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<document id="890D904C6A20046B596880A6595FB1A0" ID-DOI="10.11646/zootaxa.3670.2.11" ID-GBIF-Dataset="bc516484-c1c5-4ee3-8912-7ff89f906a8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="219876" ID-ZooBank="67402B31-9D14-4116-8752-2927A1084240" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633918393" checkinUser="plazi" docAuthor="Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando &amp; Oromí, Pedro" docDate="2013" docId="015C614EF400FF955C94FD970463FB22" docLanguage="en" docName="zt03670p299.pdf" docOrigin="Zootaxa 3670 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Roncocreagris occidentalis Zaragoza &amp; Reboleira, sp. nov." docType="treatment" docUuid="6D81C454-B078-46D0-803A-5E29B4B35345" docUuidSource="ZooBank" docVersion="9" lastPageNumber="297" masterDocId="FD651936F40BFF9B5C03FFB30766FFAE" masterDocTitle="On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species" masterLastPageNumber="299" masterPageNumber="283" pageNumber="294" updateTime="1698334986200" updateUser="plazi">
<mods:mods id="C5212AD29A50A56DA45F889B2DE3D5E5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="FD2902173EA00CBC65B4F39FA1BB4D64">On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="048857FC1B636567C73CEA5FF458A573">Reboleira, Ana Sofia P. S.</mods:namePart>
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<mods:namePart id="AAD0DD7022A154BA04733F5D28A5F903">Zaragoza, Juan A.</mods:namePart>
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<mods:namePart id="AF9EB74B251125D85096719927B0E348">Gonçalves, Fernando</mods:namePart>
</mods:name>
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<mods:namePart id="477990048C34CCB7EFD9DEDBF93EA2A3">Oromí, Pedro</mods:namePart>
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<mods:typeOfResource id="1DCDFC34868CF21252B5B9A88904D638">text</mods:typeOfResource>
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<mods:title id="5E6C4F4BAF81E4FDC9A12262019AE3A7">Zootaxa</mods:title>
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<mods:part id="EA2DC2978CFD41E2427F2A3EB5A84ABD">
<mods:date id="94011E701B5D3033CB8B70FB9A644D9D">2013</mods:date>
<mods:detail id="80B1F042346557D7BE19B376808F2E71" type="volume">
<mods:number id="2FA10293068BC9E59FBFB49BEC5C3519">3670</mods:number>
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<mods:identifier id="57D8B17F03703BD98462F6A4335440C3" type="DOI">10.11646/zootaxa.3670.2.11</mods:identifier>
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<treatment id="015C614EF400FF955C94FD970463FB22" ID-DOI="http://doi.org/10.5281/zenodo.6146636" ID-GBIF-Taxon="119570120" ID-Zenodo-Dep="6146636" LSID="urn:lsid:zoobank.org:act:6D81C454-B078-46D0-803A-5E29B4B35345" httpUri="http://treatment.plazi.org/id/015C614EF400FF955C94FD970463FB22" lastPageId="14" lastPageNumber="297" pageId="11" pageNumber="294">
<subSubSection id="C1EF83D3F400FF905C94FD970641FDF0" pageId="11" pageNumber="294" type="nomenclature">
<paragraph id="894AD058F400FF905C94FD970406FD93" blockId="11.[151,864,547,606]" box="[151,864,547,574]" pageId="11" pageNumber="294">
<heading id="D2026734F400FF905C94FD970406FD93" bold="true" box="[151,864,547,574]" fontSize="11" level="1" pageId="11" pageNumber="294" reason="1">
<emphasis id="BB810C4AF400FF905C94FD970406FD93" bold="true" box="[151,864,547,574]" pageId="11" pageNumber="294">
