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<document ID-CLB-Dataset="56876" ID-DOI="http://dx.doi.org/10.3897/asp.81.e96737" ID-GBIF-Dataset="3aaa10b0-eb01-4c20-8857-be51cd7e827d" ID-Pensoft-Pub="1864-8312-81-345" ID-Pensoft-UUID="B69B5606C9B358B9A9027E3E43F2B66D" ID-ZooBank="0C2444B0ECFB4A22ACF6F2ACEC0DDCDF" ModsDocID="1864-8312-81-345" checkinTime="1680679728434" checkinUser="pensoft" docAuthor="Brazidec, Manuel, Legendre, Frederic &amp; Perrichot, Vincent" docDate="2023" docId="028BBE845AE15A7FB432937068EC6412" docLanguage="en" docName="ArthropodSystPhyl 81: 345-369" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-04-04" docSource="http://dx.doi.org/10.3897/asp.81.e96737" docTitle="Gwesped groehni Brazidec &amp; Legendre &amp; Perrichot 2023, sp. nov." docType="treatment" docUuid="C7B02F37-05F4-4122-85FF-D54F1D8482F5" docUuidSource="ZooBank" docVersion="5" id="B69B5606C9B358B9A9027E3E43F2B66D" lastPageNumber="345" masterDocId="B69B5606C9B358B9A9027E3E43F2B66D" masterDocTitle="Diversity and phylogeny of the extinct wasp subfamily Lancepyrinae (Hymenoptera, Bethylidae) revealed by mid-Cretaceous Burmese amber" masterLastPageNumber="369" masterPageNumber="345" pageNumber="345" updateTime="1732742041388" updateUser="ExternalLinkService">
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<mods:title id="F659E45B696CA9D9E552B7185EB86C6F">Diversity and phylogeny of the extinct wasp subfamily Lancepyrinae (Hymenoptera, Bethylidae) revealed by mid-Cretaceous Burmese amber</mods:title>
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<mods:namePart id="0BD296D9C6CB0FE4BAC68A550C2BC942">Brazidec, Manuel</mods:namePart>
<mods:nameIdentifier id="80D77A2B3A13A2BF40D9E9B7CD4237E6" type="ORCID">https://orcid.org/0000-0002-0860-8972</mods:nameIdentifier>
<mods:affiliation id="75D89FB21D73EC5833961B42133F94CA">Univ. Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000 Rennes, France &amp; State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
<mods:nameIdentifier id="70723C3C7FF014FB0ED0DDDE4AAA2263" type="email">manuel.brazidec@gmail.com</mods:nameIdentifier>
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<mods:namePart id="10C8B83B8A2A6873CF2A6796947709B1">Legendre, Frederic</mods:namePart>
<mods:nameIdentifier id="E63645AD38FDEB04ACD6CCD8B10E5484" type="ORCID">https://orcid.org/0000-0001-5900-8048</mods:nameIdentifier>
<mods:affiliation id="9F62CDF53001506BAD5023C312931E70">Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, Sorbonne Universite, EPHE, Universite des Antilles, CP 50, 57, rue Cuvier, 75005 Paris, France</mods:affiliation>
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<mods:namePart id="74EDC26A443F7B3B91238DC07716D0DA">Perrichot, Vincent</mods:namePart>
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<mods:affiliation id="55E1B60BBFBB00648FBB67949E6F1C4B">Univ. Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000 Rennes, France</mods:affiliation>
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<mods:title id="D4C29AB58A64064B06C670A6AE120A15">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date id="88DC503FFD3430584D73CA4CFB6BE381">2023</mods:date>
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<mods:number id="D93704A116BA27D576CCC311ADAD3178">81</mods:number>
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<treatment id="028BBE845AE15A7FB432937068EC6412" ID-GBIF-Taxon="209487375" LSID="urn:lsid:zoobank.org:act:C7B02F37-05F4-4122-85FF-D54F1D8482F5" httpUri="http://treatment.plazi.org/id/028BBE845AE15A7FB432937068EC6412" lastPageNumber="345" pageId="0" pageNumber="345" scope_family="Bethylidae" scope_order="Hymenoptera" scope_subFamily="Lancepyrinae" scope_superFamily="Chrysidoidea">
<subSubSection id="2D33C0D8E4D21AE4726CF5CBDEB4716F" pageId="0" pageNumber="345" type="nomenclature">
<paragraph id="BD791D2CDEF255C73F0AE0DD2435F877" pageId="0" pageNumber="345">
<taxonomicName id="FDEB390FC63B0B3CC043C59311EEE5C9" LSID="https://zoobank.org/C7B02F37-05F4-4122-85FF-D54F1D8482F5" authority="Brazidec &amp; Legendre &amp; Perrichot, 2023" authorityName="Brazidec &amp; Legendre &amp; Perrichot" authorityYear="2023" class="Insecta" family="Bethylidae" genus="Gwesped" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gwesped groehni" order="Hymenoptera" pageId="0" pageNumber="345" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">Gwesped groehni</taxonomicName>
