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<document id="97532A58D0A62265C95041C17D4C17EA" ID-CLB-Dataset="299054" ID-DOI="10.5281/zenodo.12117576" ID-GBIF-Dataset="9ec266f5-1dc7-4272-9fbc-67c7f2b0f5aa" ID-ISSN="0026-279X" ID-Zenodo-Dep="12117576" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1718741462746" checkinUser="felipe" docAuthor="Lobban, Christopher S." docDate="2015" docId="03826143FF928173FF34F920151CF9C2" docLanguage="en" docName="Micronesica.2015.02.1-49.pdf" docOrigin="Micronesica 2015 (2)" docStyle="DocumentStyle:73477DD41873F4E732E2765E250AF97D.1:Micronesica.2013-.journal_article.type1" docStyleId="73477DD41873F4E732E2765E250AF97D" docStyleName="Micronesica.2013-.journal_article.type1" docStyleVersion="1" docTitle="Pinnunavis yarrensis H. Okuno" docType="treatment" docVersion="5" lastPageNumber="11" masterDocId="FFBB193BFF9B8179FFDCFF9E1460FD3F" masterDocTitle="Benthic marine diatom flora of Guam: new records, redescription of Psammodictyon pustulatum n. comb., n. stat., and three new species (Colliculoamphora gabgabensis, Lauderia excentrica, and Rhoiconeis pagoensis)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="10" updateTime="1719541204275" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="E84B47457A2EE8813AB9DA0B1A70B4BD">Benthic marine diatom flora of Guam: new records, redescription of Psammodictyon pustulatum n. comb., n. stat., and three new species (Colliculoamphora gabgabensis, Lauderia excentrica, and Rhoiconeis pagoensis)</mods:title>
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<mods:affiliation id="29D6E34BB10B80ED516471305358B852">Division of Natural Sciences, University of Guam, Mangilao, GU 83823, USA e-mail: clobban @ guam. net</mods:affiliation>
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<treatment id="03826143FF928173FF34F920151CF9C2" ID-DOI="http://doi.org/10.5281/zenodo.12576017" ID-GBIF-Taxon="233814387" ID-Zenodo-Dep="12576017" LSID="urn:lsid:plazi:treatment:03826143FF928173FF34F920151CF9C2" httpUri="http://treatment.plazi.org/id/03826143FF928173FF34F920151CF9C2" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="C33183DEFF928170FF34F9201114FBE8" box="[232,1396,1726,1751]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B94D055FF928170FF34F9201114FBE8" blockId="9.[232,1403,1726,1886]" box="[232,1396,1726,1751]" pageId="9" pageNumber="10">
<heading id="D0DC6739FF928170FF34F9201114FBE8" box="[232,1396,1726,1751]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<taxonomicName id="4C2BABD6FF928170FF34F92016B7FBE8" authority="(Grunow) H. Okuno" authorityName="H. Okuno" baseAuthorityName="Grunow" box="[232,727,1726,1751]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnunavis" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Naviculales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="yarrensis">
<emphasis id="B95F0C47FF928170FF34F9201580FBE8" box="[232,480,1726,1751]" italics="true" pageId="9" pageNumber="10">Pinnunavis yarrensis</emphasis>
(Grunow) H. Okuno
</taxonomicName>
<figureCitation id="1310CCD0FF928170FB3EF9201114FBE8" box="[1250,1396,1726,1751]" captionStart="Figures 98106" captionStartId="39.[292,372,821,844]" captionTargetId="figure-14@40.[232,1400,352,1529]" captionTargetPageId="40" captionText="Figures 98106. Figs 98105. Pinnunavis yarrensis (GU69A-1). Fig. 98. Live cell showing plastid lobes below the valve surface (DIC). Figs 99101. Three valves of different sizes (DIC). Fig. 102. Detail of areolae and central area, external (SEM). Fig. 103. Detail of girdle view showing valvocopula with single row of pores (SEM). Fig. 104. Fragment of apex, broken along the raphe slit, showing helictoglossa and thickened interstriae (SEM). Fig. Detail of valvocopula showing fluted margin of the pars interior (SEM). Fig. 106. Plagiogramma atomus, external valve view (GU44L-E) (SEM). Scale bars: Fig 98101 = 10 µm, Figs 102106 = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.12575973" httpUri="https://zenodo.org/record/12575973/files/figure.png" pageId="9" pageNumber="10">Figs 98105</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33183DEFF928173FF34F97E151CF9C2" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="8B94D055FF928170FF34F97E1053FBC6" blockId="9.[232,1403,1726,1886]" box="[232,1075,1760,1785]" pageId="9" pageNumber="10">
