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<document id="1513621D8F1AEFED3D39FB2D600FB82B" ID-DOI="10.5281/zenodo.181238" ID-GBIF-Dataset="c9731fe4-5bb9-4792-8c6a-929a0bb00e82" ID-ISSN="1175-5326" ID-Zenodo-Dep="181238" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459829043349" checkinUser="plazi" docAuthor="Nieto, Carolina &amp; Richard, Barton" docDate="2008" docId="036D87E7D539FFC9FF6CB519FD70FAAB" docLanguage="en" docName="zt01727p021.pdf" docOrigin="Zootaxa 1727" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Cloeodes penai Morihara &amp; Edmunds" docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="FF54FF9FD535FFC7FFFBB32BFF8DFFD9" masterDocTitle="The genus Cloeodes (Ephemeroptera: Baetidae) in Argentina with new generic synonymy and new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="13" updateTime="1698222712371" updateUser="plazi">
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<mods:title id="09DE8A6165CE5A7888C9EACCC595B2CA">The genus Cloeodes (Ephemeroptera: Baetidae) in Argentina with new generic synonymy and new species</mods:title>
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<mods:namePart id="973DD7AA56DFEAE29FF0B51371A4F0B9">Nieto, Carolina</mods:namePart>
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<subSubSection id="C3DE657AD539FFCBFF6CB519FEB8F9B6" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B7B36F1D539FFCBFF6CB519FD0FF995" blockId="12.[151,642,1586,1647]" box="[151,642,1586,1612]" pageId="12" pageNumber="13">
<heading id="D033819DD539FFCBFF6CB519FD0FF995" bold="true" box="[151,642,1586,1612]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="4CC44D72D539FFCBFF6CB519FD0FF995" ID-CoL="5ZC4W" authority="Morihara &amp; Edmunds" authorityName="Morihara &amp; Edmunds" box="[151,642,1586,1612]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B9B0EAE3D539FFCBFF6CB519FD0FF995" bold="true" box="[151,642,1586,1612]" pageId="12" pageNumber="13">
<emphasis id="B9B0EAE3D539FFCBFF6CB519FEC3F995" bold="true" box="[151,334,1586,1612]" italics="true" pageId="12" pageNumber="13">Cloeodes penai</emphasis>
(Morihara &amp; Edmunds)
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<paragraph id="8B7B36F1D539FFCBFF6CB57EFEB8F9B6" blockId="12.[151,642,1586,1647]" box="[151,309,1621,1647]" pageId="12" pageNumber="13">(Figs. 52–67)</paragraph>
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<subSubSection id="C3DE657AD539FFCBFF6CB58EFCB8F925" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8B7B36F1D539FFCBFF6CB58EFD30F965" blockId="12.[151,821,1701,1788]" box="[151,701,1701,1724]" pageId="12" pageNumber="13">
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<taxonomicName id="4CC44D72D539FFCBFF6CB58EFD34F965" authority="Morihara &amp; Edmunds, 1980: 606" authorityName="Morihara &amp; Edmunds" authorityPageNumber="606" authorityYear="1980" box="[151,697,1701,1724]" class="Insecta" family="Baetidae" genus="Notobaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B9B0EAE3D539FFCBFF6CB58EFEC1F965" box="[151,332,1701,1724]" italics="true" pageId="12" pageNumber="13">Notobaetis penai</emphasis>
<treatmentCitation id="0A6510E0D539FFCBFEA8B58EFD34F965" author="Morihara" box="[339,697,1701,1724]" page="606" pageId="12" pageNumber="13" year="1980">
<bibRefCitation id="EF554B00D539FFCBFEA8B58EFD0EF965" author="Morihara" box="[339,643,1701,1724]" pageId="12" pageNumber="17" refString="Morihara D. K. &amp; Edmunds G. F. (1980) Notobaetis: a new Genus of Baetidae (Ephemeroptera) from South America. International Revue gesamten Hydrobiologie, 65 (4), 605 - 610." type="journal article" year="1980">Morihara &amp; Edmunds, 1980</bibRefCitation>
: 606
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.