<taxonomicNameLabel id="6D84D5A524FDC4455F6952D7F090A9A6" pageId="0" pageNumber="345">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="078DAD5FAAD00D5F02196009006F60DA" pageId="0" pageNumber="345" type="description">
<paragraph id="B11206DD4FF8D118E658128982E75FD3" pageId="0" pageNumber="345">
<figureCitation id="6C6FAD7E0F199294AD3DCB6CF9CBC2FA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gwesped groehni gen. et sp. nov., holotype female number GPIH. 5068. A habitus in dorsal view; B habitus in ventral view (arrow = sting); C head in dorsal view; D head in ventral view; E fore wing in dorsal view; F line drawing of fore wing. Scale bars: 1 mm (A, B, E, F); 0.5 mm (C, D). Note: The apparent structure at the apex of the metasoma is the result of a bubble formed in the amber, combined with some artifacts produced during the stacking process." figureDoi="10.3897/asp.81.e96737.figure3" httpUri="https://binary.pensoft.net/fig/833458" pageId="0" pageNumber="345">Figs 3</figureCitation>
<figureCitation id="436D242F85A81A42D4947E866B1DB023" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Line drawing of Gwesped groehni gen. et sp. nov., holotype female number GPIH. 5068. Scale bar: 1 mm." figureDoi="10.3897/asp.81.e96737.figure4" httpUri="https://binary.pensoft.net/fig/833459" pageId="0" pageNumber="345">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="85E04EB089A90EE2B9530DBF753EF254" pageId="0" pageNumber="345" type="etymology">
<paragraph id="CF935A87184613184E333612BFD502D4" pageId="0" pageNumber="345">Etymology.</paragraph>
<paragraph id="4631B3C69662D9BB97A9761CB6CD0CA2" pageId="0" pageNumber="345">
The specific epithet is a patronym honouring Carsten
<normalizedToken id="1B4E6F1CE865AA113F21088D604E2046" originalValue="Gröhn">Groehn</normalizedToken>
, who generously made the specimen available for study. The specific epithet is to be treated as a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="81F8C3B0E3D09F47238D9054F96DC4E3" pageId="0" pageNumber="345" type="materials_examined">
<paragraph id="361FA2412CE0333F8BD2D7B695F31795" pageId="0" pageNumber="345">Material studied.</paragraph>
<paragraph id="F3A64460A135F9681809632866D973FC" pageId="0" pageNumber="345">
<materialsCitation id="EE8B3EF47C51AE381B650C355220A9F4" ID-GBIF-Occurrence="4071676303" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Germany" county="Analysis of Biodiversity Change" location="Carsten Groehn" municipality="Leibniz Institute" specimenCount="1" stateProvince="Hamburg" typeStatus="Holotype">
<emphasis id="45845FD55532ABF479DDA761A923B976" bold="true" italics="true" pageId="0" pageNumber="345">
<typeStatus id="9C0C9AB3AE707A9368F32F06F6AFFEEE">Holotype</typeStatus>
</emphasis>
GPIH.5068, a complete female; housed in the
<location id="F6373F6F9720A3605CBD536CFC909888" LSID="urn:lsid:plazi:treatment:028BBE845AE15A7FB432937068EC6412:F6373F6F9720A3605CBD536CFC909888" country="Germany" county="Analysis of Biodiversity Change" municipality="Leibniz Institute" name="Carsten Groehn" stateProvince="Hamburg">
Carsten
<normalizedToken id="956B7EDDA147EA051C9C33D06FCECD6C" originalValue="Gröhn">Groehn</normalizedToken>
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amber collection (under CCGG no. 11341) of the
<collectingMunicipality id="EA62D38BBE04A23FEEAA6F84675F13F0">Leibniz Institute</collectingMunicipality>
for the
<collectingCounty id="8B3E13CD2C93BC132158F36165C2B648">Analysis of Biodiversity Change</collectingCounty>
,
<collectingRegion id="6A8B112B1C3FDEEC3FE43318F9F2075C" country="Germany" name="Hamburg">Hamburg</collectingRegion>
,
<collectingCountry id="9BAA21C60AFF624BAAA4EE70E6A581B3" name="Germany">Germany</collectingCountry>
(GPIH).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9C186F5C420E0680997F753157099316" pageId="0" pageNumber="345" type="type locality">
<paragraph id="4E8B6F548DE1E3166E3FA9BBF90814D3" pageId="0" pageNumber="345">Type locality and horizon.</paragraph>