<taxonomicName id="4EF5ABDBF400FF905C94FD97059AFD93" ID-CoL="4TB4G" authority="Zaragoza &amp; Reboleira" authorityName="Zaragoza &amp; Reboleira" box="[151,764,547,574]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="BB810C4AF400FF905C94FD9706BBFD90" bold="true" box="[151,477,548,574]" italics="true" pageId="11" pageNumber="294">Roncocreagris occidentalis</emphasis>
Zaragoza &amp; Reboleira
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F400FF905F00FD900406FD93" box="[771,864,547,573]" pageId="11" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="894AD058F400FF905C94FDF50641FDF0" blockId="11.[151,864,547,606]" box="[151,295,582,606]" pageId="11" pageNumber="294">
(
<figureCitation id="11CECCDDF400FF905C9CFDF50678FDF0" box="[159,286,582,606]" captionStart-0="FIGURE 33" captionStart-1="FIGURES 34 – 43" captionStartId-0="11.[151,255,1349,1373]" captionStartId-1="12.[151,269,1870,1894]" captionTargetBox-0="[152,1436,877,1326]" captionTargetBox-1="[159,1434,199,1869]" captionTargetId-0="figure@11.[151,1436,877,1328]" captionTargetId-1="figure@12.[151,1436,194,1869]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 33. Roncocreagris occidentalis sp. nov., Algar do Javali, Montejunto massif, habitus, dorsal view. A. living specimen; B. preserved specimen." captionText-1="FIGURES 34 – 43. Roncocreagris occidentalis sp. nov., female holotype. 34. carapace; 35. anterior margin of carapace; 36. anterior and medial processes of right coxa I; 37. left chelicera; 38. partial view of fingers of left chelicera, with spinneret; 39. left pedipalp minus chela, dorsal view; 40. left chela, dorsal view; 41. left chela, lateral view; 42. claw of left leg IV; 43. subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39 – 41)." httpUri-0="https://zenodo.org/record/219882/files/figure.png" httpUri-1="https://zenodo.org/record/219883/files/figure.png" pageId="11" pageNumber="294">Figs 33–43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F400FF905C94FD3C03F9FC98" pageId="11" pageNumber="294" type="materials_examined">
<paragraph id="894AD058F400FF905C94FD3C03F9FC98" blockId="11.[151,1436,654,822]" pageId="11" pageNumber="294">
<emphasis id="BB810C4AF400FF905C94FD3C062CFD09" bold="true" box="[151,330,654,679]" pageId="11" pageNumber="294">
<typeStatus id="564E6EFAF400FF905C94FD3C07B6FD09" box="[151,208,655,679]" pageId="11" pageNumber="294">Type</typeStatus>
material.
</emphasis>
<typeStatus id="564E6EFAF400FF905D58FD3D06A5FD08" box="[347,451,654,678]" pageId="11" pageNumber="294" type="holotype">Holotype</typeStatus>
Ƥ,
<collectingCountry id="F1E290C8F400FF905DFFFD3D0538FD08" box="[508,606,654,678]" name="Portugal" pageId="11" pageNumber="294">Portugal</collectingCountry>
, Serra de Montejunto, District of Lisboa, Pragança, Algar do Javali (39º11.965ʹN, 9º01.718ʹW;
<specimenCount id="9FF31BD1F400FF905DCCFD010545FD64" box="[463,547,690,714]" pageId="11" pageNumber="294" type="generic">
<quantity id="4E0D7DBDF400FF905DCCFD010579FD64" box="[463,543,690,714]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="294" unit="m" value="380.0">380 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="FD4BF698F400FF905E6FFD010589FD64" box="[620,751,690,714]" pageId="11" pageNumber="294" value="2009-09-19">19.IX.2009</date>
, lgt. A.S.P.S. Reboleira (
<collectionCode id="EFE4489DF400FF90580FFD010306FD64" box="[1036,1120,690,714]" pageId="11" pageNumber="294">DEUA</collectionCode>
).