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</paragraph>
<paragraph id="8B7B36F1D539FFCBFF6CB5EEFD25F905" blockId="12.[151,821,1701,1788]" box="[151,680,1733,1756]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="ABD411DFD539FFCBFF6CB5EEFD25F905" box="[151,680,1733,1756]" pageId="12" pageNumber="13">
<taxonomicName id="4CC44D72D539FFCBFF6CB5EEFD29F905" ID-CoL="5ZC4W" authority="Waltz &amp; McCafferty, 1987a: 179" authorityName="Waltz &amp; McCafferty" authorityPageNumber="179" authorityYear="1987" box="[151,676,1733,1756]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B9B0EAE3D539FFCBFF6CB5EEFEB1F905" box="[151,316,1733,1756]" italics="true" pageId="12" pageNumber="13">Cloeodes penai</emphasis>
<treatmentCitation id="0A6510E0D539FFCBFEB8B5EEFD29F905" author="Waltz" box="[323,676,1733,1756]" page="179" pageId="12" pageNumber="13" year="1987">
<bibRefCitation id="EF554B00D539FFCBFEB8B5EEFDE2F905" author="Waltz" box="[323,623,1733,1756]" pageId="12" pageNumber="18" refString="Waltz R. D. &amp; McCafferty W. P. (1987 a) Generic Revision of Cloeodes and description of two new Genera (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 89 (1), 177 - 184. Waltz R. D. &amp; McCafferty W. P. (1987 b) Revision of the Genus Cloeodes Traver (Ephemeroptera: Baetidae). Annals of the Entomological Society of America, 80, 191 - 207." type="journal article" year="1987" yearSuffix="a">Waltz &amp; McCafferty, 1987a</bibRefCitation>
: 179
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.
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</paragraph>
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<treatmentCitationGroup id="ABD411DFD539FFCBFF6CB5CEFCB8F925" box="[151,821,1765,1788]" pageId="12" pageNumber="13">
<taxonomicName id="4CC44D72D539FFCBFF6CB5CEFCB8F925" ID-CoL="5ZC4W" authority="Waltz &amp; McCafferty, 1987b: 196" authorityName="Waltz &amp; McCafferty" authorityPageNumber="196" authorityYear="1987" box="[151,821,1765,1788]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penai" subGenus="Notobaetis">
<emphasis id="B9B0EAE3D539FFCBFF6CB5CEFE45F925" box="[151,456,1765,1788]" italics="true" pageId="12" pageNumber="13">Cloeodes (Notobaetis) penai</emphasis>
, 
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: 196
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</paragraph>
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<subSubSection id="C3DE657AD539FFC9FF6CB406FCF2FE2B" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8B7B36F1D539FFC9FF6CB406FCF2FE2B" blockId="12.[151,1436,1837,2023]" lastBlockId="14.[151,1436,152,698]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<emphasis id="B9B0EAE3D539FFCBFF6CB406FE9AF89E" bold="true" box="[151,279,1837,1863]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
<taxonomicName id="4CC44D72D539FFCBFEE4B406FE58F89F" box="[287,469,1837,1862]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B9B0EAE3D539FFCBFEE4B406FE58F89F" box="[287,469,1837,1862]" italics="true" pageId="12" pageNumber="13">Cloeodes penai</emphasis>
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can be distinguished from the other species of the genus by the following combination of characters. In the adult, 1) hind wings small, 0.1 times the length of fore wings (Figs. 52, 53a), costal projection located in the center of anterior margin (Fig. 53b); 2) genitalia (Fig. 54) bases of forceps close together, segment III rounded. In the nymphs, 1) antennae with scape subequal in length to pedicel (
<figureCitation id="13FF2A74D539FFCBFACEB48EFA03F866" box="[1333,1422,1957,1983]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="12" pageNumber="13">Fig. 55</figureCitation>
); 2) labrum (
<figureCitation id="13FF2A74D539FFCBFEE1B4E6FEE2F83E" box="[282,367,1997,2023]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="12" pageNumber="13">Fig. 56</figureCitation>
) with one subapical seta centrally and two setae near lateral margin, anterior margin with basally bifid setae near midline and apically bifid setae near lateral margin; 3) left mandible without setae between prostheca and mola, thumb of molar area transverse to anterior margin (
<figureCitation id="13FF2A74D53BFFC9FBA6B3EBFB3BFF03" box="[1117,1206,192,218]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 57</figureCitation>