Syn.:
<taxonomicName id="4C2BABD6FF928170FEF0F97E1681FBC6" authority="(Grunow) Jurilj" authorityName="Jurilj" baseAuthorityName="Grunow" box="[300,737,1760,1785]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnularia" kingdom="Chromista" order="Naviculales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="yarrensis">
<emphasis id="B95F0C47FF928170FEF0F97E1640FBC6" box="[300,544,1760,1785]" italics="true" pageId="9" pageNumber="10">Pinnularia yarrensis</emphasis>
(Grunow) Jurilj
</taxonomicName>
,
<taxonomicName id="4C2BABD6FF928170FD31F97E1053FBC6" authority="Grunow" authorityName="Grunow" authorityYear="1876" box="[749,1075,1760,1785]" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" order="Naviculales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="yarrensis">
<emphasis id="B95F0C47FF928170FD31F97E17ACFBC6" box="[749,972,1760,1785]" italics="true" pageId="9" pageNumber="10">Navicula yarrensis</emphasis>
Grunow
</taxonomicName>
</paragraph>
<paragraph id="8B94D055FF928170FF34F89C1648FA24" blockId="9.[232,1403,1726,1886]" box="[232,552,1794,1819]" pageId="9" pageNumber="10">
<taxonomicName id="4C2BABD6FF928170FF34F89C1517FA24" box="[232,375,1794,1819]" class="Bacillariophyceae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="order">Naviculales</taxonomicName>
:
<taxonomicName id="4C2BABD6FF928170FE5EF89C1644FA24" box="[386,548,1794,1819]" class="Bacillariophyceae" family="Naviculaceae" kingdom="Chromista" order="Naviculales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="family">Naviculaceae</taxonomicName>
.
</paragraph>
<paragraph id="8B94D055FF928173FF34F8BA170FFC14" blockId="9.[232,1403,1726,1886]" lastBlockId="10.[232,1403,240,1277]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Ref. illus.: Grunow in Schmidt et al. 18741959, pl. 46, figs 16;
<bibRefCitation id="EFBAADA4FF928170FBD2F8BA10CFFA02" author="Meister, F." box="[1038,1199,1828,1853]" pageId="9" pageNumber="10" refId="ref16415" refString="Meister, F. 1932. Kieselalgen aus Asien. Gebruder Borntraeger, Berlin." type="book" year="1932">Meister 1932</bibRefCitation>
, pl. 17, fig. 139;
<bibRefCitation id="EFBAADA4FF928170FF34F8DB15E8FA61" author="Hustedt, F." box="[232,392,1861,1886]" pageId="9" pageNumber="10" refId="ref15381" refString="Hustedt, F. 1955. Marine Littoral Diatoms of Beaufort, North Carolina. Duke University Press. Durham, North Carolina. 67 pp." type="book" year="1955">Hustedt 1955</bibRefCitation>
, pl. 9, fig. 2;
<bibRefCitation id="EFBAADA4FF928170FDF8F8DB16B5FA61" author="Navarro, J. N." box="[548,725,1861,1886]" pageId="9" pageNumber="10" pagination="119 - 136" refId="ref16583" refString="Navarro, J. N. 1983 a. A survey of the marine diatoms of Puerto Rico. VI. Suborder Raphidineae: Family Naviculaceae (Genera Haslea, Mastogloia and Navicula). Botanica Marina 26: 119 - 136." type="journal article" year="1983">Navarro 1983a</bibRefCitation>
, figs 108110;
<bibRefCitation id="EFBAADA4FF928170FC50F8DB1003FA61" author="Wah, T. T. &amp; Y. C. Wee" box="[908,1123,1861,1886]" pageId="9" pageNumber="10" pagination="317 - 327" refId="ref18131" refString="Wah, T. T. &amp; Y. C. Wee. 1988. Diatoms from mangrove environments of Singapore and southern peninsular Malaysia. Botanica Marina 31: 317 - 327." type="journal article" year="1988">Wah &amp; Wee 1988</bibRefCitation>
, fig. 64 (all as
<taxonomicName id="4C2BABD6FF928173FACDF8DB1535FC36" authorityName="Grunow" authorityYear="1876" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" lastPageId="10" lastPageNumber="11" order="Naviculales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="yarrensis">
<emphasis id="B95F0C47FF928173FACDF8DB1535FC36" italics="true" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">Navicula yarrensis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFBAADA4FF918173FEACFF6E161AFC36" author="Witkowski A. &amp; H. Lange-Bertalot &amp; D. Metzeltin" box="[368,634,240,265]" pageId="10" pageNumber="11" refId="ref18313" refString="Witkowski A., H. Lange-Bertalot &amp; D. Metzeltin. 2000. Diatom Flora of Marine Coasts, I. Gantner Verlag, Ruggell, Germany." type="book" year="2000">Witkowski et al. 2000</bibRefCitation>