<typeStatus id="564E6EFAF400FF905877FD000383FD65" box="[1140,1253,691,715]" pageId="11" pageNumber="294" type="paratype">Paratypes</typeStatus>
: 1 deutonymph, same location as
<typeStatus id="564E6EFAF400FF905D54FD6506DAFD40" box="[343,444,726,750]" pageId="11" pageNumber="294" type="holotype">holotype</typeStatus>
,
<date id="FD4BF698F400FF905DC4FD65052FFD40" box="[455,585,726,750]" pageId="11" pageNumber="294" value="2009-09-19">19.IX.2009</date>
, lgt. A.S.P.S. Reboleira (
<collectionCode id="EFE4489DF400FF905F5CFD6504D5FD40" box="[863,947,726,750]" pageId="11" pageNumber="294">DEUA</collectionCode>
); 1 Ƥ, same location as
<typeStatus id="564E6EFAF400FF9058BDFD650245FD40" box="[1214,1315,726,750]" pageId="11" pageNumber="294" type="holotype">holotype</typeStatus>
,
<date id="FD4BF698F400FF90592DFD6502FDFD40" box="[1326,1435,726,750]" pageId="11" pageNumber="294" value="2009-04-05">5.IV.2009</date>
lgt. A.S.P.S. Reboleira (
<collectionCode id="EFE4489DF400FF905DABFD49069FFCBC" box="[424,505,762,786]" httpUri="http://grbio.org/cool/x2t0-vyyc" name="Departamento de Zoologia, Universidad de La Laguna" pageId="11" pageNumber="294">DZUL</collectionCode>
); 1 Ƥ,
<collectingCountry id="F1E290C8F400FF905E4FFD4905C8FCBC" box="[588,686,762,786]" name="Portugal" pageId="11" pageNumber="294">Portugal</collectingCountry>
, Cesaredas Plateau, District of Lisboa, Bolhos, Gruta dos Bolhos (39°18ʹ31.35&quot;N, 9°16ʹ36.69&quot;W;
<specimenCount id="9FF31BD1F400FF905E07FCAD0530FC98" box="[516,598,798,822]" pageId="11" pageNumber="294" type="generic">
<quantity id="4E0D7DBDF400FF905E07FCAD0534FC98" box="[516,594,798,822]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="11" pageNumber="294" unit="m" value="145.0">145 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="FD4BF698F400FF905E9EFCAD044FFC98" box="[669,809,798,822]" pageId="11" pageNumber="294" value="2010-12-26">26.XII.2010</date>
, lgt. A.S.P.S. Reboleira (
<collectionCode id="EFE4489DF400FF90583DFCAD03F5FC98" box="[1086,1171,798,822]" pageId="11" pageNumber="294">DEUA</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DD8A80D0F400FF905C94FAF60565FAD2" httpUri="https://zenodo.org/record/219882/files/figure.png" pageId="11" pageNumber="294" targetBox="[152,1436,877,1326]" targetPageId="11">
<paragraph id="894AD058F400FF905C94FAF60565FAD2" blockId="11.[151,1436,1349,1405]" pageId="11" pageNumber="294">
<emphasis id="BB810C4AF400FF905C94FAF6064EFAF3" bold="true" box="[151,296,1349,1373]" pageId="11" pageNumber="294">FIGURE 33.</emphasis>
<taxonomicName id="4EF5ABDBF400FF905D32FAF50530FAF3" box="[305,598,1350,1373]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="BB810C4AF400FF905D32FAF50530FAF3" box="[305,598,1350,1373]" italics="true" pageId="11" pageNumber="294">Roncocreagris occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F400FF905E5DFAF605D0FAF3" box="[606,694,1349,1373]" pageId="11" pageNumber="294" rank="species">
<emphasis id="BB810C4AF400FF905E5DFAF605D0FAF3" bold="true" box="[606,694,1349,1373]" pageId="11" pageNumber="294">sp. nov.</emphasis>
</taxonomicNameLabel>
, Algar do Javali, Montejunto massif, habitus, dorsal view.
<emphasis id="BB810C4AF400FF905938FAF60233FAF3" bold="true" box="[1339,1365,1349,1373]" pageId="11" pageNumber="294">A.</emphasis>
living specimen;
<emphasis id="BB810C4AF400FF905D09FAD60644FAD3" bold="true" box="[266,290,1381,1405]" pageId="11" pageNumber="294">B.</emphasis>
preserved specimen.
</paragraph>
</caption>
<subSubSection id="C1EF83D3F400FF905CC4FA150624FA4F" pageId="11" pageNumber="294" type="etymology">
<paragraph id="894AD058F400FF905CC4FA150624FA4F" blockId="11.[151,1437,1446,1979]" pageId="11" pageNumber="294">
<emphasis id="BB810C4AF400FF905CC4FA150628FA10" bold="true" box="[199,334,1446,1470]" pageId="11" pageNumber="294">Etymology.</emphasis>
Named in reference to the westernmost location known for a hypogean species of the genus
<taxonomicName id="4EF5ABDBF400FF905C94FA78065DFA4C" box="[151,315,1483,1506]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="BB810C4AF400FF905C94FA78065DFA4C" box="[151,315,1483,1506]" italics="true" pageId="11" pageNumber="294">Roncocreagris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F400FF905CC4FA5E03CBF9D9" pageId="11" pageNumber="294" type="diagnosis">