); 4) right mandible with prostheca bifid (
<figureCitation id="13FF2A74D53BFFC9FE6CB3C3FE7CFEDB" box="[407,497,232,258]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 58</figureCitation>
); 5) lingua with a rounded projection (
<figureCitation id="13FF2A74D53BFFC9FC38B3C3FB90FEDB" box="[963,1053,232,258]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 59</figureCitation>
); 6) maxillary palpi subequal to galea-lacinia (
<figureCitation id="13FF2A74D53BFFC9FEC5B23BFE1AFEF3" box="[318,407,272,298]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 60</figureCitation>
); 7) labial palpi with segment III rounded (
<figureCitation id="13FF2A74D53BFFC9FC63B23BFC7DFEF3" box="[920,1008,272,298]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 61</figureCitation>
); 8) hind wing pads present, though very small; 9) dorsal edge of femora with a row of pointed spines, and apically rounded (
<figureCitation id="13FF2A74D53BFFC9FB5DB213FB71FE8B" box="[1190,1276,312,338]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 63</figureCitation>
), femora, tibiae and tarsi with bipectinate spines (
<figureCitation id="13FF2A74D53BFFC9FDB4B24BFD6CFEA3" box="[591,737,352,378]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Figs. 62, 64</figureCitation>
); 10) tarsal claws without denticles (
<figureCitation id="13FF2A74D53BFFC9FB6DB24BFB62FEA3" box="[1174,1263,352,378]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 64</figureCitation>
); 11) posterior margin of abdominal terga with spines (
<figureCitation id="13FF2A74D53BFFC9FD78B2A3FD52FE7B" box="[643,735,392,418]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 65</figureCitation>
); 12) gills elongate, broadly acute posteriorly, sometimes rounded (
<figureCitation id="13FF2A74D53BFFC9FEF3B29BFEEDFE13" box="[264,352,432,458]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 66</figureCitation>
); 13) posterior margin of paraprocts with spines (
<figureCitation id="13FF2A74D53BFFC9FC51B29BFB8FFE13" box="[938,1026,432,458]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="14" pageNumber="15">Fig. 67</figureCitation>
); 14) caudal filaments with simple setae, and with whorl of spines on each segment (as in 
<figureCitation id="13FF2A74D53BFFC9FCE0B2F3FCFEFE2B" box="[795,883,472,498]" captionStart="FIGURES 31 – 42" captionStartId="11.[151,269,1905,1929]" captionTargetBox="[186,1411,182,1904]" captionTargetId="figure@11.[186,1411,128,1904]" captionTargetPageId="11" captionText="FIGURES 31 – 42. Cloeodes incus n. comb. Nymph. Mouthparts (Figs. 31 – 36): 31, labrum d. v., left d. v., right v. v.; 32, left mandible v. v.; 33, right mandible v. v.; 34, hypopharynx v. v.; 35, maxilla v. v.; 36, labium, left d. v., right v. v. 37, leg I; 38, tarsal claw I. 39, posterior margin of tergum IV. 40, gill IV. 41, paraproct. 42, terminal filament, left cercus, right caudal filament." httpUri="https://zenodo.org/record/181243/files/figure.png" pageId="14" pageNumber="15">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DFBB6679D538FFCAFF6CB474FDF7F807" httpUri="https://zenodo.org/record/181244/files/figure.png" pageId="13" pageNumber="14" targetBox="[219,1274,190,1838]" targetPageId="13">
<paragraph id="8B7B36F1D538FFCAFF6CB474FDF7F807" blockId="13.[151,1436,1887,2014]" pageId="13" pageNumber="14">
<emphasis id="B9B0EAE3D538FFCAFF6CB474FED3F8AE" bold="true" box="[151,350,1887,1911]" pageId="13" pageNumber="14">FIGURES 43–51.</emphasis>
<taxonomicName id="4CC44D72D538FFCAFE9EB44BFD91F8AE" box="[357,540,1888,1911]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="opacus" status="sp. nov.">
<emphasis id="B9B0EAE3D538FFCAFE9EB44BFD91F8AE" box="[357,540,1888,1911]" italics="true" pageId="13" pageNumber="14">Cloeodes opacus</emphasis>
</taxonomicName>
<emphasis id="B9B0EAE3D538FFCAFDD8B474FDEDF8AE" bold="true" box="[547,608,1887,1911]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A2835798D538FFCAFDD8B474FDEDF8AE" box="[547,608,1887,1911]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Nymph. 43, general view, dorsal. Mouthparts (Figs. 44–46): 44, labrum, left d.v., right v.v.; 45, labrum: anterior margin; 46, labium, left d.v., right v.v. 47, leg I; 48, dorsal margin of femur I. 49, posterior margin of tergum IV. 50, gill IV. 51, paraproct. Figs. 52–54. 
