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<document id="374980C5B79CFF0F0615DDFC2ED7DC69" ID-ISSN="0024-4082" ID-ZooBank="AE201413-3845-4F95-8E92-30C5C3B46766" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630566657801" checkinUser="plazi" docAuthor="Meregalli, Massimo, Borovec, Roman, Cervella, Piero, Santovito, Alfredo, Toševski, Ivo, Ottati, Sara &amp; Nakládal, Oto" docDate="2021" docId="035487F3FFD6FFC7FF3BC90BFDE1F95E" docLanguage="en" docName="zlaa142.pdf" docOrigin="Zoological Journal of the Linnean Society 193" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Yamalaka BOROVEC &amp; MEREGALLI 2021, GEN. NOV." docType="treatment" docUuid="537D2C1A-33B5-4B55-A8DF-F625C0D3F5B6" docUuidSource="ZooBank" docVersion="2" lastPageNumber="108" masterDocId="FF6DFF8BFFDBFFCAFFF5CB26FFC2FF85" masterDocTitle="The Namaini, a new weevil tribe with six new genera from South Africa (Coleoptera: Curculionidae: Entiminae)" masterLastPageNumber="123" masterPageNumber="95" pageNumber="108" updateTime="1699281238044" updateUser="plazi">
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<mods:title id="AB08A682238719317569E9C6C3CC69A5">The Namaini, a new weevil tribe with six new genera from South Africa (Coleoptera: Curculionidae: Entiminae)</mods:title>
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<mods:namePart id="7DB25C386E089D4DCE7BF4B39E15DDD9">Borovec, Roman</mods:namePart>
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<mods:namePart id="D3D5C9D525A73CD5A165B02496F037BD">Cervella, Piero</mods:namePart>
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<mods:namePart id="6DF9CBD3F84832DC16FC5E2FD3BB6F47">Santovito, Alfredo</mods:namePart>
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<mods:namePart id="50AE25FB7F14B89E64FE3A7C23D10F15">Toševski, Ivo</mods:namePart>
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<mods:namePart id="FA67A41BA3C10E9DDCC8BA8543031ABA">Ottati, Sara</mods:namePart>
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<mods:namePart id="0E3A1AA23E28BFCB4034BDD37E431A86">Nakládal, Oto</mods:namePart>
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<mods:title id="87A3BC39E2D3C1B32F7D6335995ED456">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="7DA88F9081BA97A46CF309DE8E0B3CF7">2021</mods:date>
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<treatment id="035487F3FFD6FFC7FF3BC90BFDE1F95E" LSID="urn:lsid:zoobank.org:act:537D2C1A-33B5-4B55-A8DF-F625C0D3F5B6" httpUri="http://treatment.plazi.org/id/035487F3FFD6FFC7FF3BC90BFDE1F95E" lastPageNumber="108" pageId="13" pageNumber="108">
<subSubSection id="C3E7656EFFD6FFC7FF3BC90BFCC6FD1D" pageId="13" pageNumber="108" type="nomenclature">
<paragraph id="8B4236E5FFD6FFC7FF3BC90BFD1DFDC7" blockId="13.[206,735,557,579]" box="[206,735,557,579]" pageId="13" pageNumber="108">
<heading id="D00A8189FFD6FFC7FF3BC90BFD1DFDC7" box="[206,735,557,579]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="108" reason="2">
<taxonomicName id="4CFD4D66FFD6FFC7FF3BC90BFD9CFDC7" ID-CoL="8KGD4" authority="BOROVEC &amp; MEREGALLI" authorityName="BOROVEC &amp; MEREGALLI" authorityYear="2021" box="[206,606,557,579]" class="Insecta" family="Curculionidae" genus="Yamalaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B989EAF7FFD6FFC7FF3BC90BFE88FDC7" bold="true" box="[206,330,557,578]" italics="true" pageId="13" pageNumber="108">YAMALAKA</emphasis>