<paragraph id="5936AA779BA62726847C5A15FEB230CD" pageId="0" pageNumber="345">Noije Bum, Hukawng Valley, Kachin State, northern Myanmar; late Albian-early Cenomanian, ca. 99 Ma, mid-Cretaceous.</paragraph>
</subSubSection>
<subSubSection id="A443785B61E5284DA5CC686ABCE67FD5" pageId="0" pageNumber="345" type="diagnosis">
<paragraph id="0BA04CE76C232E24F9593E65B79A7F93" pageId="0" pageNumber="345">Diagnosis.</paragraph>
<paragraph id="C4162262F676B2BE53BC8BE2A323BFD6" pageId="0" pageNumber="345">As for genus.</paragraph>
</subSubSection>
<subSubSection id="EA4DC6995EBF19E0EB521E646A96F32A" pageId="0" pageNumber="345" type="description">
<paragraph id="28E9F54828FFDB52BF5FD6468EE90AAD" pageId="0" pageNumber="345">Description.</paragraph>
<paragraph id="1738696B42908DE3B891B1B444ABCA57" pageId="0" pageNumber="345">
Body depressed and poorly pubescent (length 3.82 mm). -
<emphasis id="EE1DF62E264F99F709B600FDD081F031" bold="true" pageId="0" pageNumber="345">Head</emphasis>
prognathous, quadrate, about as long as wide, with metallic reflections; LH: 0.65 mm, WH: 0.56 mm; WF: 0.39 mm, HE: ca. 0.19 mm; frons flat, slightly punctate; compound eye rounded, located anteriorly on head; clypeus with median lobe only slightly projecting forward, lateral lobe poorly developed; antenna short, barely reaching pronotum posteriorly; scape 2.2 times as long as pedicel (length 0.20 mm); pedicel longer than flagellomeres 1-9; 10 flagellomeres, all compact, as long as wide; flagellomere tapering at apex, flagellomere 10 slightly longer than flagellomeres 1-9 (0.05 mm vs. 0.11 mm); occipital carina present. -
<emphasis id="7532479701E39BB69FAC1ABEC4C21E3A" bold="true" pageId="0" pageNumber="345">Mesosoma</emphasis>
narrow with dorsum smooth (length 1.56 mm); propleuron elongate,
<normalizedToken id="F82B9A152D564A701C5B7EDFE531E88C" originalValue="neck-shaped">'neck-shaped'</normalizedToken>
, visible dorsally; dorsal pronotal area narrower anteriorly, lateral margin slightly incurved, posterior margin straight; anteromesoscutum short, notaulus and parapsidal signum present; mesoscuto-mesoscutellar sulcus conspicuous, widely connecting lateral grooves; mesoscutellum subquadrate; metanotum developed, overlapping mesoscutellum posteriorly; metapectal-propodeal complex with metapostnotal median carina, lateral margin slightly outcurved, posterior corner with small lateral dentiform projection. Fore wing hyaline, posteriorly reaching fourth metasomal segment (LFW = 2.19 mm); C, Sc+R, M+Cu, 1A tubular; 1Rs and 1M aligned; cu-a slightly post-furcal to 1M; Rs+M tubular, reaching apex of pterostigma; pterostigma rounded; 2r-rs arising from distal half of pterostigma; 2r-rs&amp;Rs tubular, reaching wing margin; [2R1] cell large; post-stigmal abscissa of R1 tubular for half-length of [2R1] cell. Legs slender; tibial spur configuration 1-2-2; tarsal claws slightly curved; arolium small. -
<emphasis id="9647222202B4A6427FF551F7ECD933EF" bold="true" pageId="0" pageNumber="345">Metasoma</emphasis>
fusiform, smooth (length 1.61 mm); petiole relatively long, conspicuous; six tergites visible, convex; short sting exserted.
</paragraph>
<caption id="2A693105C8353D62CAC648669974CD7F" doi="10.3897/asp.81.e96737.figure4" httpUri="https://binary.pensoft.net/fig/833459" pageId="0" pageNumber="345" start="Figure 4" startId="F4">
<paragraph id="D9BED7E57F4FAA83CD628313E547A9B0" pageId="0" pageNumber="345">
<emphasis id="A3E73480095EDA999BA5C9BA22A475CE" bold="true" pageId="0" pageNumber="345">Figure 4.</emphasis>
Line drawing of
<taxonomicName id="0123098C55E281B6DADEA8775FA6D86E" authorityName="Brazidec &amp; Legendre &amp; Perrichot" authorityYear="2023" class="Insecta" family="Bethylidae" genus="Gwesped" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gwesped groehni" order="Hymenoptera" pageId="0" pageNumber="345" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="DAAB159FCBEF8A64B34F25E4CDB9DC62" italics="true" pageId="0" pageNumber="345">Gwesped groehni</emphasis>
</taxonomicName>
<emphasis id="EB216694144BA190E30E0EC639F8A82D" bold="true" pageId="0" pageNumber="345">gen. et sp. nov.</emphasis>
, holotype female number GPIH.5068. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
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