<taxonomicName id="4CC44D72D538FFCAFCA0B48EFB8DF865" box="[859,1024,1957,1980]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B9B0EAE3D538FFCAFCA0B48EFB8DF865" box="[859,1024,1957,1980]" italics="true" pageId="13" pageNumber="14">Cloeodes penai</emphasis>
</taxonomicName>
. Male imago: 52, fore wing; 53a, hind wing; 53b, hind wing detail; 54, genitalia v.v.
</paragraph>
</caption>
<subSubSection id="C3DE657AD53BFFC9FF3DB2D4FC88FDC3" box="[198,773,511,538]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8B7B36F1D53BFFC9FF3DB2D4FC88FDC3" blockId="14.[151,1436,152,698]" box="[198,773,511,538]" pageId="14" pageNumber="15">
<emphasis id="B9B0EAE3D53BFFC9FF3DB2D4FED2FDC0" bold="true" box="[198,351,511,537]" pageId="14" pageNumber="15">Distribution</emphasis>
. 
<collectingCountry id="F3D37661D53BFFC9FE95B12BFE6AFDC3" box="[366,487,512,538]" name="Argentina" pageId="14" pageNumber="15">Argentina</collectingCountry>
: Córdoba and Tucumán.
</paragraph>
</subSubSection>
<subSubSection id="C3DE657AD53BFFC9FF3DB10CFD5AFCE8" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8B7B36F1D53BFFC9FF3DB10CFC7DFD63" blockId="14.[151,1436,152,698]" pageId="14" pageNumber="15">
<emphasis id="B9B0EAE3D53BFFC9FF3DB10CFE36FD98" bold="true" box="[198,443,551,577]" pageId="14" pageNumber="15">Material examined.</emphasis>
<typeStatus id="547F8853D53BFFC9FE38B103FDB7FD9B" box="[451,570,552,578]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
: 2 nymphs: 
<collectingCountry id="F3D37661D53BFFC9FD33B103FCFBFD9B" box="[712,886,552,578]" name="Argentina" pageId="14" pageNumber="15">ARGENTINA</collectingCountry>
, Córdoba prov, Copina (ca. 
<quantity id="4C3C9B14D53BFFC9FB39B103FA80FD9B" box="[1218,1293,552,578]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="15" unit="km" value="25.0">25 km</quantity>
WNW Alta Gracia) elev. 
