treatments-xml/data/03/82/34/0382346DFF84FFE7A8C9FDF81C058EBF.xml

406 lines
63 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="C869A8AEDE63285944FF3EA473D34EFE" ID-DOI="10.11646/zootaxa.5476.1.31" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681863" ID-ZooBank="B3C36A64-00AF-4D48-B2B3-74AFE9793CB0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720375684251" checkinUser="plazi" docAuthor="Takeda, Masatsune &amp; Komatsu, Hironori" docDate="2024" docId="0382346DFF84FFE7A8C9FDF81C058EBF" docLanguage="en" docName="zootaxa.5476.1.31.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.31" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Actumnus ngankeeae Takeda &amp; Komatsu, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="428" masterDocId="FFBB4C15FF85FFE3A85EFFC41D438F6E" masterDocTitle="Actumnus ngankeeae sp. nov. and Pilumnus swajayai Ng &amp; Rahayu, 2021, pilumnid crabs from Japan (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="433" masterPageNumber="424" pageNumber="425" updateTime="1720379351963" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A6DC4298577421915FBBAC6E423E1BF2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AA036C202FC6CB073EF7ABAE90B8237C">
<mods:title id="FB8458B2B4636B67E2D1C21128EE0010">Actumnus ngankeeae sp. nov. and Pilumnus swajayai Ng &amp; Rahayu, 2021, pilumnid crabs from Japan (Crustacea: Decapoda: Brachyura)</mods:title>
</mods:titleInfo>
<mods:name id="058639D2C344CD0A10728823EAE17AAB" type="personal">
<mods:role id="8DF062C9D64719D4785CA08339BDB076">
<mods:roleTerm id="6AC119F2D6AA77322C1DF61EA571AB4E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B5BDF8379728B6672DB7469EE2EC684">Takeda, Masatsune</mods:namePart>
<mods:nameIdentifier id="73DB59DEC865577BC16F18C155FB5F4E" type="ORCID">0000-0002-0028-1397</mods:nameIdentifier>
<mods:affiliation id="D0D947C186881B80B80E3070B401FEAC">Department of Zoology, National Museum of Nature and Science, Tokyo, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
<mods:nameIdentifier id="FB26D0E852424CF58CBCD2F9E44860C1" type="email">takeda@kahaku.go.jp</mods:nameIdentifier>
</mods:name>
<mods:name id="BC6B4DE9C29A018936F32CB0FDF67BEB" type="personal">
<mods:role id="A6BC2868402A92850656F9C03158F00F">
<mods:roleTerm id="7A73C4F205C0A78C0B49A6164E4414E3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E701800987CF4993DDC1E2EABA93343E">Komatsu, Hironori</mods:namePart>
<mods:nameIdentifier id="7C75A83C7D1D1930DBB726CC0FCA5B9A" type="ORCID">0000-0002-7573-4127</mods:nameIdentifier>
<mods:affiliation id="C4EA075DE650459CEE007970AE319667">Department of Zoology, National Museum of Nature and Science, Tokyo, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
<mods:nameIdentifier id="862BFD20BD596C1C3F7E20AB52EC50D9" type="email">h-komatu@kahaku.go.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1D2065B968CA199F17EF404525D31942">text</mods:typeOfResource>
<mods:relatedItem id="BDF58750AA9542BC38C042AEC327F434" type="host">
<mods:titleInfo id="24F5094F84E1A877AFB7ECA77C52FDE7">
<mods:title id="ED9488F07AC9CA661059526B6A5A21B6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="562AB68DA644B9196A5480FBF3BE46A2">
<mods:date id="B0400781579CD5EE2057928A16C56E2B">2024</mods:date>
<mods:detail id="1E90F0C656154FAEF82E112B209BE8F8" type="pubDate">
<mods:number id="8B0A4E7E64B5F59159F5FF4A78B4F316">2024-07-04</mods:number>
</mods:detail>
<mods:detail id="F07715176158CC98B9C79988CF46BFCA" type="volume">
<mods:number id="2C9BA8CC2F74AD4C9439EA024CE73B43">5476</mods:number>
</mods:detail>
<mods:detail id="90F04096A342125A3B7E912C58948E9D" type="issue">
<mods:number id="514389E8CD262A97F01ABCA928AAC106">1</mods:number>
</mods:detail>
<mods:extent id="C0EB5AC058903363656220E89AA6A704" unit="page">
<mods:start id="08589FDB2CB11A7FDF46EE60BCA1AF14">424</mods:start>
<mods:end id="9E22DD2F20C168BD8C1F27AF0AEE0134">433</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="57F086EA60AA812A9A737204A347D509">
<mods:url id="58D531931251529DCFF97589966AD8B4">http://dx.doi.org/10.11646/zootaxa.5476.1.31</mods:url>
</mods:location>
<mods:classification id="7BEB181AE6E6B3B9144527C01024008B">journal article</mods:classification>
<mods:identifier id="1B66BEEE86DEC3DE76E1E60645131E39" type="DOI">10.11646/zootaxa.5476.1.31</mods:identifier>
<mods:identifier id="3D538B673281D594B5CD1AA6F4B42C16" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="40BFFF148446995C2E98433BD57C4F9F" type="Zenodo-Dep">12681863</mods:identifier>
<mods:identifier id="1F0AE6CAFD4E7A14719693487006346A" type="ZooBank">B3C36A64-00AF-4D48-B2B3-74AFE9793CB0</mods:identifier>
</mods:mods>
<treatment id="0382346DFF84FFE7A8C9FDF81C058EBF" ID-DOI="http://doi.org/10.5281/zenodo.12681406" ID-Zenodo-Dep="12681406" LSID="urn:lsid:plazi:treatment:0382346DFF84FFE7A8C9FDF81C058EBF" httpUri="http://treatment.plazi.org/id/0382346DFF84FFE7A8C9FDF81C058EBF" lastPageId="4" lastPageNumber="428" pageId="1" pageNumber="425">
<subSubSection id="C331D6F0FF84FFE2A8C9FDF81F418D39" box="[151,514,572,599]" pageId="1" pageNumber="425" type="nomenclature">
<paragraph id="8B94857BFF84FFE2A8C9FDF81F418D39" blockId="1.[151,514,572,635]" box="[151,514,572,599]" pageId="1" pageNumber="425">
<heading id="D0DC3217FF84FFE2A8C9FDF81F418D39" bold="true" box="[151,514,572,599]" fontSize="11" level="1" pageId="1" pageNumber="425" reason="1">
<emphasis id="B95F5969FF84FFE2A8C9FDF81F418D39" bold="true" box="[151,514,572,599]" pageId="1" pageNumber="425">
<taxonomicName id="4C2BFEF8FF84FFE2A8C9FDF81CDF8D39" authorityName="Takeda &amp; Komatsu" authorityYear="2024" box="[151,412,572,599]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="425" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="B95F5969FF84FFE2A8C9FDF81CDF8D39" bold="true" box="[151,412,572,599]" italics="true" pageId="1" pageNumber="425">Actumnus ngankeeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CE412FF84FFE2A9FAFDF91F418D39" box="[420,514,573,599]" pageId="1" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331D6F0FF84FFE2A8C9FDA41C3E8D15" box="[151,381,608,635]" pageId="1" pageNumber="425" type="description">
<paragraph id="8B94857BFF84FFE2A8C9FDA41C3E8D15" blockId="1.[151,514,572,635]" box="[151,381,608,635]" pageId="1" pageNumber="425">
(
<figureCitation id="131099FEFF84FFE2A8C1FDA41DAE8D15" box="[159,237,608,635]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Figs. 1</figureCitation>
,
<figureCitation id="131099FEFF84FFE2A8A5FDA51C488D15" box="[251,267,609,635]" captionStart="FIGURE 2" captionStartId="3.[151,250,709,733]" captionTargetBox="[151,1436,181,686]" captionTargetId="figure-20@3.[151,1436,181,686]" captionTargetPageId="3" captionText="FIGURE 2. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, chelae, outer view; B, pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681869" httpUri="https://zenodo.org/record/12681869/files/figure.png" pageId="1" pageNumber="425">2</figureCitation>
,
<figureCitation id="131099FEFF84FFE2A949FDA51C7A8D15" box="[279,313,609,635]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. AC, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng &amp; Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; CD, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="1" pageNumber="425">3A</figureCitation>
⎼C, 4)
</paragraph>
</subSubSection>
<subSubSection id="C331D6F0FF84FFE2A8C9FD6D181E8D89" pageId="1" pageNumber="425" type="materials_examined">
<paragraph id="8B94857BFF84FFE2A8C9FD6D181E8D89" blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="425">
<emphasis id="B95F5969FF84FFE2A8C9FD6D1C018DAC" bold="true" box="[151,322,680,707]" pageId="1" pageNumber="425">Type material</emphasis>
.