<paragraph id="894AD058F400FF905CC4FA5E03CBF9D9" blockId="11.[151,1437,1446,1979]" pageId="11" pageNumber="294">
<emphasis id="BB810C4AF400FF905CC4FA5E0627F9A8" bold="true" box="[199,321,1517,1542]" pageId="11" pageNumber="294">Diagnosis.</emphasis>
No eyes or eye-spots. Strongly troglomorphic. Carapace with 20 setae, 6 on posterior margin. Tergite
<date id="FD4BF698F400FF905CEEF9A1064EF987" box="[237,296,1553,1577]" pageId="11" pageNumber="294">I 6–7</date>
setae. Female cheliceral spinneret reduced to a hyaline dome. Pedipalp femur ratio about 5.7, femur shorter than the movable finger; chela+ ratio about 5.5; chelal hand broadest proximad of middle, ratio movable finger/hand+ about 1.4; trichobothium
<emphasis id="BB810C4AF400FF905E47F9D30539F9D9" box="[580,607,1632,1655]" italics="true" pageId="11" pageNumber="294">ist</emphasis>
close to finger base, ratio
<emphasis id="BB810C4AF400FF905F84F9EC04A0F9D9" box="[903,966,1631,1655]" italics="true" pageId="11" pageNumber="294">ib–ist</emphasis>
/
<emphasis id="BB810C4AF400FF905FCEF9D30362F9D9" box="[973,1028,1632,1655]" italics="true" pageId="11" pageNumber="294">ist–it</emphasis>
about 0.5–0.6.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F400FF955CC4F930037DFDE2" lastPageId="14" lastPageNumber="297" pageId="11" pageNumber="294" type="description">
<paragraph id="894AD058F400FF905CC4F93006D8F94D" blockId="11.[151,1437,1446,1979]" pageId="11" pageNumber="294">
<emphasis id="BB810C4AF400FF905CC4F930063FF932" bold="true" box="[199,345,1667,1692]" pageId="11" pageNumber="294">Description.</emphasis>
<emphasis id="BB810C4AF400FF905D6AF9300327F935" box="[361,1089,1667,1691]" italics="true" pageId="11" pageNumber="294">
Female
<typeStatus id="564E6EFAF400FF905DCDF9300557F935" box="[462,561,1667,1691]" pageId="11" pageNumber="294" type="holotype">holotype</typeStatus>
, followed by female
<typeStatus id="564E6EFAF400FF905F3EF93704CAF935" box="[829,940,1668,1691]" pageId="11" pageNumber="294" type="paratype">paratypes</typeStatus>
in brackets
</emphasis>
. Large pseudoscorpion with troglomorphic adaptations (
<figureCitation id="11CECCDDF400FF905DCFF9140546F911" box="[460,544,1703,1727]" captionStart="FIGURE 33" captionStartId="11.[151,255,1349,1373]" captionTargetBox="[152,1436,877,1326]" captionTargetId="figure@11.[151,1436,877,1328]" captionTargetPageId="11" captionText="FIGURE 33. Roncocreagris occidentalis sp. nov., Algar do Javali, Montejunto massif, habitus, dorsal view. A. living specimen; B. preserved specimen." httpUri="https://zenodo.org/record/219882/files/figure.png" pageId="11" pageNumber="294">Fig. 33</figureCitation>
). Opisthosoma elongate. Opisthosoma and legs yellowish. Carapace, chelicerae and pedipalps pale brown.
</paragraph>
<paragraph id="894AD058F400FF905CC4F95C04BEF8E1" blockId="11.[151,1437,1446,1979]" pageId="11" pageNumber="294">
Carapace (
<figureCitation id="11CECCDDF400FF905D3CF95C06A7F8A9" box="[319,449,1775,1799]" captionStart="FIGURES 34 – 43" captionStartId="12.[151,269,1870,1894]" captionTargetBox="[159,1434,199,1869]" captionTargetId="figure@12.[151,1436,194,1869]" captionTargetPageId="12" captionText="FIGURES 34 – 43. Roncocreagris occidentalis sp. nov., female holotype. 34. carapace; 35. anterior margin of carapace; 36. anterior and medial processes of right coxa I; 37. left chelicera; 38. partial view of fingers of left chelicera, with spinneret; 39. left pedipalp minus chela, dorsal view; 40. left chela, dorsal view; 41. left chela, lateral view; 42. claw of left leg IV; 43. subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39 – 41)." httpUri="https://zenodo.org/record/219883/files/figure.png" pageId="11" pageNumber="294">Figs 34–35</figureCitation>
) longer than broad. Without eyes or eye-spots. Anterior margin moderately prominent medially, epistome blunt and almost indistinguishable. Chaetotaxy: 20 setae, formula 4:4:6:6. Three (four) microlyrifissures in ocular zone, two between median and posterior zones.