, pl. 146, fig. 11; López-Fuerte et al. 2010, pl. 29, figs 1, 2 (both as
<taxonomicName id="4C2BABD6FF918173FED4FE8C159BFC14" box="[264,507,274,299]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnularia" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="yarrensis">
<emphasis id="B95F0C47FF918173FED4FE8C159BFC14" box="[264,507,274,299]" italics="true" pageId="10" pageNumber="11">Pinnularia yarrensis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFBAADA4FF918173FDC0FE8C16D7FC14" author="Okuno, H." box="[540,695,274,299]" pageId="10" pageNumber="11" pagination="97 - 113" refId="ref16769" refString="Okuno, H. 1975. The fine structure of the frustules of the Bacillariophyta. In J. Tokida &amp; H. Hirose (eds), Advance of Phycology in Japan. Junk. The Hague. pp. 97 - 113." type="book chapter" year="1975">Okuno, 1975</bibRefCitation>
, fig. 8-2 and 2a
</paragraph>
<paragraph id="8B94D055FF918173FF34FEAD15AFFC73" blockId="10.[232,1403,240,1277]" box="[232,463,307,332]" pageId="10" pageNumber="11">Samples: GU69A-1</paragraph>
<paragraph id="8B94D055FF918173FF34FECB1068FC51" blockId="10.[232,1403,240,1277]" box="[232,1032,341,366]" pageId="10" pageNumber="11">Dimensions: 36105 µm long, 1532 µm wide, 47 striae in 10 µm.</paragraph>
<paragraph id="8B94D055FF918173FF34FEE91749FF28" blockId="10.[232,1403,240,1277]" pageId="10" pageNumber="11">
Diagnostics: Striae radiate along most of the valve but convergent at the apices. Striae consisting of single alveolus with numerous small poroids in the outer membrane and no inner membrane (
<figureCitation id="1310CCD0FF918173FF2CFE2415F9FCEC" box="[240,409,442,467]" captionStart="Figures 98106" captionStartId="39.[292,372,821,844]" captionTargetId="figure-14@40.[232,1400,352,1529]" captionTargetPageId="40" captionText="Figures 98106. Figs 98105. Pinnunavis yarrensis (GU69A-1). Fig. 98. Live cell showing plastid lobes below the valve surface (DIC). Figs 99101. Three valves of different sizes (DIC). Fig. 102. Detail of areolae and central area, external (SEM). Fig. 103. Detail of girdle view showing valvocopula with single row of pores (SEM). Fig. 104. Fragment of apex, broken along the raphe slit, showing helictoglossa and thickened interstriae (SEM). Fig. Detail of valvocopula showing fluted margin of the pars interior (SEM). Fig. 106. Plagiogramma atomus, external valve view (GU44L-E) (SEM). Scale bars: Fig 98101 = 10 µm, Figs 102106 = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.12575973" httpUri="https://zenodo.org/record/12575973/files/figure.png" pageId="10" pageNumber="11">Figs 102104</figureCitation>
). The two plastids (
<figureCitation id="1310CCD0FF918173FD44FE241690FCEC" box="[664,752,442,467]" captionStart="Figures 98106" captionStartId="39.[292,372,821,844]" captionTargetId="figure-14@40.[232,1400,352,1529]" captionTargetPageId="40" captionText="Figures 98106. Figs 98105. Pinnunavis yarrensis (GU69A-1). Fig. 98. Live cell showing plastid lobes below the valve surface (DIC). Figs 99101. Three valves of different sizes (DIC). Fig. 102. Detail of areolae and central area, external (SEM). Fig. 103. Detail of girdle view showing valvocopula with single row of pores (SEM). Fig. 104. Fragment of apex, broken along the raphe slit, showing helictoglossa and thickened interstriae (SEM). Fig. Detail of valvocopula showing fluted margin of the pars interior (SEM). Fig. 106. Plagiogramma atomus, external valve view (GU44L-E) (SEM). Scale bars: Fig 98101 = 10 µm, Figs 102106 = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.12575973" httpUri="https://zenodo.org/record/12575973/files/figure.png" pageId="10" pageNumber="11">Fig. 98</figureCitation>
) extend onto the valve face nearly to the raphe, with closely lobed margins. Valve shape varied from elliptical-lanceolate to linear with broadly rounded apices; size also varied widely in this population.