<materialsCitation id="3B438F26FF84FFE2A90AFD6C181E8D89" collectingDate="2015-07-04" collectorName="H. Komatsu" latitude="26.588612" location="Haha-jima Island" longLatPrecision="20" longitude="142.14471" municipality="Ogasawara Islands" pageId="1" pageNumber="425" specimenCount="1" typeStatus="holotype">
<emphasis id="B95F5969FF84FFE2A90AFD6C1CF88DAC" box="[340,443,680,706]" italics="true" pageId="1" pageNumber="425">
<typeStatus id="54903BD9FF84FFE2A90AFD6C1CF88DAC" box="[340,443,680,706]" pageId="1" pageNumber="425" type="holotype">Holotype</typeStatus>
</emphasis>
: male (cb 4.6 mm, cl 3.5 mm), NSMT-Cr 31014, diving site
<emphasis id="B95F5969FF84FFE2ACCAFD6C19BC8DAC" box="[1172,1279,680,706]" italics="true" pageId="1" pageNumber="425">Zakuzaku</emphasis>
,
<geoCoordinate id="EE1FE3BCFF84FFE2AD4FFD6C18D48DAD" box="[1297,1431,680,707]" degrees="26" direction="north" minutes="35" orientation="latitude" pageId="1" pageNumber="425" precision="15" seconds="19" value="26.588612">26°35'19&quot;N</geoCoordinate>
,
<geoCoordinate id="EE1FE3BCFF84FFE2A8C9FD091C6B8D89" box="[151,296,717,743]" degrees="142" direction="east" minutes="08" orientation="longitude" pageId="1" pageNumber="425" precision="15" seconds="41" value="142.14471">142°08'41&quot;E</geoCoordinate>
, south of
<location id="8EF4D3A0FF84FFE2A9C8FD081F198D88" LSID="urn:lsid:plazi:treatment:0382346DFF84FFE7A8C9FDF81C058EBF:8EF4D3A0FF84FFE2A9C8FD081F198D88" box="[406,602,716,742]" latitude="26.588612" longLatPrecision="20" longitude="142.14471" municipality="Ogasawara Islands" name="Haha-jima Island" pageId="1" pageNumber="425">Haha-jima Island</location>
,
<collectingMunicipality id="6BF01F01FF84FFE2AA38FD091E788D88" box="[614,827,716,743]" pageId="1" pageNumber="425">Ogasawara Islands</collectingMunicipality>
, scuba diving, coll. by
<collectorName id="26DEE0ADFF84FFE2AC60FD09198B8D89" box="[1086,1224,717,743]" pageId="1" pageNumber="425">H. Komatsu</collectorName>
,
<date id="FF95A3BBFF84FFE2AC8DFD0918198D89" box="[1235,1370,716,743]" pageId="1" pageNumber="425" value="2015-07-04">
<collectingDate id="EFD15A53FF84FFE2AC8DFD0918198D89" box="[1235,1370,716,743]" pageId="1" pageNumber="425" value="2015-07-04">4 July 2015</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C331D6F0FF84FFE2A899FD341CE08CF4" pageId="1" pageNumber="425" type="diagnosis">
<paragraph id="8B94857BFF84FFE2A899FD341CE08CF4" blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="425">
<emphasis id="B95F5969FF84FFE2A899FD341C798C64" bold="true" box="[199,314,752,778]" pageId="1" pageNumber="425">Diagnosis</emphasis>
. Small species. Carapace transversely ovate, strongly convex, uniformly covered with short setae; dorsal surface microscopically granulated, with regions sharply defined by shallow, narrow furrows; front convex, medially developed, divided into two lobes by median deep fissure; anterolateral margin fringed with narrow crest, divided into four lobes by three V-shaped notches. Both chelipeds robust, subequal, similar in shape, with short, strongly curved fingers.