</paragraph>
<paragraph id="894AD058F400FF905CC4F8E805F9F815" blockId="11.[151,1437,1446,1979]" pageId="11" pageNumber="294">
Coxal area. Manducatory process with 3 setae. Anterior process of coxa I simple, long and apically pointed; medial process straight with some small, blunt protuberances (
<figureCitation id="11CECCDDF400FF905F5EF8CC04D5F839" box="[861,947,1919,1943]" captionStart="FIGURES 34 – 43" captionStartId="12.[151,269,1870,1894]" captionTargetBox="[159,1434,199,1869]" captionTargetId="figure@12.[151,1436,194,1869]" captionTargetPageId="12" captionText="FIGURES 34 – 43. Roncocreagris occidentalis sp. nov., female holotype. 34. carapace; 35. anterior margin of carapace; 36. anterior and medial processes of right coxa I; 37. left chelicera; 38. partial view of fingers of left chelicera, with spinneret; 39. left pedipalp minus chela, dorsal view; 40. left chela, dorsal view; 41. left chela, lateral view; 42. claw of left leg IV; 43. subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39 – 41)." httpUri="https://zenodo.org/record/219883/files/figure.png" pageId="11" pageNumber="294">Fig. 36</figureCitation>
). Pedipalpal coxa with 7 (6–8) setae, pedal coxa I with 7 (7–8), II: 6–7, III: 4 (5), IV: 6–7.
</paragraph>
<caption id="DD8A80D0F407FF975C94F8FD054BF85F" httpUri="https://zenodo.org/record/219883/files/figure.png" pageId="12" pageNumber="295" targetBox="[159,1434,199,1869]" targetPageId="12">
<paragraph id="894AD058F407FF975C94F8FD054BF85F" blockId="12.[151,1436,1870,2033]" pageId="12" pageNumber="295">
<emphasis id="BB810C4AF407FF975C94F8FD0638F8C8" bold="true" box="[151,350,1870,1894]" pageId="12" pageNumber="295">FIGURES 34–43.</emphasis>
<taxonomicName id="4EF5ABDBF407FF975D66F8FC05ECF8C8" box="[357,650,1871,1894]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="295" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="BB810C4AF407FF975D66F8FC05ECF8C8" box="[357,650,1871,1894]" italics="true" pageId="12" pageNumber="295">Roncocreagris occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F407FF975E92F8FD0581F8C8" box="[657,743,1870,1894]" pageId="12" pageNumber="295" rank="species">
<emphasis id="BB810C4AF407FF975E92F8FD0581F8C8" bold="true" box="[657,743,1870,1894]" pageId="12" pageNumber="295">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype.
<emphasis id="BB810C4AF407FF975FB3F8FD04B7F8C8" bold="true" box="[944,977,1870,1894]" pageId="12" pageNumber="295">34.</emphasis>
carapace;
<emphasis id="BB810C4AF407FF975847F8FD0300F8C8" bold="true" box="[1092,1126,1870,1894]" pageId="12" pageNumber="295">35.</emphasis>
anterior margin of carapace;
<emphasis id="BB810C4AF407FF975C94F8C207DEF827" bold="true" box="[151,184,1905,1929]" pageId="12" pageNumber="295">36.</emphasis>
anterior and medial processes of right coxa I;
<emphasis id="BB810C4AF407FF975EBCF8C20587F827" bold="true" box="[703,737,1905,1929]" pageId="12" pageNumber="295">37.</emphasis>
left chelicera;
<emphasis id="BB810C4AF407FF975F8BF8C204CFF827" bold="true" box="[904,937,1905,1929]" pageId="12" pageNumber="295">38.</emphasis>
partial view of fingers of left chelicera, with spinneret;
<emphasis id="BB810C4AF407FF975D04F827064FF802" bold="true" box="[263,297,1940,1964]" pageId="12" pageNumber="295">39.</emphasis>
left pedipalp minus chela, dorsal view;
<emphasis id="BB810C4AF407FF975EDBF827059FF802" bold="true" box="[728,761,1940,1964]" pageId="12" pageNumber="295">40.</emphasis>
left chela, dorsal view;
<emphasis id="BB810C4AF407FF975FFEF8270378F802" bold="true" box="[1021,1054,1940,1964]" pageId="12" pageNumber="295">41.</emphasis>
left chela, lateral view;
<emphasis id="BB810C4AF407FF975920F8270222F802" bold="true" box="[1315,1348,1940,1964]" pageId="12" pageNumber="295">42.</emphasis>
claw of left leg IV;
<emphasis id="BB810C4AF407FF975D11F8050655F860" bold="true" box="[274,307,1974,1998]" pageId="12" pageNumber="295">43.</emphasis>
subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39–41).