</paragraph>
<paragraph id="8B94D055FF918173FF34FD811099FF43" blockId="10.[232,1403,240,1277]" pageId="10" pageNumber="11">
Comments:
<collectionCode id="ED3A4890FF918173FEA4FD8115D4FF07" box="[376,436,543,568]" pageId="10" pageNumber="11">SEM</collectionCode>
views confirm the images shown by
<bibRefCitation id="EFBAADA4FF918173FCA8FD81105CFF07" author="Navarro, J. N." box="[884,1084,543,568]" pageId="10" pageNumber="11" pagination="119 - 136" refId="ref16583" refString="Navarro, J. N. 1983 a. A survey of the marine diatoms of Puerto Rico. VI. Suborder Raphidineae: Family Naviculaceae (Genera Haslea, Mastogloia and Navicula). Botanica Marina 26: 119 - 136." type="journal article" year="1983">Navarro (1983a)</bibRefCitation>
. I also noted that the inner margin of the valvocopula is scalloped to fit against the outer edges of the interstriae (
<figureCitation id="1310CCD0FF918173FAD8FDDF110EFF65" box="[1284,1390,577,602]" captionStart="Figures 98106" captionStartId="39.[292,372,821,844]" captionTargetId="figure-14@40.[232,1400,352,1529]" captionTargetPageId="40" captionText="Figures 98106. Figs 98105. Pinnunavis yarrensis (GU69A-1). Fig. 98. Live cell showing plastid lobes below the valve surface (DIC). Figs 99101. Three valves of different sizes (DIC). Fig. 102. Detail of areolae and central area, external (SEM). Fig. 103. Detail of girdle view showing valvocopula with single row of pores (SEM). Fig. 104. Fragment of apex, broken along the raphe slit, showing helictoglossa and thickened interstriae (SEM). Fig. Detail of valvocopula showing fluted margin of the pars interior (SEM). Fig. 106. Plagiogramma atomus, external valve view (GU44L-E) (SEM). Scale bars: Fig 98101 = 10 µm, Figs 102106 = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.12575973" httpUri="https://zenodo.org/record/12575973/files/figure.png" pageId="10" pageNumber="11">Fig. 105</figureCitation>
). There is a single line of pores near the abvalvar edge of the wide valvocopula (
<figureCitation id="1310CCD0FF918173FB59FDFD108CFF43" box="[1157,1260,611,636]" captionStart="Figures 98106" captionStartId="39.[292,372,821,844]" captionTargetId="figure-14@40.[232,1400,352,1529]" captionTargetPageId="40" captionText="Figures 98106. Figs 98105. Pinnunavis yarrensis (GU69A-1). Fig. 98. Live cell showing plastid lobes below the valve surface (DIC). Figs 99101. Three valves of different sizes (DIC). Fig. 102. Detail of areolae and central area, external (SEM). Fig. 103. Detail of girdle view showing valvocopula with single row of pores (SEM). Fig. 104. Fragment of apex, broken along the raphe slit, showing helictoglossa and thickened interstriae (SEM). Fig. Detail of valvocopula showing fluted margin of the pars interior (SEM). Fig. 106. Plagiogramma atomus, external valve view (GU44L-E) (SEM). Scale bars: Fig 98101 = 10 µm, Figs 102106 = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.12575973" httpUri="https://zenodo.org/record/12575973/files/figure.png" pageId="10" pageNumber="11">Fig. 103</figureCitation>
).