BOROVEC &amp; MEREGALLI
</taxonomicName>
,
<emphasis id="B989EAF7FFD6FFC7FD9EC917FD1DFDC7" bold="true" box="[619,735,557,578]" pageId="13" pageNumber="108">
<taxonomicNameLabel id="A2BA578CFFD6FFC7FD9EC917FD1DFDC7" box="[619,735,557,578]" pageId="13" pageNumber="108" rank="genus">GEN. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B4236E5FFD6FFC7FF56C943FCC6FD1D" blockId="13.[163,779,612,664]" pageId="13" pageNumber="108">
<emphasis id="B989EAF7FFD6FFC7FF56C943FE26FDFF" box="[163,484,612,634]" italics="true" pageId="13" pageNumber="108">Z o o b a n k r e g i s t r a t i o n</emphasis>
: u r n: l s i d: z o o b a n k. org:act:
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</paragraph>
</subSubSection>
<subSubSection id="C3E7656EFFD6FFC7FF56C99CFDE1F95E" pageId="13" pageNumber="108" type="materials_examined">
<paragraph id="8B4236E5FFD6FFC7FF56C99CFD38FD68" blockId="13.[163,780,697,750]" pageId="13" pageNumber="108">
<typeStatus id="54468847FFD6FFC7FF56C99CFF19FD4A" box="[163,219,698,719]" pageId="13" pageNumber="108">Type</typeStatus>
species:
<taxonomicName id="4CFD4D66FFD6FFC7FECAC99CFF20FD6B" authority="Borovec &amp; Meregalli, 2013: p. 508" authorityName="Borovec &amp; Meregalli" authorityPageNumber="508" authorityYear="2013" class="Magnoliopsida" family="Namaceae" genus="Nama" kingdom="Plantae" order="Boraginales" pageId="13" pageNumber="108" phylum="Tracheophyta" rank="species" species="erikae">
<emphasis id="B989EAF7FFD6FFC7FECAC99CFE13FD4B" box="[319,465,697,719]" italics="true" pageId="13" pageNumber="108">Nama erikae</emphasis>
<bibRefCitation id="EF6C4B14FFD6FFC7FE2DC99CFCC7FD4B" author="Borovec R &amp; Meregalli M" box="[472,773,697,719]" pageId="13" pageNumber="108" pagination="501 - 515" refId="ref13857" refString="Borovec R, Meregalli M. 2013. Soil insect research in South Africa. 1. A new genus of edaphic weevils with four new species from the Richtersveld National Park (Coleoptera: Curculionidae: Entiminae: Trachyphloeini). Zootaxa 3646: 501 - 515." type="journal article" year="2013">Borovec &amp; Meregalli, 2013</bibRefCitation>
: p.508
</taxonomicName>
, here designated (
<figureCitation id="13C62A60FFD6FFC7FE47C9FEFE35FD6B" box="[434,503,728,750]" captionStart="Figure 6" captionStartId="14.[145,223,1593,1615]" captionTargetBox="[272,1298,196,1552]" captionTargetId="figure-96@14.[271,1298,195,1553]" captionTargetPageId="14" captionText="Figure 6. Yamalaka erikae, paratype. A, body, dorsal view. B, body, lateral view. C, antenna. D, elytra, dorso-lateral view. E, protibia. F, pronotum. G, pronotum, detail of scales at side. H, ventrites, male. I, J, K, rostrum respectively dorsal, dorsolateral and lateral view. Bar: 1 mm." pageId="13" pageNumber="108">Figs 6</figureCitation>
,