</paragraph>
</caption>
<paragraph id="894AD058F406FF965CC4FF240527FE91" blockId="13.[151,1437,151,2005]" pageId="13" pageNumber="296">Tergal chaetotaxy I–X: 7:7:8:9:9:10:10:11:11:10 (6–7:7–8:8:9:9–10:9–10:11:10:10–11:9), most hairs lacking. Genital area with 5 (6) thin setae on sternite II. Sternal chaetotaxy III–X: 10:8:11:13:13:11:11:11 (10:7:11:12:12:11:11:9–10); sternites VI and VII each with 2 discal setae (included in sternal formula); discal setae glandular, with visible duct. Segment XI: 9 setae. Stigmata III and IV with 3 microsetae on each side. Anal cone with two dorsal and two ventral setae.</paragraph>
<paragraph id="894AD058F406FF965CC4FEFF0780FDB9" blockId="13.[151,1437,151,2005]" pageId="13" pageNumber="296">
Chelicera (
<figureCitation id="11CECCDDF406FF965D47FEFF06ACFECA" box="[324,458,332,356]" captionStart="FIGURES 34 – 43" captionStartId="12.[151,269,1870,1894]" captionTargetBox="[159,1434,199,1869]" captionTargetId="figure@12.[151,1436,194,1869]" captionTargetPageId="12" captionText="FIGURES 34 – 43. Roncocreagris occidentalis sp. nov., female holotype. 34. carapace; 35. anterior margin of carapace; 36. anterior and medial processes of right coxa I; 37. left chelicera; 38. partial view of fingers of left chelicera, with spinneret; 39. left pedipalp minus chela, dorsal view; 40. left chela, dorsal view; 41. left chela, lateral view; 42. claw of left leg IV; 43. subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39 – 41)." httpUri="https://zenodo.org/record/219883/files/figure.png" pageId="13" pageNumber="296">Figs 37–38</figureCitation>
). Palm with 6–7 setae, subgaleal seta 0.66 (0.62–0.68) from base of movable finger. Spinneret a low hyaline dome. Fingers with teeth worn (fixed finger with 5 distal denticles and 7–12 small and medium sized teeth; movable finger with 3 distal denticles and 14 teeth, one large medial tooth, the others medium or small, dental row ending proximad of subgaleal seta. Rallum with 8 blades, all unilaterally pinnate on anterior face, basal blade about half length of others. Serrula exterior with 32 (31–33) blades, serrula interior 24 (25–26) blades.
</paragraph>
<paragraph id="894AD058F406FF965CC4FD970230FC6B" blockId="13.[151,1437,151,2005]" pageId="13" pageNumber="296">
Pedipalp (
<figureCitation id="11CECCDDF406FF965D3BFD9706DAFD92" box="[312,444,548,572]" captionStart="FIGURES 34 – 43" captionStartId="12.[151,269,1870,1894]" captionTargetBox="[159,1434,199,1869]" captionTargetId="figure@12.[151,1436,194,1869]" captionTargetPageId="12" captionText="FIGURES 34 – 43. Roncocreagris occidentalis sp. nov., female holotype. 34. carapace; 35. anterior margin of carapace; 36. anterior and medial processes of right coxa I; 37. left chelicera; 38. partial view of fingers of left chelicera, with spinneret; 39. left pedipalp minus chela, dorsal view; 40. left chela, dorsal view; 41. left chela, lateral view; 42. claw of left leg IV; 43. subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39 – 41)." httpUri="https://zenodo.org/record/219883/files/figure.png" pageId="13" pageNumber="296">Figs 39–41</figureCitation>
). Femur weakly granulated on paraxial face, more pronounced in basal half; chelal hand weakly granulated at base of finger; trochanter and patella smooth. Lyrifissures as in
<figureCitation id="11CECCDDF406FF96585FFDF403BBFDF1" box="[1116,1245,583,607]" captionStart="FIGURES 34 – 43" captionStartId="12.[151,269,1870,1894]" captionTargetBox="[159,1434,199,1869]" captionTargetId="figure@12.[151,1436,194,1869]" captionTargetPageId="12" captionText="FIGURES 34 – 43. Roncocreagris occidentalis sp. nov., female holotype. 34. carapace; 35. anterior margin of carapace; 36. anterior and medial processes of right coxa I; 37. left chelicera; 38. partial view of fingers of left chelicera, with spinneret; 39. left pedipalp minus chela, dorsal view; 40. left chela, dorsal view; 41. left chela, lateral view; 42. claw of left leg IV; 43. subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39 – 41)." httpUri="https://zenodo.org/record/219883/files/figure.png" pageId="13" pageNumber="296">Figs 39–41</figureCitation>