</paragraph>
<paragraph id="8B94D055FF918173FE94FD1A151CF9C2" blockId="10.[232,1403,240,1277]" pageId="10" pageNumber="11">
A “striking species of massive sculpture” (
<bibRefCitation id="EFBAADA4FF918173FC97FD1A106DFFA2" author="Mann, A." box="[843,1037,644,669]" pageId="10" pageNumber="11" pagination="1 - 182" refId="ref16276" refString="Mann, A. 1925. Marine diatoms of the Philippine Islands. Smithsonian Institution Bulletin 100, 6 (1): 1 - 182 + 39 pl." type="journal article" year="1925">Mann 1925:125</bibRefCitation>
), and equally handsome alive. This species is usually noted as widespread but scarce (
<bibRefCitation id="EFBAADA4FF918173FC1EFD3810F2FF80" author="Hustedt, F." box="[962,1170,678,703]" pageId="10" pageNumber="11" refId="ref15381" refString="Hustedt, F. 1955. Marine Littoral Diatoms of Beaufort, North Carolina. Duke University Press. Durham, North Carolina. 67 pp." type="book" year="1955">Hustedt 1955:32</bibRefCitation>
;
<bibRefCitation id="EFBAADA4FF918173FB74FD38153AFFDE" author="Witkowski A. &amp; H. Lange-Bertalot &amp; D. Metzeltin" pageId="10" pageNumber="11" refId="ref18313" refString="Witkowski A., H. Lange-Bertalot &amp; D. Metzeltin. 2000. Diatom Flora of Marine Coasts, I. Gantner Verlag, Ruggell, Germany." type="book" year="2000">Witkowski et al. 2000:338</bibRefCitation>
), but was abundant in GU69A-1 from surface sediment at the edge of a mangrove.
<bibRefCitation id="EFBAADA4FF918173FF34FD7415D6FE3C" author="Navarro, J. N." box="[232,438,746,771]" pageId="10" pageNumber="11" pagination="1 - 151" refId="ref16503" refString="Navarro, J. N. 1982 a. Marine diatoms associated with mangrove prop roots in the Indian River, Florida, U. S. A. Bibliotheca Phycologica 61: 1 - 151." type="journal article" year="1982">Navarro (1982a)</bibRefCitation>
,
<bibRefCitation id="EFBAADA4FF918173FE16FD7416AAFE3C" author="Wah, T. T. &amp; Y. C. Wee" box="[458,714,746,771]" pageId="10" pageNumber="11" pagination="317 - 327" refId="ref18131" refString="Wah, T. T. &amp; Y. C. Wee. 1988. Diatoms from mangrove environments of Singapore and southern peninsular Malaysia. Botanica Marina 31: 317 - 327." type="journal article" year="1988">Wah &amp; Wee (1988)</bibRefCitation>
and López-Fuerte et al. (2010) also reported it from mangroves.