</paragraph>
</subSubSection>
<subSubSection id="C331D6F0FF84FFE0A899FC601FBD8C00" lastPageId="3" lastPageNumber="427" pageId="1" pageNumber="425" type="description">
<paragraph id="8B94857BFF84FFE2A899FC601F9A8AFC" blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="425">
<emphasis id="B95F5969FF84FFE2A899FC601C998CD1" bold="true" box="[199,474,932,959]" pageId="1" pageNumber="425">
Description of
<typeStatus id="54903BD9FF84FFE2A92DFC611C998CD1" box="[371,474,933,959]" pageId="1" pageNumber="425" type="holotype">holotype</typeStatus>
</emphasis>
. Carapace (
<figureCitation id="131099FEFF84FFE2AA02FC601FE48CD1" box="[604,679,932,959]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Figs. 1</figureCitation>
,
<figureCitation id="131099FEFF84FFE2AAEDFC611F818CD1" box="[691,706,933,959]" captionStart="FIGURE 4" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[272,1315,885,1909]" captionTargetId="figure-344@6.[272,1315,885,1909]" captionTargetPageId="6" captionText="FIGURE 4. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life." figureDoi="http://doi.org/10.5281/zenodo.12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="1" pageNumber="425">4</figureCitation>
) transversely ovate, narrow, ca. 1.3 times as long as wide, strongly convex longitudinally and transversely; dorsal surface uniformly covered with microscopic granules, sparsely with short setae; regions sharply defined by shallow, narrow furrows. Carapace anterolateral margin (
<figureCitation id="131099FEFF84FFE2AC9FFC28184E8B69" box="[1217,1293,1004,1031]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Figs. 1</figureCitation>
,
<figureCitation id="131099FEFF84FFE2AD44FC29186A8B69" box="[1306,1321,1005,1031]" captionStart="FIGURE 4" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[272,1315,885,1909]" captionTargetId="figure-344@6.[272,1315,885,1909]" captionTargetPageId="6" captionText="FIGURE 4. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life." figureDoi="http://doi.org/10.5281/zenodo.12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="1" pageNumber="425">4</figureCitation>
) narrowly crested along its whole length, cut into four lobes by three V-shaped notches; first lobe as long as third lobe, longer than second and fourth lobes, anterior end of first lobe forming external orbital angle; margin of each lobe lined with some small granules, with sharpened anterior end. Greatest breadth of carapace at fourth lobes of both sides. Carapace posterolateral margin (
<figureCitation id="131099FEFF84FFE2AA0AFBB81FF98BF9" box="[596,698,1148,1175]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Figs. 1A</figureCitation>
,
<figureCitation id="131099FEFF84FFE2AA96FBB91F948BF9" box="[712,727,1149,1175]" captionStart="FIGURE 4" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[272,1315,885,1909]" captionTargetId="figure-344@6.[272,1315,885,1909]" captionTargetPageId="6" captionText="FIGURE 4. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life." figureDoi="http://doi.org/10.5281/zenodo.12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="1" pageNumber="425">4</figureCitation>
) strongly convergent toward lateral end of carapace posterior margin. Carapace posterior margin (
<figureCitation id="131099FEFF84FFE2AA68FB641FDF8BD5" box="[566,668,1184,1211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Figs. 1A</figureCitation>
,
<figureCitation id="131099FEFF84FFE2AAF4FB651FFA8BD5" box="[682,697,1185,1211]" captionStart="FIGURE 4" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[272,1315,885,1909]" captionTargetId="figure-344@6.[272,1315,885,1909]" captionTargetPageId="6" captionText="FIGURE 4. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life." figureDoi="http://doi.org/10.5281/zenodo.12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="1" pageNumber="425">4</figureCitation>
) as wide as frontal margin. Front (
<figureCitation id="131099FEFF84FFE2AC10FB6419DD8BD5" box="[1102,1182,1184,1211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Figs. 1</figureCitation>
,
<figureCitation id="131099FEFF84FFE2ACF0FB6519FE8BD5" box="[1198,1213,1185,1211]" captionStart="FIGURE 4" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[272,1315,885,1909]" captionTargetId="figure-344@6.[272,1315,885,1909]" captionTargetPageId="6" captionText="FIGURE 4. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life." figureDoi="http://doi.org/10.5281/zenodo.12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="1" pageNumber="425">4</figureCitation>
) directed obliquely downward, bilobed, lobes separated by median narrow, deep slit; each lobe fringed with sharp, close-set granules, most prominently developed close to median slit, outer one-third weakly concave, ending as small lateral angle directed obliquely downward, widely separated from inner angle of supraorbital margin. Orbit (
<figureCitation id="131099FEFF84FFE2AC8CFAC818508A49" box="[1234,1299,1292,1319]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Fig. 1</figureCitation>
) deep, with inner part of supraorbital margin strongly concave; supraorbital margin narrowly raised, fringed with small closelyset granules, separated to three parts by two V-shaped notches; infraorbital margin transverse, thinly edged, without interruption; inner infraorbital angle not developed.
</paragraph>
<paragraph id="8B94857BFF84FFE2A899FA581E1A8994" blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="425">
Both chelipeds (
<figureCitation id="131099FEFF84FFE2A924FA581C868AD9" box="[378,453,1436,1463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Figs. 1</figureCitation>
,
<figureCitation id="131099FEFF84FFE2A98FFA591CA28AD9" box="[465,481,1437,1463]" captionStart="FIGURE 2" captionStartId="3.[151,250,709,733]" captionTargetBox="[151,1436,181,686]" captionTargetId="figure-20@3.[151,1436,181,686]" captionTargetPageId="3" captionText="FIGURE 2. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, chelae, outer view; B, pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681869" httpUri="https://zenodo.org/record/12681869/files/figure.png" pageId="1" pageNumber="425">2</figureCitation>
,
<figureCitation id="131099FEFF84FFE2A9B5FA591F4C8AD9" box="[491,527,1437,1463]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. AC, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng &amp; Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; CD, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="1" pageNumber="425">3A</figureCitation>
,
<figureCitation id="131099FEFF84FFE2AA47FA591F6B8AD9" box="[537,552,1437,1463]" captionStart="FIGURE 4" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[272,1315,885,1909]" captionTargetId="figure-344@6.[272,1315,885,1909]" captionTargetPageId="6" captionText="FIGURE 4. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life." figureDoi="http://doi.org/10.5281/zenodo.12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="1" pageNumber="425">4</figureCitation>
) robust, subequal. Merus short, entirely concealed under carapace, when viewed dorsally; both with lower margins armed with rows of several obtuse granules. Carpus thickly covered with small sharp granules sparsely interspaced with short setae, with inner surface touching with carapace epimeral surface; inner angle blunt, without tooth. Palm (
<figureCitation id="131099FEFF84FFE2AA01F9CC1F86894D" box="[607,709,1544,1571]" captionStart="FIGURE 2" captionStartId="3.[151,250,709,733]" captionTargetBox="[151,1436,181,686]" captionTargetId="figure-20@3.[151,1436,181,686]" captionTargetPageId="3" captionText="FIGURE 2. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, chelae, outer view; B, pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681869" httpUri="https://zenodo.org/record/12681869/files/figure.png" pageId="1" pageNumber="425">Figs. 2A</figureCitation>
,
<figureCitation id="131099FEFF84FFE2AA8DF9CD1FB4894D" box="[723,759,1545,1571]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. AC, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng &amp; Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; CD, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="1" pageNumber="425">3A</figureCitation>
) short, not strongly bulged outward; outer surface covered with small conical granules of various sizes; upper margin sharp, with row of several larger, sharper granules; outer surface thickly covered with short setae sparsely interspaced with long setae. Both fingers (
<figureCitation id="131099FEFF84FFE2ACA2F99418218905" box="[1276,1378,1616,1643]" captionStart="FIGURE 2" captionStartId="3.[151,250,709,733]" captionTargetBox="[151,1436,181,686]" captionTargetId="figure-20@3.[151,1436,181,686]" captionTargetPageId="3" captionText="FIGURE 2. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, chelae, outer view; B, pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681869" httpUri="https://zenodo.org/record/12681869/files/figure.png" pageId="1" pageNumber="425">Figs. 2A</figureCitation>
,
<figureCitation id="131099FEFF84FFE2AD2FF99518D78905" box="[1393,1428,1617,1643]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. AC, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng &amp; Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; CD, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="1" pageNumber="425">3A</figureCitation>
) short, stout, with prominent tuft of setae on each external surface; immovable finger one-third as long as palm, with triangular tooth occupying more than half length of cutting edge; movable finger stout, length slightly less than upper margin of palm, strongly curved at distal one-third toward to tip of immovable finger; dorsal surface of movable finger armed with several sharp, tuberculate granules.