. Femur with one tiny tubercle just distad of middle of antiaxial face, dorsally with one distal glandular pore. Patella with two dorsal micropores at base of pedicel. Chelal hand dorsal gently ovoid, widest proximad of middle; pedicel bears two dorsal micropores; antiaxial face with 4 glandular pores close to base of finger. Fixed finger with 100 teeth, most of them apically cusped, dental row reaching base of finger;
<emphasis id="BB810C4AF406FF965F58FD6B036DFD5E" box="[859,1035,728,752]" italics="true" pageId="13" pageNumber="296">nodus ramosus</emphasis>
short, level with 4th distal tooth; trichobothrium
<emphasis id="BB810C4AF406FF965D45FD4E0630FCBA" box="[326,342,765,788]" italics="true" pageId="13" pageNumber="296">it</emphasis>
proximad of
<emphasis id="BB810C4AF406FF965DF3FD4E0562FCBA" box="[496,516,765,788]" italics="true" pageId="13" pageNumber="296">et</emphasis>
, halfway between
<emphasis id="BB810C4AF406FF965ED9FD4E059FFCBA" box="[730,761,765,788]" italics="true" pageId="13" pageNumber="296">est</emphasis>
and
<emphasis id="BB810C4AF406FF965F32FD4E0423FCBA" box="[817,837,765,788]" italics="true" pageId="13" pageNumber="296">et</emphasis>
; trichobothrium
<emphasis id="BB810C4AF406FF965807FD4E0379FCBA" box="[1028,1055,765,788]" italics="true" pageId="13" pageNumber="296">ist</emphasis>
markedly closer to base of finger than to apex; distance between trichobothria
<emphasis id="BB810C4AF406FF965EA2FCAC05D1FC99" box="[673,695,799,823]" italics="true" pageId="13" pageNumber="296">ib</emphasis>
and
<emphasis id="BB810C4AF406FF965EF7FC930469FC99" box="[756,783,800,823]" italics="true" pageId="13" pageNumber="296">ist</emphasis>
0.5 (0.5–0.6) times shorter than that between
<emphasis id="BB810C4AF406FF96592EFC930221FC99" box="[1325,1351,800,823]" italics="true" pageId="13" pageNumber="296">ist</emphasis>
and
<emphasis id="BB810C4AF406FF965986FC9302F2FC99" box="[1413,1428,800,823]" italics="true" pageId="13" pageNumber="296">it</emphasis>
. Chelal microsetae pattern: all groups present, extending distad of trichobothrium
<emphasis id="BB810C4AF406FF965847FCF10302FCF4" box="[1092,1124,834,858]" italics="true" pageId="13" pageNumber="296">isb</emphasis>
,
<emphasis id="BB810C4AF406FF965876FCF003FDFCF4" box="[1141,1179,835,858]" italics="true" pageId="13" pageNumber="296">Em</emphasis>
5 (4–5),
<emphasis id="BB810C4AF406FF965908FCF00251FCF4" box="[1291,1335,835,858]" italics="true" pageId="13" pageNumber="296">Mm</emphasis>
2,
<emphasis id="BB810C4AF406FF96595CFCF0021AFCF4" box="[1375,1404,835,858]" italics="true" pageId="13" pageNumber="296">Im</emphasis>
2. Movable finger with 97 fingers, most of them cusped, dental row shorter than on fixed finger, ending just distad of trichobothrium
<emphasis id="BB810C4AF406FF965D46FC3A0635FC0F" box="[325,339,905,929]" italics="true" pageId="13" pageNumber="296">b</emphasis>
; distance between trichobothria
<emphasis id="BB810C4AF406FF965EBDFC3A05B1FC0F" box="[702,727,905,929]" italics="true" pageId="13" pageNumber="296">sb</emphasis>
and
<emphasis id="BB810C4AF406FF965F0FFC390479FC0F" box="[780,799,906,929]" italics="true" pageId="13" pageNumber="296">st</emphasis>
1.1 (0.9–1.2) times that between
<emphasis id="BB810C4AF406FF965891FC3A03CDFC0F" box="[1170,1195,905,929]" italics="true" pageId="13" pageNumber="296">sb</emphasis>
and
<emphasis id="BB810C4AF406FF9658E3FC3A0388FC0F" box="[1248,1262,905,929]" italics="true" pageId="13" pageNumber="296">b</emphasis>
. One sensillum near tip of both fingers; diploid sensillum
<emphasis id="BB810C4AF406FF965E6CFC1D05ECFC6B" box="[623,650,942,965]" italics="true" pageId="13" pageNumber="296">pc</emphasis>
not raised, close to dental margin, level with trichobothrium
<emphasis id="BB810C4AF406FF965935FC1E0229FC6B" box="[1334,1359,941,965]" italics="true" pageId="13" pageNumber="296">sb</emphasis>
.