<bibRefCitation id="EFBAADA4FF918173FEA0FC951636FE1B" author="Meister, F." box="[380,598,779,804]" pageId="10" pageNumber="11" refId="ref16415" refString="Meister, F. 1932. Kieselalgen aus Asien. Gebruder Borntraeger, Berlin." type="book" year="1932">Meister (1932:43)</bibRefCitation>
reported linear valves of the nominate
<taxonomicName id="4C2BABD6FF918173FB94FC9510BFFE1B" box="[1096,1247,779,804]" pageId="10" pageNumber="11" rank="variety" variety="from">variety from</taxonomicName>
<collectingRegion id="49EF1EB7FF918173FB37FC951157FE1B" box="[1259,1335,779,804]" country="Philippines" name="Cavite" pageId="10" pageNumber="11">Cavite</collectingRegion>
Bay, Luzon,
<collectingCountry id="F33C90C5FF918173FE9FFCB315A7FE79" box="[323,455,813,838]" name="Philippines" pageId="10" pageNumber="11">Philippines</collectingCountry>
(habitat not given). The number of striae reported varies from 33.5 in 10 µm (
<bibRefCitation id="EFBAADA4FF918173FF33FCD115F0FE57" author="Meister, F." box="[239,400,847,872]" pageId="10" pageNumber="11" refId="ref16415" refString="Meister, F. 1932. Kieselalgen aus Asien. Gebruder Borntraeger, Berlin." type="book" year="1932">Meister 1932</bibRefCitation>
) and
<quantity id="4CD37DB0FF918173FE0AFCD11665FE57" box="[470,517,847,872]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="11" unit="in" value="4.0">4 in</quantity>
10 µm (
<bibRefCitation id="EFBAADA4FF918173FDB2FCD1171DFE57" author="Witkowski A. &amp; H. Lange-Bertalot &amp; D. Metzeltin" box="[622,893,847,872]" pageId="10" pageNumber="11" refId="ref18313" refString="Witkowski A., H. Lange-Bertalot &amp; D. Metzeltin. 2000. Diatom Flora of Marine Coasts, I. Gantner Verlag, Ruggell, Germany." type="book" year="2000">Witkowski et al. 2000</bibRefCitation>
), to
<quantity id="4CD37DB0FF918173FC6FFCD11791FE57" box="[947,1009,847,872]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="11" unit="in" value="10.0">10 in</quantity>
10 µm (
<bibRefCitation id="EFBAADA4FF918173FB86FCD11547FEB5" author="Podzorski, A. C. &amp; H. Hakansson" pageId="10" pageNumber="11" pagination="1 - 245" refId="ref17030" refString="Podzorski, A. C. &amp; H. Hakansson. 1987. Freshwater and marine diatoms from Palawan (a Philippine island). Bibliotheca Diatomologica 13: 1 - 245. J. Cramer, Berlin." type="journal article" year="1987">Podzorski &amp; Håkansson 1987</bibRefCitation>
); the more finely striate forms are not so readily distinguished from
<emphasis id="B95F0C47FF918173FB8BFCEF10A9FEB5" box="[1111,1225,881,906]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C2BABD6FF918173FB8BFCEF10A5FEB5" box="[1111,1221,881,906]" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="genus">Navicula</taxonomicName>
.
</emphasis>
In my sample, the smaller valves (
<figureCitation id="1310CCD0FF918173FE11FC0C1652FE94" box="[461,562,914,939]" captionStart="Figures 98106" captionStartId="39.[292,372,821,844]" captionTargetId="figure-14@40.[232,1400,352,1529]" captionTargetPageId="40" captionText="Figures 98106. Figs 98105. Pinnunavis yarrensis (GU69A-1). Fig. 98. Live cell showing plastid lobes below the valve surface (DIC). Figs 99101. Three valves of different sizes (DIC). Fig. 102. Detail of areolae and central area, external (SEM). Fig. 103. Detail of girdle view showing valvocopula with single row of pores (SEM). Fig. 104. Fragment of apex, broken along the raphe slit, showing helictoglossa and thickened interstriae (SEM). Fig. Detail of valvocopula showing fluted margin of the pars interior (SEM). Fig. 106. Plagiogramma atomus, external valve view (GU44L-E) (SEM). Scale bars: Fig 98101 = 10 µm, Figs 102106 = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.12575973" httpUri="https://zenodo.org/record/12575973/files/figure.png" pageId="10" pageNumber="11">Fig. 104</figureCitation>
) had denser striae but appear to still be the same species. Length range in the literature is 60200 µm (
<bibRefCitation id="EFBAADA4FF918173FD84FC2A1705FEF2" author="Witkowski A. &amp; H. Lange-Bertalot &amp; D. Metzeltin" box="[600,869,948,973]" pageId="10" pageNumber="11" refId="ref18313" refString="Witkowski A., H. Lange-Bertalot &amp; D. Metzeltin. 2000. Diatom Flora of Marine Coasts, I. Gantner Verlag, Ruggell, Germany." type="book" year="2000">Witkowski et al. 2000</bibRefCitation>
); the smallest valve I found was 36 µm long. Several varieties have been named, including three from Miocene deposits (
<bibRefCitation id="EFBAADA4FF918173FBB7FC481097FED0" author="Hajos, M." box="[1131,1271,982,1007]" pageId="10" pageNumber="11" pagination="1 - 401" refId="ref14889" refString="Hajos, M. 1968 Matraalja Miocen Uledekeinek Diatomai [Die diatomeen der Miozanen Ablagerungen des Matravorlandes]. Geologica Hungarica, Series Paleontologica, Institutum Geologicum Hungaricum, Budapestini 37: 1 - 401." type="journal article" year="1968">Hajós 1968</bibRefCitation>
;
<bibRefCitation id="EFBAADA4FF918173FAD8FC481657F92E" author="California Academy of Sciences" pageId="10" pageNumber="11" refId="ref14156" refString="California Academy of Sciences. 2011. Catalogue of diatom names, on-line version updated 19 Sept. 2011. Compiled by Elisabeth Fourtanier &amp; J. Patrick Kociolek. Available online at http: // research. calacademy. org / research / diatoms / names / index. asp. Accessed 22 June 2014." type="url" year="2011">California Academy of Sciences 2011</bibRefCitation>
).