<quantity id="4C3C9B14D53BFFC9FECEB17BFE1FFDB3" box="[309,402,592,618]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="14" pageNumber="15" unit="m" value="1650.0">1650 m</quantity>
, 
<date id="FF7A1031D53BFFC9FE65B17BFDEEFDB3" box="[414,611,592,618]" pageId="14" pageNumber="15" value="1967-04-11" valueMax="1967-04-14" valueMin="1967-04-11">11–14/ IV/ 1967</date>
, L. Peña. (
<collectionCode id="EDD5AE34D53BFFC9FD19B17BFCA0FDB3" box="[738,813,592,618]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="14" pageNumber="15">FSCA</collectionCode>
E1010T); 
<specimenCount id="9DC2FD78D53BFFC9FC55B17BFC73FDB3" box="[942,1022,592,618]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
imago (reared): Tucumán prov, La Cascada, La 
<collectingCountry id="F3D37661D53BFFC9FED7B153FEECFD4B" box="[300,353,632,658]" name="Iceland" pageId="14" pageNumber="15">Sala</collectingCountry>
, 
<date id="FF7A1031D53BFFC9FE96B153FE7BFD4B" box="[365,502,632,658]" pageId="14" pageNumber="15" value="1981-11-07">7/ XI/ 1981</date>
, Domínguez col.; 8 nymphs: Tucumán, San Javier, Arroyo Los Noques, 
<date id="FF7A1031D53BFFC9FABFB153FF59FD63" pageId="14" pageNumber="15" value="1978-08-03">3/ VIII/ 1978</date>
, Bertikian col. 
<typeStatus id="547F8853D53BFFC9FE7CB18BFE74FD63" box="[391,505,672,698]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
housed in 
<collectionCode id="EDD5AE34D53BFFC9FD83B18BFD43FD63" box="[632,718,672,698]" pageId="14" pageNumber="15">FAMU</collectionCode>
, other material in 
<collectionCode id="EDD5AE34D53BFFC9FC59B18BFC61FD63" box="[930,1004,672,698]" pageId="14" pageNumber="15">IFML</collectionCode>
.
</paragraph>
<paragraph id="8B7B36F1D53BFFC9FF6CB03CFD5AFCE8" blockId="14.[151,727,791,818]" box="[151,727,791,818]" pageId="14" pageNumber="15">
<taxonomicName id="4CC44D72D53BFFC9FF6CB03CFDF5FCE8" authority="(Weyenbergh) Nomen" authorityName="(Weyenbergh) Nomen" box="[151,632,791,818]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stelzneri">
<emphasis id="B9B0EAE3D53BFFC9FF6CB03CFEFDFCE8" bold="true" box="[151,368,791,817]" italics="true" pageId="14" pageNumber="15">Cloeodes stelzneri</emphasis>
(Weyenbergh) 
<emphasis id="B9B0EAE3D53BFFC9FDDEB033FDF5FCE8" box="[549,632,792,817]" italics="true" pageId="14" pageNumber="15">Nomen</emphasis>
</taxonomicName>
dubium
</paragraph>
</subSubSection>
<subSubSection id="C3DE657AD53BFFC9FF6CB043FD8BFC46" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8B7B36F1D53BFFC9FF6CB043FDBBFCA6" blockId="14.[151,765,872,959]" box="[151,566,872,895]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="ABD411DFD53BFFC9FF6CB043FDBBFCA6" box="[151,566,872,895]" pageId="14" pageNumber="15">
<taxonomicName id="4CC44D72D53BFFC9FF6CB043FDBFFCA6" authority="Weyenbergh, 1883: 170" authorityName="Weyenbergh" authorityPageNumber="170" authorityYear="1883" box="[151,562,872,895]" class="Insecta" family="Baetidae" genus="Cloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stelzneri">
<emphasis id="B9B0EAE3D53BFFC9FF6CB043FEA7FCA6" box="[151,298,872,895]" italics="true" pageId="14" pageNumber="15">Cloë stelzneri</emphasis>
<treatmentCitation id="0A6510E0D53BFFC9FECBB043FDBFFCA6" author="Weyenbergh" box="[304,562,872,895]" page="170" pageId="14" pageNumber="15" year="1883">
<bibRefCitation id="EF554B00D53BFFC9FECBB043FE71FCA6" author="Weyenbergh" box="[304,508,872,895]" pageId="14" pageNumber="18" refString="Weyenbergh H. (1883) Bjdrage tot de kennis der Zuid-Amerikaansche Ephemeriden. Tijdschrift voor Entomologie, 26, 159 - 174." type="journal article" year="1883">Weyenbergh, 1883</bibRefCitation>
: 170
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7B36F1D53BFFC9FF6CB0A3FD8BFC46" blockId="14.[151,765,872,959]" box="[151,518,904,927]" pageId="14" pageNumber="15">
<taxonomicName id="4CC44D72D53BFFC9FF6CB0A3FEB7FC46" box="[151,314,904,927]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stelzneri">
<emphasis id="B9B0EAE3D53BFFC9FF6CB0A3FEB7FC46" box="[151,314,904,927]" italics="true" pageId="14" pageNumber="15">Baetis stelzneri</emphasis>
</taxonomicName>
; Eaton, 1885: 171.