<figureCitation id="13C62A60FFD6FFC7FDF1C9FEFDB2FD68" box="[516,624,728,749]" captionStart="Figure 10" captionStartId="19.[163,243,1625,1647]" captionTargetBox="[322,1282,196,1583]" captionTargetId="figure-26@19.[321,1284,195,1585]" captionTargetPageId="19" captionText="Figure 10. Male terminalia. Penis with temones: (A) Cederbergia indecora, paratype, (B) Cervellaea coheni, paratype, (C) Nama richtersveldiana, paratype, (D) Namaquania andreai, paratype, (E) Pentamerica pentamera, paratype, (F) Springbokia sacculus, paratype, (G) Yamalaka erikae, paratype. Scale bars: 0.50 mm. Sternite IX: (H) Cederbergia indecora, paratype, (I) Cervellaea coheni, paratype, (J) Nama richtersveldiana, paratype, (K) Namaquania andreai, paratype, (L) Pentamerica pentamera, paratype, (M) Springbokia sacculus, paratype, (N) Yamalaka erikae, paratype. Scale bars: 0.50 mm. Tegmen: (O) Cederbergia indecora, paratype, (P) Cervellaea coheni, paratype, (Q) Nama richtersveldiana, paratype, (R) Namaquania andreai, paratype, (S) Pentamerica pentamera, paratype, (T) Springbokia sacculus, paratype, (U) Yamalaka erikae, paratype. Scale bars: 0.25 mm." pageId="13" pageNumber="108">10G, N, U</figureCitation>
,
<figureCitation id="13C62A60FFD6FFC7FD89C9FEFD28FD68" box="[636,746,728,749]" captionStart="Figure 11" captionStartId="20.[145,223,1634,1656]" captionTargetBox="[336,1232,196,1592]" captionTargetId="figure-26@20.[335,1234,195,1594]" captionTargetPageId="20" captionText="Figure 11. Female terminalia. Spermatheca: (A) Cederbergia indecora, paratype, (B) Cervellaea coheni, paratype, (C) Nama richtersveldiana, paratype, (D) Namaquania andreai, paratype, (E) Pentamerica pentamera, paratype, (F) Springbokia sacculus, paratype, (G) Yamalaka erikae, paratype. Scale bars: 0.10 mm. Gonocoxites: (H) Cederbergia indecora, paratype, (I) Cervellaea coheni, paratype, (J) Nama richtersveldiana, paratype, (K) Namaquania andreai, paratype, (L) Pentamerica pentamera, paratype, (M) Springbokia sacculus, paratype, (N) Yamalaka erikae, paratype. Scale bars: 0.25 mm. Sternite VIII: (O) Cederbergia indecora, paratype, (P) Cervellaea coheni, paratype, (Q) Nama richtersveldiana, paratype, (R) Namaquania andreai, paratype, (S) Pentamerica pentamera, paratype, (T) Springbokia sacculus, paratype, (U) Yamalaka erikae, paratype. Scale bars: 0.50 mm." pageId="13" pageNumber="108">11G, N, U</figureCitation>
).
</paragraph>
<paragraph id="8B4236E5FFD6FFC7FF56C829FD10FB11" blockId="13.[163,779,782,1172]" pageId="13" pageNumber="108">
<emphasis id="B989EAF7FFD6FFC7FF56C829FE68FCA6" box="[163,426,782,804]" italics="true" pageId="13" pageNumber="108">Diagnostic description</emphasis>
: Very small to small
<taxonomicName id="4CFD4D66FFD6FFC7FD51C828FCC8FCA1" authorityName="BOROVEC &amp; MEREGALLI" authorityYear="2021" box="[676,778,782,804]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="tribe" tribe="Namaini">Namaini</taxonomicName>
<quantity id="4C059B00FFD6FFC7FF56C80BFEE8FCC6" box="[163,298,813,835]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="13" pageNumber="108" unit="mm" value="2.05" valueMax="2.4" valueMin="1.7">1.7–2.4 mm</quantity>
long; rostrum continuous with head with U-shaped stria on epifrons; ventral border of rostrum in profile short, conspicuously shorter than rostral thickness; frons squamose; gena and subgena densely squamose; antennal sockets in dorsal view narrowly reniform, laterally subtriangular, longer than wide, not reaching eyes; procoxal cavities placed in middle of prosternum; metatibiae with apical surface densely squamose; ventrites squamose; tegmen with parameres; female sternite VIII with long and slender apodeme, terminating just inside of plate, plate oval to widely oval; gonocoxites with long apical styli.
