,1914]" italics="true" pageId="11" pageNumber="12">B. longistriga</emphasis>
<bibRefCitation id="EFBA4B7FD37E6463FB5DF8DDADD9F825" author="Regimbart M." pageId="11" pageNumber="12" pagination="1 - 244" refId="ref7828" refString="Regimbart M. 1895. Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. Memoires de la Societe entomologique de Belgique 4: 1 - 244." type="journal article" year="1895">Régimbart, 1895</bibRefCitation>
</taxonomicName>
.
<emphasis id="B95FEA9CD37E6463FEA8F83EAC3BF826" bold="true" box="[262,282,1923,1949]" pageId="11" pageNumber="12">B</emphasis>
.
<taxonomicName id="4C2B4D0DD37E6463FE87F839AF45F826" authority="Kolbe, 1883" authorityName="Kolbe" authorityYear="1883" box="[297,612,1923,1949]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perexiguus">
<emphasis id="B95FEA9CD37E6463FE87F839ACE8F826" box="[297,457,1924,1949]" italics="true" pageId="11" pageNumber="12">B. perexiguus</emphasis>
<bibRefCitation id="EFBA4B7FD37E6463FE7EF83EAF45F826" author="Kolbe H. J." box="[464,612,1923,1949]" pageId="11" pageNumber="12" pagination="383 - 427" refId="ref7538" refString="Kolbe H. J. 1883. Uber die madagascarischen Dytisciden des K ˆ nigl. entomologischen Museums zu Berlin. Archiv fur Naturgeschichte 49 (1): 383 - 427." type="journal article" year="1883">Kolbe, 1883</bibRefCitation>
</taxonomicName>
.
<emphasis id="B95FEA9CD37E6463FDDEF83EAFA4F826" bold="true" box="[624,645,1923,1949]" pageId="11" pageNumber="12">C</emphasis>
.
<taxonomicName id="4C2B4D0DD37E6463FD3AF839A926F826" authority="Gschwendtner, 1933" authorityName="Gschwendtner" authorityYear="1933" box="[660,1031,1923,1949]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nero">
<emphasis id="B95FEA9CD37E6463FD3AF839AF8CF826" box="[660,685,1924,1949]" italics="true" pageId="11" pageNumber="12">B.</emphasis>
cf.
<emphasis id="B95FEA9CD37E6463FD74F839AE2EF826" box="[730,783,1924,1949]" italics="true" pageId="11" pageNumber="12">nero</emphasis>
<bibRefCitation id="EFBA4B7FD37E6463FCB8F83EA926F826" author="Gschwendtner L." box="[790,1031,1923,1949]" pageId="11" pageNumber="12" pagination="84 - 95" refId="ref7416" refString="Gschwendtner L. 1933. Neue Schwimmkafer aus Belgisch Kongo. IV. Teil. Revue de Zoologie et botanique africaines 23: 84 - 95." type="journal article" year="1933">Gschwendtner, 1933</bibRefCitation>
</taxonomicName>
(circle),
<taxonomicName id="4C2B4D0DD37E6463FBDCF839ADD9F87A" authority="Guignot, 1956" authorityName="Guignot" authorityYear="1956" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nesioticus">
<emphasis id="B95FEA9CD37E6463FBDCF839A826F826" box="[1138,1287,1924,1949]" italics="true" pageId="11" pageNumber="12">B. nesioticus</emphasis>
<bibRefCitation id="EFBA4B7FD37E6463FAA1F83EADD9F87A" author="Guignot F." pageId="11" pageNumber="12" pagination="75 - 80" refId="ref7478" refString="Guignot F. 1956. Contribution a la connaissance de la faune dystiscidienne Malgache. Le Naturaliste malgache 8: 75 - 80." type="journal article" year="1956">Guignot, 1956</bibRefCitation>
</taxonomicName>
(square),
<taxonomicName id="4C2B4D0DD37E6463FEDEF81AAF34F87B" authority="Bergsten &amp; Ranarilalatiana &amp; Biström, 2020" authorityName="Bergsten &amp; Ranarilalatiana &amp; Biström" authorityYear="2020" box="[368,533,1958,1984]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
<emphasis id="B95FEA9CD37E6463FEDEF81AAF34F87B" box="[368,533,1958,1984]" italics="true" pageId="11" pageNumber="12">B. anjozorobe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57E7D37E6463FDB3F81AAF58F87A" box="[541,633,1959,1985]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
(triangle).
<emphasis id="B95FEA9CD37E6463FD50F81BAE32F87B" bold="true" box="[766,787,1958,1984]" pageId="11" pageNumber="12">D</emphasis>
.