</paragraph>
<paragraph id="8B94857BFF84FFE2A899F8C0193088E4" blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="425">
Ambulatory legs (
<figureCitation id="131099FEFF84FFE2A9CEF8C01C9E8871" box="[400,477,1796,1823]" captionStart="FIGURE 1" captionStartId="2.[151,250,1974,1998]" captionTargetBox="[226,1361,181,1951]" captionTargetId="figure-17@2.[226,1361,181,1951]" captionTargetPageId="2" captionText="FIGURE 1. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="1" pageNumber="425">Figs. 1</figureCitation>
,
<figureCitation id="131099FEFF84FFE2A9B4F8C11F4E8871" box="[490,525,1797,1823]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. AC, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng &amp; Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; CD, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="1" pageNumber="425">3B</figureCitation>
,
<figureCitation id="131099FEFF84FFE2AA46F8C11F648871" box="[536,551,1797,1823]" captionStart="FIGURE 4" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[272,1315,885,1909]" captionTargetId="figure-344@6.[272,1315,885,1909]" captionTargetPageId="6" captionText="FIGURE 4. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life." figureDoi="http://doi.org/10.5281/zenodo.12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="1" pageNumber="425">4</figureCitation>
) stout, moderate in length, covered with silky setae of various lengths on dorsal and ventral margins of meri, carpi and propodi of all pairs; dorsal margins of meri, carpi and propodi subacute, not sharply crested; both lower margins of meri of first three pairs sharp-edged, armed with several minute, sharp granules; merus of last pair with denser setae and smaller granules along lower margins.
</paragraph>
<paragraph id="8B94857BFF84FFE2A899F8501FF18898" blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="425">
Male pleon narrow (
<figureCitation id="131099FEFF84FFE2A9EEF8501F4B88C1" box="[432,520,1940,1967]" captionStart="FIGURE 2" captionStartId="3.[151,250,709,733]" captionTargetBox="[151,1436,181,686]" captionTargetId="figure-20@3.[151,1436,181,686]" captionTargetPageId="3" captionText="FIGURE 2. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, chelae, outer view; B, pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681869" httpUri="https://zenodo.org/record/12681869/files/figure.png" pageId="1" pageNumber="425">Fig. 2B</figureCitation>
) throughout six somites, first two somites setose; telson as long as sixth somite, weakly tapering distally. G1 (
<figureCitation id="131099FEFF84FFE2A9B9F87C1F7D88BD" box="[487,574,1976,2003]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. AC, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng &amp; Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; CD, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="1" pageNumber="425">Fig. 3C</figureCitation>
) of typical pilumnid shape, long, weakly curved, occupying whole space of abdominal trench; tip sharply pointed. G2 short.
</paragraph>
<caption id="DF54D5F3FF87FFE1A8C9F8721FF2889D" ID-DOI="http://doi.org/10.5281/zenodo.12681865" ID-Zenodo-Dep="12681865" httpUri="https://zenodo.org/record/12681865/files/figure.png" pageId="2" pageNumber="426" startId="2.[151,250,1974,1998]" targetBox="[226,1361,181,1951]" targetPageId="2" targetType="figure">
<paragraph id="8B94857BFF87FFE1A8C9F8721FF2889D" blockId="2.[151,1436,1974,2035]" pageId="2" pageNumber="426">
<emphasis id="B95F5969FF87FFE1A8C9F8721C5788A0" bold="true" box="[151,276,1974,1998]" pageId="2" pageNumber="426">FIGURE 1.</emphasis>
<taxonomicName id="4C2BFEF8FF87FFE1A942F8731CB088A1" authorityName="Takeda &amp; Komatsu" authorityYear="2024" box="[284,499,1975,1999]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="426" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="B95F5969FF87FFE1A942F8731CB088A1" box="[284,499,1975,1999]" italics="true" pageId="2" pageNumber="426">Actumnus ngankeeae</emphasis>
</taxonomicName>
<emphasis id="B95F5969FF87FFE1A9A5F8731F0E88A1" bold="true" box="[507,589,1975,1999]" pageId="2" pageNumber="426">
<taxonomicNameLabel id="A26CE412FF87FFE1A9A5F8731F0E88A1" box="[507,589,1975,1999]" pageId="2" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, habitus, dorsal view; B, habitus, oblique dorsal view.
</paragraph>
</caption>
<caption id="DF54D5F3FF86FFE0A8C9FD011F078C6C" ID-DOI="http://doi.org/10.5281/zenodo.12681869" ID-Zenodo-Dep="12681869" httpUri="https://zenodo.org/record/12681869/files/figure.png" pageId="3" pageNumber="427" startId="3.[151,250,709,733]" targetBox="[151,1436,181,686]" targetPageId="3" targetType="figure">
<paragraph id="8B94857BFF86FFE0A8C9FD011F078C6C" blockId="3.[151,1436,709,770]" pageId="3" pageNumber="427">
<emphasis id="B95F5969FF86FFE0A8C9FD011C578DB0" bold="true" box="[151,276,709,734]" pageId="3" pageNumber="427">FIGURE 2.</emphasis>
<taxonomicName id="4C2BFEF8FF86FFE0A942FD021CB08DB0" authorityName="Takeda &amp; Komatsu" authorityYear="2024" box="[284,499,710,734]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="B95F5969FF86FFE0A942FD021CB08DB0" box="[284,499,710,734]" italics="true" pageId="3" pageNumber="427">Actumnus ngankeeae</emphasis>
</taxonomicName>
<emphasis id="B95F5969FF86FFE0A9A5FD021F0E8DB0" bold="true" box="[507,589,710,734]" pageId="3" pageNumber="427">
<taxonomicNameLabel id="A26CE412FF86FFE0A9A5FD021F0E8DB0" box="[507,589,710,734]" pageId="3" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. A, chelae, outer view; B, pleon, ventral view.
</paragraph>
</caption>
<paragraph id="8B94857BFF86FFE0A899FCF41FBD8C00" blockId="3.[151,1437,816,2030]" pageId="3" pageNumber="427">
<emphasis id="B95F5969FF86FFE0A899FCF41C148C24" bold="true" box="[199,343,816,842]" pageId="3" pageNumber="427">Color in life</emphasis>
(
<figureCitation id="131099FEFF86FFE0A939FCF41CEE8C24" box="[359,429,816,842]" captionStart="FIGURE 4" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[272,1315,885,1909]" captionTargetId="figure-344@6.[272,1315,885,1909]" captionTargetPageId="6" captionText="FIGURE 4. Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life." figureDoi="http://doi.org/10.5281/zenodo.12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="3" pageNumber="427">Fig. 4</figureCitation>
). Yellowish tan, sparsely speckled with blurry reddish spots; setae on carapace, chelipeds and ambulatory legs reddish to yellow; cornea reddish.