</paragraph>
<paragraph id="8B4236E5FFD6FFC7FF56CF93FD63FA24" blockId="13.[163,779,1205,1656]" pageId="13" pageNumber="108">
<emphasis id="B989EAF7FFD6FFC7FF56CF93FEDCFB4F" box="[163,286,1205,1226]" italics="true" pageId="13" pageNumber="108">Etymology</emphasis>
: During our 2013 trip with our colleague Andrea Battisti, we camped one night in an area with several large burrows of an unknown animal, possibly an aardvark [
<taxonomicName id="4CFD4D66FFD6FFC7FEB7CE37FD68FAA2" authority="(Pallas, 1766)" baseAuthorityName="Pallas" baseAuthorityYear="1766" box="[322,682,1296,1319]" class="Mammalia" family="Orycteropodidae" genus="Orycteropus" kingdom="Animalia" order="Tubulidentata" pageId="13" pageNumber="108" phylum="Chordata" rank="species" species="afer">
<emphasis id="B989EAF7FFD6FFC7FEB7CE37FDC0FAA3" box="[322,514,1297,1318]" italics="true" pageId="13" pageNumber="108">Orycteropus afer</emphasis>
(Pallas, 1766)
</taxonomicName>
]. We did not know what it was and the fantasy name
<taxonomicName id="4CFD4D66FFD6FFC7FD6CCE16FCC9FAC3" authorityName="BOROVEC &amp; MEREGALLI" authorityYear="2021" box="[665,779,1328,1350]" class="Insecta" family="Curculionidae" genus="Yamalaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">Yamalaka</taxonomicName>
was coined. Since then we always think of a
<taxonomicName id="4CFD4D66FFD6FFC7FD6CCE68FCC9FAE1" authorityName="BOROVEC &amp; MEREGALLI" authorityYear="2021" box="[665,779,1358,1380]" class="Insecta" family="Curculionidae" genus="Yamalaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">Yamalaka</taxonomicName>
following and protecting us during our expeditions. We name this genus after our fictional protector.
</paragraph>
<paragraph id="8B4236E5FFD6FFC7FF4ECE8CFDB1F9FD" blockId="13.[163,779,1205,1656]" pageId="13" pageNumber="108">
<emphasis id="B989EAF7FFD6FFC7FF4ECE8CFDE8FA3A" box="[187,554,1450,1472]" italics="true" pageId="13" pageNumber="108">Included taxa and distribution</emphasis>
: In addition to the
<typeStatus id="54468847FFD6FFC7FF56CEEFFF16FA5B" box="[163,212,1481,1502]" pageId="13" pageNumber="108">type</typeStatus>
species,
<taxonomicName id="4CFD4D66FFD6FFC7FEC3CEEFFF25FA78" authority="(Borovec &amp; Meregalli, 2013)" authorityName="Meregalli &amp; Borovec &amp; Cervella &amp; Santovito &amp; Toševski &amp; Ottati &amp; Nakládal" authorityYear="2021" baseAuthorityName="Borovec &amp; Meregalli" baseAuthorityYear="2013" class="Insecta" family="Curculionidae" genus="Yamalaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="erikae" status="comb. nov.">
<emphasis id="B989EAF7FFD6FFC7FEC3CEEFFDDCFA5A" bold="true" box="[310,542,1481,1503]" pageId="13" pageNumber="108">
<emphasis id="B989EAF7FFD6FFC7FEC3CEEFFDCCFA5A" bold="true" box="[310,526,1481,1503]" italics="true" pageId="13" pageNumber="108">Yamalaka erikae</emphasis>
(
</emphasis>
<bibRefCitation id="EF6C4B14FFD6FFC7FDEBCEEFFF1FFA78" author="Borovec R &amp; Meregalli M" pageId="13" pageNumber="108" pagination="501 - 515" refId="ref13857" refString="Borovec R, Meregalli M. 2013. Soil insect research in South Africa. 1. A new genus of edaphic weevils with four new species from the Richtersveld National Park (Coleoptera: Curculionidae: Entiminae: Trachyphloeini). Zootaxa 3646: 501 - 515." type="journal article" year="2013">Borovec &amp; Meregalli, 2013</bibRefCitation>
<emphasis id="B989EAF7FFD6FFC7FF28CEC1FF25FA78" bold="true" box="[221,231,1511,1533]" pageId="13" pageNumber="108">)</emphasis>
</taxonomicName>
<emphasis id="B989EAF7FFD6FFC7FF07CECEFE43FA78" bold="true" box="[242,385,1512,1533]" pageId="13" pageNumber="108">