</paragraph>
</subSubSection>
<subSubSection id="C3DE657AD53BFFC9FF6CB083FD70FC66" box="[151,765,936,959]" pageId="14" pageNumber="15" type="description">
<paragraph id="8B7B36F1D53BFFC9FF6CB083FD70FC66" blockId="14.[151,765,872,959]" box="[151,765,936,959]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="ABD411DFD53BFFC9FF6CB083FD70FC66" box="[151,765,936,959]" pageId="14" pageNumber="15">
<taxonomicName id="4CC44D72D53BFFC9FF6CB083FED6FC66" ID-CoL="5ZC4Y" box="[151,347,936,959]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stelzneri">
<emphasis id="B9B0EAE3D53BFFC9FF6CB083FED6FC66" box="[151,347,936,959]" italics="true" pageId="14" pageNumber="15">Cloeodes stelzneri</emphasis>
</taxonomicName>
; 
<treatmentCitation id="0A6510E0D53BFFC9FE92B083FD74FC66" author="Lugo-Ortiz" box="[361,761,936,959]" page="259" pageId="14" pageNumber="15" year="1999">
<bibRefCitation id="EF554B00D53BFFC9FE92B083FD49FC66" author="Lugo-Ortiz" box="[361,708,936,959]" pageId="14" pageNumber="17" refString="Lugo-Ortiz C. R. &amp; McCafferty W. P. (1999) Revision of South American species of Baetidae (Ephemeroptera) previously placed in Baetis Leach and Pseudocloeon Klapalek. Annales of Limnologie, 35 (4), 257 - 262." type="journal article" year="1999">Lugo-Ortiz &amp; McCafferty, 1999</bibRefCitation>
: 259
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3DE657AD53BFFC9FF6CB0C4FF7BFA93" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B7B36F1D53BFFC9FF6CB0C4FF7BFA93" blockId="14.[151,1436,1007,1394]" pageId="14" pageNumber="15">
<emphasis id="B9B0EAE3D53BFFC9FF6CB0C4FEAFFBD0" bold="true" box="[151,290,1007,1033]" pageId="14" pageNumber="15">Discussion.</emphasis>
This species was originally described by 
<bibRefCitation id="EF554B00D53BFFC9FCF5B0DBFC7AFBD3" author="Weyenbergh" box="[782,1015,1008,1034]" pageId="14" pageNumber="18" refString="Weyenbergh H. (1883) Bjdrage tot de kennis der Zuid-Amerikaansche Ephemeriden. Tijdschrift voor Entomologie, 26, 159 - 174." type="journal article" year="1883">Weyenbergh (1883)</bibRefCitation>
from adults from Santa Fe and was placed in 
<taxonomicName id="4CC44D72D53BFFC9FEF7B733FECEFBE8" box="[268,323,1048,1073]" class="Insecta" family="Baetidae" genus="Cloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B9B0EAE3D53BFFC9FEF7B733FECEFBE8" box="[268,323,1048,1073]" italics="true" pageId="14" pageNumber="15">Cloë</emphasis>
</taxonomicName>
. Eaton (1885) transferred this species to the genus 
<taxonomicName id="4CC44D72D53BFFC9FC55B733FC7AFBE8" box="[942,1015,1048,1073]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B9B0EAE3D53BFFC9FC55B733FC7AFBE8" box="[942,1015,1048,1073]" italics="true" pageId="14" pageNumber="15">Baetis</emphasis>
</taxonomicName>
, but he worked with material from Córdoba, 
<collectingCountry id="F3D37661D53BFFC9FEF1B76BFEF3FB83" box="[266,382,1088,1114]" name="Argentina" pageId="14" pageNumber="15">Argentina</collectingCountry>
(see Eaton, 1885: 171). Later, 