</paragraph>
</subSubSection>
<subSubSection id="C331D6F0FF86FFE0A899FCBC182C8CFC" box="[199,1391,888,914]" pageId="3" pageNumber="427" type="etymology">
<paragraph id="8B94857BFF86FFE0A899FCBC182C8CFC" blockId="3.[151,1437,816,2030]" box="[199,1391,888,914]" pageId="3" pageNumber="427">
<emphasis id="B95F5969FF86FFE0A899FCBC1C0B8CFC" bold="true" box="[199,328,888,914]" pageId="3" pageNumber="427">Etymology</emphasis>
. This new species is named after the late Dr. Ng Ngan Kee, who was our friend and colleague.
</paragraph>
</subSubSection>
<subSubSection id="C331D6F0FF86FFE7A899FC581F148EE7" lastPageId="4" lastPageNumber="428" pageId="3" pageNumber="427" type="discussion">
<paragraph id="8B94857BFF86FFE0A899FC5818388B94" blockId="3.[151,1437,816,2030]" pageId="3" pageNumber="427">
<emphasis id="B95F5969FF86FFE0A899FC581C778CD8" bold="true" box="[199,308,924,950]" pageId="3" pageNumber="427">Remarks</emphasis>
.
<taxonomicName id="4C2BFEF8FF86FFE0A91CFC581F6C8CD8" authorityName="Takeda &amp; Komatsu" authorityYear="2024" box="[322,559,924,950]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="B95F5969FF86FFE0A91CFC581F6C8CD8" box="[322,559,924,950]" italics="true" pageId="3" pageNumber="427">Actumnus ngankeeae</emphasis>
</taxonomicName>
<emphasis id="B95F5969FF86FFE0AA68FC581FD38CD8" bold="true" box="[566,656,924,950]" pageId="3" pageNumber="427">
<taxonomicNameLabel id="A26CE412FF86FFE0AA68FC581FD38CD8" box="[566,656,924,950]" pageId="3" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is close to
<taxonomicName id="4C2BFEF8FF86FFE0AB50FC5819048CD8" authority="(Rathbun, 1907)" baseAuthorityName="Rathbun" baseAuthorityYear="1907" box="[782,1095,924,950]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="digitalis">
<emphasis id="B95F5969FF86FFE0AB50FC581ECA8CD8" box="[782,905,924,950]" italics="true" pageId="3" pageNumber="427">A. digitalis</emphasis>
(
<bibRefCitation id="EFBAF88AFF86FFE0ABC9FC58197C8CD8" author="Rathbun, M. J." box="[919,1087,924,950]" pageId="3" pageNumber="427" refId="ref6254" refString="Rathbun, M. J. (1907) The Brachyura. IX. Reports on the scientific results. of the expedition to the tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from August, 1899, to March, 1900, Commander Jefferson F. Moser, U. S. N., commanding. X. Reports on the scientific results of the expedition to the eastern tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from October, 1904, to March, 1905, Lieut. - Commander L. M. Garrett, U. S. N., commanding. Memoirs of the Museum of Comparative Zoology at Harvard College, 35, 23 - 74, pls. 1 - 9." type="journal volume" year="1907">Rathbun, 1907</bibRefCitation>
)
</taxonomicName>
and its synonym,
<taxonomicName id="4C2BFEF8FF86FFE0AD4DFC581C788CB5" authority="Bouvier, 1915" authorityName="Bouvier" authorityYear="1915" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B95F5969FF86FFE0AD4DFC5818DF8CD8" box="[1299,1436,924,950]" italics="true" pageId="3" pageNumber="427">A. carinatus</emphasis>
<bibRefCitation id="EFBAF88AFF86FFE0A8C9FC041C788CB5" author="Bouvier, E. - L." box="[151,315,960,987]" pageId="3" pageNumber="427" refId="ref5183" refString="Bouvier, E. - L. (1915) Decapodes marcheurs (Reptantia) et stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique, 48, 178 - 318, pls. 4 - 7." type="journal volume" year="1915">Bouvier, 1915</bibRefCitation>
</taxonomicName>
, in the general appearance of the carapace, chelipeds and ambulatory legs.
<taxonomicName id="4C2BFEF8FF86FFE0ACCBFC04182B8CB4" baseAuthorityName="Rathbun" baseAuthorityYear="1907" box="[1173,1384,960,986]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="digitalis">
<emphasis id="B95F5969FF86FFE0ACCBFC04182B8CB4" box="[1173,1384,960,986]" italics="true" pageId="3" pageNumber="427">Actumnus digitalis</emphasis>
</taxonomicName>
was originally referred to the genus
<taxonomicName id="4C2BFEF8FF86FFE0AA5CFC201FBC8C91" authority="Bell, 1835" authorityName="Bell" authorityYear="1835" box="[514,767,996,1023]" class="Malacostraca" family="Xanthidae" genus="Platypodia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="B95F5969FF86FFE0AA5CFC201F3E8C90" box="[514,637,996,1022]" italics="true" pageId="3" pageNumber="427">Platypodia</emphasis>
<bibRefCitation id="EFBAF88AFF86FFE0AADBFC201FBC8C91" author="Bell, T." box="[645,767,996,1023]" pageId="3" pageNumber="427" refId="ref5113" refString="Bell, T. (1835) Observations on the genus Cancer of Dr. Leach (Platycarcinus, Latr.,), with descriptions of three new species. Transactions of the Zoological Society of London, 1, 335 - 342, pls. 43 - 47. https: // doi. org / 10.1111 / j. 1096 - 3642.1835. tb 00634. x" type="journal volume" year="1835">Bell, 1835</bibRefCitation>
</taxonomicName>
, but is really a pilumnid species and accommodated in the genus
<taxonomicName id="4C2BFEF8FF86FFE0A880FBCC1C0F8B4C" authorityName="Dana" authorityYear="1851" box="[222,332,1032,1058]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="B95F5969FF86FFE0A880FBCC1C0F8B4C" box="[222,332,1032,1058]" italics="true" pageId="3" pageNumber="427">Actumnus</emphasis>
</taxonomicName>
due to the dorsally convex carapace with the regions defined by narrow furrows, the stout chelipeds having the short and convex fingers, the sharp but not crested dorsal margins of the ambulatory legs, the narrow thoracic sternum, the slender and freely segmented male pleon, and the pilumnid
<typeStatus id="54903BD9FF86FFE0AC6CFB9419218B04" box="[1074,1122,1104,1130]" pageId="3" pageNumber="427">type</typeStatus>
G1. As figured by
<bibRefCitation id="EFBAF88AFF86FFE0AD63FB941D9E8BE0" author="Rathbun, M. J." pageId="3" pageNumber="427" refId="ref6254" refString="Rathbun, M. J. (1907) The Brachyura. IX. Reports on the scientific results. of the expedition to the tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from August, 1899, to March, 1900, Commander Jefferson F. Moser, U. S. N., commanding. X. Reports on the scientific results of the expedition to the eastern tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from October, 1904, to March, 1905, Lieut. - Commander L. M. Garrett, U. S. N., commanding. Memoirs of the Museum of Comparative Zoology at Harvard College, 35, 23 - 74, pls. 1 - 9." type="journal volume" year="1907">Rathbun (1907</bibRefCitation>