<taxonomicName id="4C2BABD6FF918173FD93FC661751F92E" authorityName="Grunow" authorityYear="1876" box="[591,817,1016,1041]" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="yarrensis">
<emphasis id="B95F0C47FF918173FD93FC661751F92E" box="[591,817,1016,1041]" italics="true" pageId="10" pageNumber="11">Navicula yarrensis</emphasis>
</taxonomicName>
was first transferred to
<taxonomicName id="4C2BABD6FF918173FB87FC66111AF92E" authority="(Jurilj 1957)" baseAuthorityName="Jurilj" baseAuthorityYear="1957" box="[1115,1402,1016,1041]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnularia" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF918173FB87FC6610BAF92E" box="[1115,1242,1016,1041]" italics="true" pageId="10" pageNumber="11">Pinnularia</emphasis>
(
<bibRefCitation id="EFBAADA4FF918173FB31FC661112F92E" author="Jurilj, A." box="[1261,1394,1016,1041]" pageId="10" pageNumber="11" pagination="5 - 153" refId="ref15487" refString="Jurilj, A. 1957. Dijatomeje samatskog mora okoline Zagreba. (Flora of Diatoms of Sarmatic Sea in Environs of Zagreb). Jugoslavenska Akademija Znanosti i Umjetnosi, Zagreb. (Acta Biologica 1) 28: 5 - 153, 40 pls." type="book chapter" year="1957">Jurilj 1957</bibRefCitation>
)
</taxonomicName>
and later to a new genus,
<taxonomicName id="4C2BABD6FF918173FDC4FB84172DF90C" authority="Okuno (1975)" authorityName="Okuno" authorityYear="1975" box="[536,845,1050,1075]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnunavis" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF918173FDC4FB8416FCF90C" box="[536,668,1050,1075]" italics="true" pageId="10" pageNumber="11">Pinnunavis</emphasis>
<bibRefCitation id="EFBAADA4FF918173FD79FB84172DF90C" author="Okuno, H." box="[677,845,1050,1075]" pageId="10" pageNumber="11" pagination="97 - 113" refId="ref16769" refString="Okuno, H. 1975. The fine structure of the frustules of the Bacillariophyta. In J. Tokida &amp; H. Hirose (eds), Advance of Phycology in Japan. Junk. The Hague. pp. 97 - 113." type="book chapter" year="1975">Okuno (1975)</bibRefCitation>
</taxonomicName>
, along with
<taxonomicName id="4C2BABD6FF918173FC3CFB841124F90C" authority="(W. Smith) Okuno" authorityName="Okuno" authorityYear="1975" baseAuthorityName="W. Smith" box="[992,1348,1050,1075]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnunavis" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="elegans">
<emphasis id="B95F0C47FF918173FC3CFB84103BF90C" box="[992,1115,1050,1075]" italics="true" pageId="10" pageNumber="11">P. elegans</emphasis>
(W. Smith) Okuno
</taxonomicName>
, and this is the currently accepted name for
<taxonomicName id="4C2BABD6FF918173FD69FBA51723F96B" authorityName="Grunow" authorityYear="1876" box="[693,835,1083,1108]" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="yarrensis">
<emphasis id="B95F0C47FF918173FD69FBA51723F96B" box="[693,835,1083,1108]" italics="true" pageId="10" pageNumber="11">P. yarrensis</emphasis>
</taxonomicName>
according to AlgaeBase (
<bibRefCitation id="EFBAADA4FF918173FBA9FBA5110FF96B" author="Guiry, M. D. &amp; G. M. Guiry" box="[1141,1391,1083,1108]" pageId="10" pageNumber="11" refId="ref14752" refString="Guiry, M. D. &amp; G. M. Guiry 2014. AlgaeBase. World-wide electronic publication, National University of Ireland, Galway. http: // www. algaebase. org; accessed 22 Dec. 2013." type="url" year="2014">Guiry &amp; Guiry 2014</bibRefCitation>