<taxonomicName id="4C2B4D0DD37E6463FC8DF81AA826F87B" authority="Bistrom &amp; Sanfilippo, 1986" authorityName="Bistrom &amp; Sanfilippo" authorityYear="1986" box="[803,1287,1958,1984]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apicidens">
<emphasis id="B95FEA9CD37E6463FC8DF81AAE95F87B" box="[803,948,1958,1984]" italics="true" pageId="11" pageNumber="12">B. apicidens</emphasis>
<bibRefCitation id="EFBA4B7FD37E6463FC13F81BA826F87B" author="Bistrom O. &amp; Sanfilippo N." box="[957,1287,1958,1984]" pageId="11" pageNumber="12" pagination="46 - 47" refId="ref7377" refString="Bistrom O. &amp; Sanfilippo N. 1986. Bidessus apicidens sp. n., and faunistic notes on the genus from Africa (Coleoptera, Dytiscidae). Annales Entomologici Fennici 52: 46 - 47." type="journal article" year="1986">Biström &amp; Sanfilippo, 1986</bibRefCitation>
</taxonomicName>
(square),
<taxonomicName id="4C2B4D0DD37E6463FF13F877AF2AF85F" authority="Guignot, 1941" authorityName="Guignot" authorityYear="1941" box="[189,523,1993,2020]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ceratus">
<emphasis id="B95FEA9CD37E6463FF13F877ADF7F858" box="[189,214,1994,2019]" italics="true" pageId="11" pageNumber="12">B.</emphasis>
cf.
<emphasis id="B95FEA9CD37E6463FEADF877AC78F858" box="[259,345,1994,2019]" italics="true" pageId="11" pageNumber="12">ceratus</emphasis>
<bibRefCitation id="EFBA4B7FD37E6463FECFF874AF2AF85F" author="Guignot F." box="[353,523,1993,2020]" pageId="11" pageNumber="12" pagination="31 - 39" refId="ref7443" refString="Guignot F. 1941. Description de nouveaux Dytiscidae d'Afrique. Bulletin de la Societe d'Etude des Sciences naturelles de Vaucluse 11 (3 - 4) (1940): 31 - 39." type="journal article" year="1941">Guignot, 1941</bibRefCitation>
</taxonomicName>
(circle).
</paragraph>
</caption>
<subSubSection id="C3316505D3796464FF13FEB5A856FDBC" pageId="12" pageNumber="13" type="description">
<paragraph id="8B94368ED3796464FF13FEB5ACD5FE99" blockId="12.[189,500,264,290]" box="[189,500,264,290]" pageId="12" pageNumber="13">
<heading id="D0DC81E2D3796464FF13FEB5ACD5FE99" bold="true" box="[189,500,264,290]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B95FEA9CD3796464FF13FEB5ACD5FE99" bold="true" box="[189,500,264,290]" pageId="12" pageNumber="13">Collecting circumstances</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED3796464FF13FE8AAFDDFE2C" blockId="12.[189,1399,311,408]" pageId="12" pageNumber="13">
On
<collectingCountry id="F33C761ED3796464FF43FE8AAC5AFEEA" box="[237,379,311,337]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
found at elevations between
<quantity id="4CD39B6BD3796464FD44FE8AAEA2FEE9" box="[746,899,311,338]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.7" metricValueMin="0.9" pageId="12" pageNumber="13" unit="m" value="1300.0" valueMax="1700.0" valueMin="900.0">900–1700 m</quantity>
a.s.l. The localities are characterized by being vegetation-rich stagnant waters, a pool next to a river, a pond/marsh and small lake with floating vegetation edges. Semi-open to open landscapes.
</paragraph>
<paragraph id="8B94368ED3796464FF13FE02ACC0FE62" blockId="12.[189,481,447,473]" box="[189,481,447,473]" pageId="12" pageNumber="13">
<heading id="D0DC81E2D3796464FF13FE02ACC0FE62" bold="true" box="[189,481,447,473]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B95FEA9CD3796464FF13FE02ACC0FE62" bold="true" box="[189,481,447,473]" pageId="12" pageNumber="13">Taxonomic observation</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED3796464FF13FE50A856FDBC" blockId="12.[189,1399,493,520]" box="[189,1399,493,520]" pageId="12" pageNumber="13">
Further study on molecular level may reveal that
<taxonomicName id="4C2B4D0DD3796464FD50FE53AE58FDBC" authorityName="Gschwendtner" authorityYear="1933" box="[766,889,493,519]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nero">
<emphasis id="B95FEA9CD3796464FD50FE53AE39FDBC" box="[766,792,494,519]" italics="true" pageId="12" pageNumber="13">B.</emphasis>
cf.
<emphasis id="B95FEA9CD3796464FCEAFE53AE58FDBC" box="[836,889,494,519]" italics="true" pageId="12" pageNumber="13">nero</emphasis>
</taxonomicName>
in
<collectingCountry id="F33C761ED3796464FC30FE50A90DFDBC" box="[926,1068,493,519]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
is in fact a separate species.
</paragraph>
</subSubSection>
</treatment>
</document>
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