: pl. 1 fig. 6, as
<taxonomicName id="4C2BFEF8FF86FFE0A9CFFBB11F498BE0" baseAuthorityName="Rathbun" baseAuthorityYear="1907" box="[401,522,1140,1166]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="digitalis">
<emphasis id="B95F5969FF86FFE0A9CFFBB11F498BE0" box="[401,522,1140,1166]" italics="true" pageId="3" pageNumber="427">P. digitalis</emphasis>
</taxonomicName>
from Kusaie, now
<collectingRegion id="49EF4B99FF86FFE0AAB6FBB01E748BE0" box="[744,823,1140,1166]" country="Micronesia" name="Kosrae" pageId="3" pageNumber="427">Kosrae</collectingRegion>
in the
<collectingCountry id="F33CC5EBFF86FFE0ABD7FBB019078BE0" box="[905,1092,1140,1166]" name="Micronesia" pageId="3" pageNumber="427">Caroline Islands</collectingCountry>
, and Papeete, Tahiti),
<bibRefCitation id="EFBAF88AFF86FFE0AD1CFBB01D9E8BDD" author="Bouvier, E. - L." pageId="3" pageNumber="427" refId="ref5183" refString="Bouvier, E. - L. (1915) Decapodes marcheurs (Reptantia) et stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique, 48, 178 - 318, pls. 4 - 7." type="journal volume" year="1915">Bouvier (1915</bibRefCitation>
: fig. 26, as
<taxonomicName id="4C2BFEF8FF86FFE0A907FB5C1CA28BDC" authorityName="Bouvier" authorityYear="1915" box="[345,481,1176,1202]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B95F5969FF86FFE0A907FB5C1CA28BDC" box="[345,481,1176,1202]" italics="true" pageId="3" pageNumber="427">A. carinatus</emphasis>
</taxonomicName>
from
<collectingRegion id="49EF4B99FF86FFE0AA7DFB5C1FDB8BDC" box="[547,664,1176,1202]" country="Mauritius" name="Port Louis" pageId="3" pageNumber="427">Port Louis</collectingRegion>
in
<collectingCountry id="F33CC5EBFF86FFE0AAE4FB5C1E6A8BDC" box="[698,809,1176,1202]" name="Mauritius" pageId="3" pageNumber="427">Mauritius</collectingCountry>
) and
<bibRefCitation id="EFBAF88AFF86FFE0AB3AFB5C19348BDC" author="Takeda, M. &amp; Miyake, S." box="[868,1143,1176,1202]" pageId="3" pageNumber="427" refId="ref6738" refString="Takeda, M. &amp; Miyake, S. (1969) Pilumnid crabs of the family Xanthidae from the West Pacific. II. Twenty-one species of four genera, with descriptions of four new species. Ohmu, 2, 93 - 156." type="journal volume" year="1969">Takeda &amp; Miyake (1969</bibRefCitation>
: fig. 3, as
<taxonomicName id="4C2BFEF8FF86FFE0ACB8FB5C18238BDC" baseAuthorityName="Rathbun" baseAuthorityYear="1907" box="[1254,1376,1176,1202]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="digitalis">
<emphasis id="B95F5969FF86FFE0ACB8FB5C18238BDC" box="[1254,1376,1176,1202]" italics="true" pageId="3" pageNumber="427">A. digitalis</emphasis>
</taxonomicName>
from Kamiyama-jima Island in the Ryukyu Islands), the supraorbital and anterolateral margins of the carapace are sharply crested, with two closed, deep fissures on the supraorbital margin and three fissures on the anterolateral margin.
</paragraph>
<paragraph id="8B94857BFF86FFE0A899FAC01EF88AE4" blockId="3.[151,1437,816,2030]" pageId="3" pageNumber="427">
In
<taxonomicName id="4C2BFEF8FF86FFE0A8B4FAC01C9F8A70" authorityName="Takeda &amp; Komatsu" authorityYear="2024" box="[234,476,1284,1310]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="B95F5969FF86FFE0A8B4FAC01C9F8A70" box="[234,476,1284,1310]" italics="true" pageId="3" pageNumber="427">Actumnus ngankeeae</emphasis>
</taxonomicName>
, the carapace anterolateral margin is narrowly crested throughout the length, and divided into four distinctly isolated lobes by three deep, V-shaped notches. Furthermore, in the new species, the carapace dorsal surface is uniformly covered only with short setae, but in
<taxonomicName id="4C2BFEF8FF86FFE0AB8DFA88190D8A08" baseAuthorityName="Rathbun" baseAuthorityYear="1907" box="[979,1102,1356,1382]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="digitalis">
<emphasis id="B95F5969FF86FFE0AB8DFA88190D8A08" box="[979,1102,1356,1382]" italics="true" pageId="3" pageNumber="427">A. digitalis</emphasis>
</taxonomicName>
, the setae are restricted to the regions which are weakly convex dorsally and isolated from each other.
</paragraph>
<paragraph id="8B94857BFF86FFE0A899FA501C4B8974" blockId="3.[151,1437,816,2030]" pageId="3" pageNumber="427">
It is noteworthy that
<taxonomicName id="4C2BFEF8FF86FFE0A9E3FA501E068AC0" authority="Deb, 1992" authorityName="Deb" authorityYear="1992" box="[445,837,1428,1454]" class="Malacostraca" family="Xanthidae" genus="Platypodia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="andamania">
<emphasis id="B95F5969FF86FFE0A9E3FA501F838AC0" box="[445,704,1428,1454]" italics="true" pageId="3" pageNumber="427">Platypodia andamania</emphasis>
<bibRefCitation id="EFBAF88AFF86FFE0AA94FA501E068AC0" author="Deb, M." box="[714,837,1428,1454]" pageId="3" pageNumber="427" refId="ref5393" refString="Deb, M. (1992) Two new species of xanthid crabs from Bay Islands. Journal of the Andaman Science Association, 8, 121 - 124." type="journal volume" year="1992">Deb, 1992</bibRefCitation>
</taxonomicName>
, may be synonymous with
<taxonomicName id="4C2BFEF8FF86FFE0ACDBFA5018198AC0" baseAuthorityName="Rathbun" baseAuthorityYear="1907" box="[1157,1370,1428,1454]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="digitalis">
<emphasis id="B95F5969FF86FFE0ACDBFA5018198AC0" box="[1157,1370,1428,1454]" italics="true" pageId="3" pageNumber="427">Actumnus digitalis</emphasis>
</taxonomicName>
. This species, described from the Andaman Islands, was left behind the recent studies since its original description. The original figure is so schematic that it is difficult to make a definitive conclusion, but the figure is similar to
<taxonomicName id="4C2BFEF8FF86FFE0ADDAFA181C428974" authorityName="Bouvier" authorityYear="1915" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B95F5969FF86FFE0ADDAFA181C428974" italics="true" pageId="3" pageNumber="427">A. carinatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B94857BFF86FFE0A899F9E01F128878" blockId="3.[151,1437,816,2030]" pageId="3" pageNumber="427">
<taxonomicName id="4C2BFEF8FF86FFE0A899F9E01CF98950" authorityName="Takeda &amp; Komatsu" authorityYear="2024" box="[199,442,1572,1598]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="B95F5969FF86FFE0A899F9E01CF98950" box="[199,442,1572,1598]" italics="true" pageId="3" pageNumber="427">Actumnus ngankeeae</emphasis>