<bibRefCitation id="EF554B00D53BFFC9FD1DB76BFBE1FB83" author="Lugo-Ortiz" box="[742,1132,1088,1114]" pageId="14" pageNumber="17" refString="Lugo-Ortiz C. R. &amp; McCafferty W. P. (1999) Revision of South American species of Baetidae (Ephemeroptera) previously placed in Baetis Leach and Pseudocloeon Klapalek. Annales of Limnologie, 35 (4), 257 - 262." type="journal article" year="1999">Lugo-Ortiz &amp; McCafferty (1999)</bibRefCitation>
concluded, based on size, distribution, and possession of a small lancetlike costal process on the hind wings, that this species belonged to 
<taxonomicName id="4CC44D72D53BFFC9FF43B7BBFEA8FB70" box="[184,293,1168,1193]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B9B0EAE3D53BFFC9FF43B7BBFEA8FB70" box="[184,293,1168,1193]" italics="true" pageId="14" pageNumber="15">Cloeodes</emphasis>
</taxonomicName>
. They also suggested a possible synonymy between this species and 
<taxonomicName id="4CC44D72D53BFFC9FB94B7BBFAAAFB70" box="[1135,1319,1168,1193]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B9B0EAE3D53BFFC9FB94B7BBFAAAFB70" box="[1135,1319,1168,1193]" italics="true" pageId="14" pageNumber="15">Cloeodes penai</emphasis>
</taxonomicName>
based on similarity of the hind wings. However, this single characteristic is not sufficient for the synonymy of the two species. Moreover, the 
<typeStatus id="547F8853D53BFFC9FE5DB7CBFE55FB23" box="[422,472,1248,1274]" pageId="14" pageNumber="15">type</typeStatus>
material is apparently lost (
<bibRefCitation id="EF554B00D53BFFC9FCE6B7CBFB15FB23" author="Lugo-Ortiz" box="[797,1176,1248,1274]" pageId="14" pageNumber="17" refString="Lugo-Ortiz C. R. &amp; McCafferty W. P. (1999) Revision of South American species of Baetidae (Ephemeroptera) previously placed in Baetis Leach and Pseudocloeon Klapalek. Annales of Limnologie, 35 (4), 257 - 262." type="journal article" year="1999">Lugo-Ortiz &amp; McCafferty, 1999</bibRefCitation>
) so it will be impossible to assign fresh material to this species. For these reasons, 
<taxonomicName id="4CC44D72D53BFFC9FC95B623FBCAFAF8" box="[878,1095,1288,1313]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stelzneri">
<emphasis id="B9B0EAE3D53BFFC9FC95B623FBCAFAF8" box="[878,1095,1288,1313]" italics="true" pageId="14" pageNumber="15">Cloeodes stelzneri</emphasis>
</taxonomicName>
is proposed here as a 
<emphasis id="B9B0EAE3D53BFFC9FAB5B623FF62FA90" italics="true" pageId="14" pageNumber="15">nomen dubium</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3DE657AD53BFFC9FF3DB67CFD70FAAB" box="[198,765,1367,1394]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8B7B36F1D53BFFC9FF3DB67CFD70FAAB" blockId="14.[151,1436,1007,1394]" box="[198,765,1367,1394]" pageId="14" pageNumber="15">
<emphasis id="B9B0EAE3D53BFFC9FF3DB67CFEEAFAA8" bold="true" box="[198,359,1367,1393]" pageId="14" pageNumber="15">Distribution.</emphasis>
<collectingCountry id="F3D37661D53BFFC9FE96B673FE6BFAAB" box="[365,486,1368,1394]" name="Argentina" pageId="14" pageNumber="15">Argentina</collectingCountry>
: Santa Fe and Córdoba.
</paragraph>
</subSubSection>
</treatment>
</document>