). However,
<taxonomicName id="4C2BABD6FF918173FEB8FBC31753F949" authority="(W. Smith) Okuno" authorityName="Okuno" authorityYear="1975" baseAuthorityName="W. Smith" box="[356,819,1117,1142]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnunavis" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="elegans">
<emphasis id="B95F0C47FF918173FEB8FBC3162AF949" box="[356,586,1117,1142]" italics="true" pageId="10" pageNumber="11">Pinnunavis elegans</emphasis>
(W. Smith) Okuno
</taxonomicName>
was transferred back to
<taxonomicName id="4C2BABD6FF918173FB80FBC310BBF949" baseAuthorityName="Jurilj" baseAuthorityYear="1957" box="[1116,1243,1117,1142]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnularia" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF918173FB80FBC310BBF949" box="[1116,1243,1117,1142]" italics="true" pageId="10" pageNumber="11">Pinnularia</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFBAADA4FF918173FAD7FBC31559F9A7" author="Krammer, K." pageId="10" pageNumber="11" pagination="1 - 353" refId="ref15534" refString="Krammer, K. 1992. Pinnularia. Eine Monographie der europaischen Taxa. Bibliotheca Diatomologica. 26: 1 - 353. J. Cramer. Berlin-Stuttgart." type="journal article" year="1992">Krammer (1992)</bibRefCitation>
. The plastids of
<taxonomicName id="4C2BABD6FF918173FE27FBE11614F9A7" authorityName="Okuno" authorityYear="1975" baseAuthorityName="W. Smith" box="[507,628,1151,1176]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnunavis" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="elegans">
<emphasis id="B95F0C47FF918173FE27FBE11614F9A7" box="[507,628,1151,1176]" italics="true" pageId="10" pageNumber="11">P. elegans</emphasis>
</taxonomicName>
lie along the girdle bands, typical of
<taxonomicName id="4C2BABD6FF918173FBF7FBE110F5F9A7" box="[1067,1173,1151,1176]" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF918173FBF7FBE110F5F9A7" box="[1067,1173,1151,1176]" italics="true" pageId="10" pageNumber="11">Navicula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2BABD6FF918173FB12FBE1112DF9A7" baseAuthorityName="Jurilj" baseAuthorityYear="1957" box="[1230,1357,1151,1176]" class="Bacillariophyceae" family="Naviculaceae" genus="Pinnularia" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF918173FB12FBE1112DF9A7" box="[1230,1357,1151,1176]" italics="true" pageId="10" pageNumber="11">Pinnularia</emphasis>
</taxonomicName>
(A. Witkowski, personal communication), whereas those seen here for
<taxonomicName id="4C2BABD6FF918173FBCEFB3F10C3F985" authorityName="Grunow" authorityYear="1876" box="[1042,1187,1185,1210]" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="yarrensis">
<emphasis id="B95F0C47FF918173FBCEFB3F10C3F985" box="[1042,1187,1185,1210]" italics="true" pageId="10" pageNumber="11">P. yarrensis</emphasis>
</taxonomicName>
spread across the valve face, more like those of some
<taxonomicName id="4C2BABD6FF918173FD42FB5C1770F9E4" authorityName="P.T.Cleve" authorityYear="1894" box="[670,784,1218,1243]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Naviculales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF918173FD42FB5C1770F9E4" box="[670,784,1218,1243]" italics="true" pageId="10" pageNumber="11">Diploneis</emphasis>
</taxonomicName>
. It remains to be seen how these genera are related molecularly.
</paragraph>
</subSubSection>
</treatment>
</document>