</taxonomicName>
is somewhat similar to
<taxonomicName id="4C2BFEF8FF86FFE0AABEF9E0194C8950" authority="(Herbst, 1801)" baseAuthorityName="Herbst" baseAuthorityYear="1801" box="[736,1039,1572,1598]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="B95F5969FF86FFE0AABEF9E01E188950" box="[736,859,1572,1598]" italics="true" pageId="3" pageNumber="427">A. calypso</emphasis>
(
<bibRefCitation id="EFBAF88AFF86FFE0AB30F9E019448950" author="Herbst, J. F. W." box="[878,1031,1572,1598]" pageId="3" pageNumber="427" refId="ref5647" refString="Herbst, J. F. W. (1801) Versuch einer Naturgeschichte der Krabben und Krebse nebst einer systematischen Beschreibung ihrer verschiedenen Arten, 3 (3), 1 - 54, pls. 55 - 58. Gottlieb August Lange, Berlin und Stralsund." type="book" year="1801">Herbst, 1801</bibRefCitation>
)
</taxonomicName>
in the general appearance of the carapace. The original figure of
<taxonomicName id="4C2BFEF8FF86FFE0A9A1F98C1F37890C" baseAuthorityName="Herbst" baseAuthorityYear="1801" box="[511,628,1608,1634]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="B95F5969FF86FFE0A9A1F98C1F37890C" box="[511,628,1608,1634]" italics="true" pageId="3" pageNumber="427">A. calypso</emphasis>
</taxonomicName>
is schematic, but indicates the characteristic formation of the proto- and mesogastric regions that is distinguished in the photograph of the
<typeStatus id="54903BD9FF86FFE0ABD8F9A81EF589E8" box="[902,950,1644,1670]" pageId="3" pageNumber="427">type</typeStatus>
specimen of
<taxonomicName id="4C2BFEF8FF86FFE0AC0FF9A8198489E8" baseAuthorityName="Herbst" baseAuthorityYear="1801" box="[1105,1223,1644,1670]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="B95F5969FF86FFE0AC0FF9A8198489E8" box="[1105,1223,1644,1670]" italics="true" pageId="3" pageNumber="427">A. calypso</emphasis>
</taxonomicName>
given by K.
<bibRefCitation id="EFBAF88AFF86FFE0AD00F9A81D9E89C4" author="Sakai, K." pageId="3" pageNumber="427" refId="ref6413" refString="Sakai, K. (1999) J. F. W. Herbst-Collection of decapod Crustacea of the Berlin Zoological Museum, with remarks on certain species. Naturalists, 6, 1 ⎼ 45, pls. 1 ⎼ 21." type="journal volume" year="1999">Sakai (1999</bibRefCitation>
: pl. 18 fig. E), and also in the illustration of the synonymous species,
<taxonomicName id="4C2BFEF8FF86FFE0ABB0F954181089C4" authority="Henderson, 1893" authorityName="Henderson" authorityYear="1893" box="[1006,1363,1680,1706]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="B95F5969FF86FFE0ABB0F95419CA89C4" box="[1006,1161,1680,1706]" italics="true" pageId="3" pageNumber="427">A. verrucosus</emphasis>
<bibRefCitation id="EFBAF88AFF86FFE0ACCEF954181089C4" author="Henderson, J. R." box="[1168,1363,1680,1706]" pageId="3" pageNumber="427" refId="ref5590" refString="Henderson, J. R. (1893) A contribution to Indian Carcinology. Transactions of the Linnean Society of London, Series 2, 5, 325 ⎼ 458, pls. 36 ⎼ 40. https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal volume" year="1893">Henderson, 1893</bibRefCitation>
</taxonomicName>
, given by
<bibRefCitation id="EFBAF88AFF86FFE0A8E5F9701CC189A0" author="Henderson, J. R." box="[187,386,1716,1742]" pageId="3" pageNumber="427" refId="ref5590" refString="Henderson, J. R. (1893) A contribution to Indian Carcinology. Transactions of the Linnean Society of London, Series 2, 5, 325 ⎼ 458, pls. 36 ⎼ 40. https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal volume" year="1893">Henderson (1893</bibRefCitation>
: pl. 36 fig. 15). In
<taxonomicName id="4C2BFEF8FF86FFE0AA07F9701F8C89A0" baseAuthorityName="Herbst" baseAuthorityYear="1801" box="[601,719,1716,1742]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="B95F5969FF86FFE0AA07F9701F8C89A0" box="[601,719,1716,1742]" italics="true" pageId="3" pageNumber="427">A. calypso</emphasis>
</taxonomicName>
, the protogastric region is imperfectly subdivided into three by two oblique furrows, with three-forked image, and the mesogastric region is symmetrically subdivided into anterior one and posterior two by wide furrows.
</paragraph>
<paragraph id="8B94857BFF86FFE7A899F8E41F148EE7" blockId="3.[151,1437,816,2030]" lastBlockId="4.[151,1437,151,465]" lastPageId="4" lastPageNumber="428" pageId="3" pageNumber="427">
<taxonomicName id="4C2BFEF8FF86FFE0A899F8E41F628855" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[199,545,1824,1851]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B95F5969FF86FFE0A899F8E41CCE8854" box="[199,397,1824,1850]" italics="true" pageId="3" pageNumber="427">Actumnus obesus</emphasis>
<bibRefCitation id="EFBAF88AFF86FFE0A9C6F8E41F628855" author="Dana, J. D." box="[408,545,1824,1851]" pageId="3" pageNumber="427" refId="ref5264" refString="Dana, J. D. (1852, 1855) Crustacea. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842 under the command of Charles Wilkes, U. S. N., 13 (1), i - viii, 1 - 685. [Atlas, Crustacea (1855). C. Sherman, Philadelphia, 27 pp., 96 pls.]" type="book" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
, reported by
<bibRefCitation id="EFBAF88AFF86FFE0AA9EF8E41E2C8854" author="Rathbun, M. J." box="[704,879,1824,1850]" pageId="3" pageNumber="427" refId="ref6216" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23, 827 ⎼ 930, pls. 1 ⎼ 24." type="journal volume" year="1906">Rathbun (1906</bibRefCitation>
: pl. 11 fig. 2) and
<bibRefCitation id="EFBAF88AFF86FFE0AC0AF8E418698854" author="Edmondson, C. H." box="[1108,1322,1824,1850]" pageId="3" pageNumber="427" refId="ref5423" refString="Edmondson, C. H. (1962) Xanthidae of Hawaii. Occasional Papers of Bernice P. Bishop Museum, 22, 215 ⎼ 309." type="journal volume" year="1962">Edmondson (1962</bibRefCitation>
: fig. 28a) from the Hawaiian Islands, seem to be close to
<taxonomicName id="4C2BFEF8FF86FFE0AAE2F8801E168830" authorityName="Takeda &amp; Komatsu" authorityYear="2024" box="[700,853,1860,1886]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="B95F5969FF86FFE0AAE2F8801E168830" box="[700,853,1860,1886]" italics="true" pageId="3" pageNumber="427">A. ngankeeae</emphasis>
</taxonomicName>
in the general form of the carapace, chelipeds and ambulatory legs.