<taxonomicNameLabel id="A2BA578CFFD6FFC7FF07CECEFE43FA78" box="[242,385,1512,1533]" pageId="13" pageNumber="108" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4CFD4D66FFD6FFC7FE66CEC1FEA2F999" authority="(Borovec &amp; Meregalli, 2013)" authorityName="Meregalli &amp; Borovec &amp; Cervella &amp; Santovito &amp; Toševski &amp; Ottati &amp; Nakládal" authorityYear="2021" baseAuthorityName="Borovec &amp; Meregalli" baseAuthorityYear="2013" class="Insecta" family="Curculionidae" genus="Yamalaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="iuliae" status="comb. nov.">
<emphasis id="B989EAF7FFD6FFC7FE66CEC1FD4EFA78" bold="true" box="[403,652,1511,1533]" pageId="13" pageNumber="108">
<emphasis id="B989EAF7FFD6FFC7FE66CEC1FDB5FA78" bold="true" box="[403,631,1511,1533]" italics="true" pageId="13" pageNumber="108">Yamalaka iuliae</emphasis>
(
</emphasis>
<bibRefCitation id="EF6C4B14FFD6FFC7FD79CECEFE94F99E" author="Borovec R &amp; Meregalli M" pageId="13" pageNumber="108" pagination="501 - 515" refId="ref13857" refString="Borovec R, Meregalli M. 2013. Soil insect research in South Africa. 1. A new genus of edaphic weevils with four new species from the Richtersveld National Park (Coleoptera: Curculionidae: Entiminae: Trachyphloeini). Zootaxa 3646: 501 - 515." type="journal article" year="2013">Borovec &amp; Meregalli, 2013</bibRefCitation>
<emphasis id="B989EAF7FFD6FFC7FEA3CD20FEA2F999" bold="true" box="[342,352,1542,1564]" pageId="13" pageNumber="108">)</emphasis>
</taxonomicName>
<emphasis id="B989EAF7FFD6FFC7FE9DCD20FE2DF99E" bold="true" box="[360,495,1542,1563]" pageId="13" pageNumber="108">
<taxonomicNameLabel id="A2BA578CFFD6FFC7FE9DCD20FE2DF99E" box="[360,495,1542,1563]" pageId="13" pageNumber="108" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and five additional taxa are included in
<emphasis id="B989EAF7FFD6FFC7FE99CD03FE2DF9BF" box="[364,495,1573,1594]" italics="true" pageId="13" pageNumber="108">
<taxonomicName id="4CFD4D66FFD6FFC7FE99CD03FE28F9BF" authorityName="BOROVEC &amp; MEREGALLI" authorityYear="2021" box="[364,490,1573,1594]" class="Insecta" family="Curculionidae" genus="Yamalaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">Yamalaka</taxonomicName>
,
</emphasis>
based on the
<emphasis id="B989EAF7FFD6FFC7FD50CD03FCC9F9BF" box="[677,779,1573,1594]" italics="true" pageId="13" pageNumber="108">mt-Cox1</emphasis>
analysis and/or their morphology. They are distributed in
<collectingCountry id="F3EA7675FFD6FFC7FF35CD44FE90F9FD" box="[192,338,1634,1656]" name="South Africa" pageId="13" pageNumber="108">South Africa</collectingCountry>
:
<collectingRegion id="4939F807FFD6FFC7FEA9CD44FDC5F9F2" box="[348,519,1634,1656]" country="South Africa" name="Northern Cape" pageId="13" pageNumber="108">Northern Cape</collectingRegion>
(
<figureCitation id="13C62A60FFD6FFC7FDE2CD44FDA1F9FD" box="[535,611,1634,1656]" captionStart="Figure 15" captionStartId="24.[145,224,1560,1582]" captionTargetBox="[273,1297,195,1520]" captionTargetId="figure-134@24.[273,1297,195,1520]" captionTargetPageId="24" captionText="Figure 15. Distribution map of the genus Yamalaka. * = type species. Map data: Google Earth, Maxar Technologies, used according to Google Earth Terms of Service." pageId="13" pageNumber="108">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8B4236E5FFD6FFC7FE7FCDE5FDE1F95E" blockId="13.[394,547,1731,1755]" box="[394,547,1731,1755]" pageId="13" pageNumber="108">SUBCLADE B</paragraph>
</subSubSection>
</treatment>
</document>
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