<bibRefCitation id="EFBAF88AFF86FFE0A903F8AC1CA888ED" author="Dana, J. D." box="[349,491,1896,1923]" pageId="3" pageNumber="427" refId="ref5264" refString="Dana, J. D. (1852, 1855) Crustacea. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842 under the command of Charles Wilkes, U. S. N., 13 (1), i - viii, 1 - 685. [Atlas, Crustacea (1855). C. Sherman, Philadelphia, 27 pp., 96 pls.]" type="book" year="1852">Dana (1852)</bibRefCitation>
described the specimen from Maui Island, the Hawaiian Islands, as “Anterolateral margin arcuate, almost entire, very faintly four-lobed, lobes minutely denticulate,” but as
<bibRefCitation id="EFBAF88AFF86FFE0ACF2F84818DF88C9" author="Rathbun, M. J." box="[1196,1436,1932,1959]" pageId="3" pageNumber="427" refId="ref6216" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23, 827 ⎼ 930, pls. 1 ⎼ 24." type="journal volume" year="1906">Rathbun (1906: 865)</bibRefCitation>
mentioned, no lobes are indicated by Dana (1855: pl. 14 fig. 3a). In spite of some discrepancies with the original description,
<bibRefCitation id="EFBAF88AFF86FFE0A97BF8101C9A8880" author="Rathbun, M. J." box="[293,473,2004,2030]" pageId="3" pageNumber="427" refId="ref6216" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23, 827 ⎼ 930, pls. 1 ⎼ 24." type="journal volume" year="1906">Rathbun (1906)</bibRefCitation>
recorded the specimens from Molokai and Maui Islands as
<taxonomicName id="4C2BFEF8FF86FFE0ACC2F81018488880" authorityName="Dana" authorityYear="1852" box="[1180,1291,2004,2030]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B95F5969FF86FFE0ACC2F81018488880" box="[1180,1291,2004,2030]" italics="true" pageId="3" pageNumber="427">A. obesus</emphasis>
</taxonomicName>
, with a fine photograph that is apparently different from the Danas original figure. Later,
<taxonomicName id="4C2BFEF8FF81FFE7AC7AFF5319D08FDF" authorityName="Dana" authorityYear="1852" box="[1060,1171,151,177]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="428" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B95F5969FF81FFE7AC7AFF5319D08FDF" box="[1060,1171,151,177]" italics="true" pageId="4" pageNumber="428">A. obesus</emphasis>
</taxonomicName>
was recorded, without figures, by many authors from the wide area of the Indo-Pacific waters, most remarkably by
<bibRefCitation id="EFBAF88AFF81FFE7AC9DFF7F18108FBB" author="Balss, H." box="[1219,1363,187,213]" pageId="4" pageNumber="428" refId="ref5021" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattung Pilumnus (Crustacea Dekapoda) und verwandter Gattungen. Capita Zoologica, 4 (3), 1 ⎼ 47, pls. 1 - 7." type="journal volume" year="1933">Balss (1933)</bibRefCitation>
based on the specimens from
<collectingCountry id="F33CC5EBFF81FFE7A9C1FF1B1F6B8F97" box="[415,552,223,249]" name="Madagascar" pageId="4" pageNumber="428">Madagascar</collectingCountry>
, the Persian Gulf, Western Australia, the Andamans, New
<collectingCountry id="F33CC5EBFF81FFE7ACE5FF1B184C8F97" box="[1211,1295,223,249]" name="Guinea-Bissau" pageId="4" pageNumber="428">Guinea</collectingCountry>
,
<collectingCountry id="F33CC5EBFF81FFE7AD44FF1B182A8F97" box="[1306,1385,223,249]" name="Samoa" pageId="4" pageNumber="428">Samoa</collectingCountry>
,
<collectingCountry id="F33CC5EBFF81FFE7AD2BFF1B18DF8F97" box="[1397,1436,223,249]" name="Fiji" pageId="4" pageNumber="428">Fiji</collectingCountry>
and the Marquesas. It is highly probable that most of the old records followed
<bibRefCitation id="EFBAF88AFF81FFE7AC7DFEC7199B8E73" author="Rathbun, M. J." box="[1059,1240,259,285]" pageId="4" pageNumber="428" refId="ref6216" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23, 827 ⎼ 930, pls. 1 ⎼ 24." type="journal volume" year="1906">Rathbun (1906)</bibRefCitation>
. The
<typeStatus id="54903BD9FF81FFE7AD44FEC7183F8E73" box="[1306,1404,259,285]" pageId="4" pageNumber="428" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2BFEF8FF81FFE7A8C9FEE31C6C8E2F" authorityName="Takeda &amp; Komatsu" authorityYear="2024" box="[151,303,295,321]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="428" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="B95F5969FF81FFE7A8C9FEE31C6C8E2F" box="[151,303,295,321]" italics="true" pageId="4" pageNumber="428">A. ngankeeae</emphasis>
</taxonomicName>
is rather small, but such characters as the fully developed G1, the distinct carapace dorsal areolation and anterolateral armature, the stout chelipeds and the ambulatory legs embody the adult form and are valid for the subsequent identification of the species.
</paragraph>
</subSubSection>
<subSubSection id="C331D6F0FF81FFE7A899FE561C058EBF" pageId="4" pageNumber="428" type="distribution">
<paragraph id="8B94857BFF81FFE7A899FE561C058EBF" blockId="4.[151,1437,151,465]" pageId="4" pageNumber="428">
<emphasis id="B95F5969FF81FFE7A899FE561C228EC2" bold="true" box="[199,353,402,428]" pageId="4" pageNumber="428">Distribution.</emphasis>
The
<typeStatus id="54903BD9FF81FFE7A9C0FE571C8D8EC3" box="[414,462,403,429]" pageId="4" pageNumber="428">type</typeStatus>
locality is Haha-jima Island, Ogasawara (Bonin) Islands (
<geoCoordinate id="EE1FE3BCFF81FFE7AC37FE5619AC8EC3" box="[1129,1263,402,429]" degrees="26" direction="north" minutes="35" orientation="latitude" pageId="4" pageNumber="428" precision="15" seconds="19" value="26.588612">26°35'19&quot;N</geoCoordinate>
,
<geoCoordinate id="EE1FE3BCFF81FFE7ACA3FE5718CC8EC3" box="[1277,1423,403,429]" degrees="142" direction="east" minutes="08" orientation="longitude" pageId="4" pageNumber="428" precision="15" seconds="41" value="142.14471">142°08'41&quot;E</geoCoordinate>
), southern
<collectingCountry id="F33CC5EBFF81FFE7A8A1FE731C018EBF" box="[255,322,439,465]" name="Japan" pageId="4" pageNumber="428">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>