</paragraph>
<paragraph id="894AD058F406FF965CC4FC620784FB81" blockId="13.[151,1437,151,2005]" pageId="13" pageNumber="296">
Legs. Claws of legs I and IV with a tiny dorsal tooth proximad of middle (
<figureCitation id="11CECCDDF406FF965804FC63033DFC46" box="[1031,1115,976,1000]" captionStart="FIGURES 34 – 43" captionStartId="12.[151,269,1870,1894]" captionTargetBox="[159,1434,199,1869]" captionTargetId="figure@12.[151,1436,194,1869]" captionTargetPageId="12" captionText="FIGURES 34 – 43. Roncocreagris occidentalis sp. nov., female holotype. 34. carapace; 35. anterior margin of carapace; 36. anterior and medial processes of right coxa I; 37. left chelicera; 38. partial view of fingers of left chelicera, with spinneret; 39. left pedipalp minus chela, dorsal view; 40. left chela, dorsal view; 41. left chela, lateral view; 42. claw of left leg IV; 43. subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39 – 41)." httpUri="https://zenodo.org/record/219883/files/figure.png" pageId="13" pageNumber="296">Fig. 42</figureCitation>
), subterminal setae with two rami and some spinules (
<figureCitation id="11CECCDDF406FF965DB0FC40056FFBA5" box="[435,521,1011,1035]" captionStart="FIGURES 34 – 43" captionStartId="12.[151,269,1870,1894]" captionTargetBox="[159,1434,199,1869]" captionTargetId="figure@12.[151,1436,194,1869]" captionTargetPageId="12" captionText="FIGURES 34 – 43. Roncocreagris occidentalis sp. nov., female holotype. 34. carapace; 35. anterior margin of carapace; 36. anterior and medial processes of right coxa I; 37. left chelicera; 38. partial view of fingers of left chelicera, with spinneret; 39. left pedipalp minus chela, dorsal view; 40. left chela, dorsal view; 41. left chela, lateral view; 42. claw of left leg IV; 43. subterminal seta of right left IV. Scale bars (in mm): 0.05 (Figs 42, 43), 0.10 (Figs 35, 36, 38), 0.20 (Fig. 37), 0.30 (Fig. 34), 0.50 (Figs 39 – 41)." httpUri="https://zenodo.org/record/219883/files/figure.png" pageId="13" pageNumber="296">Fig. 43</figureCitation>
). Leg IV tibia TS ratio 0.49 (0.45), basitarsus TS 0.14 (0.13), telotarsus TS 0.33 (0.44).
</paragraph>
<paragraph id="894AD058F406FF965CC4FB8803B6FA52" blockId="13.[151,1437,151,2005]" pageId="13" pageNumber="296">
<emphasis id="BB810C4AF406FF965CC4FB880630FBFD" box="[199,342,1083,1107]" italics="true" pageId="13" pageNumber="296">Deutonymph</emphasis>
. Body and appendages yellowish. Carapace longer than broad, no eyes or eye-spots, epistome absent; chaetotaxy 20 setae, formula 4:4:6:6. Manducatory process 3 setae, pedipalp coxa 5, pedal coxa
<date id="FD4BF698F406FF96591CFBEC0259FBD8" box="[1311,1343,1118,1142]" pageId="13" pageNumber="296">I 4</date>
,
<date id="FD4BF698F406FF965948FBEC0212FBD8" box="[1355,1396,1118,1142]" pageId="13" pageNumber="296">II 4</date>
,
<date id="FD4BF698F406FF965983FBEC07C1FB37" pageId="13" pageNumber="296">III 3</date>
,
<date id="FD4BF698F406FF965CB4FB310797FB37" box="[183,241,1153,1177]" pageId="13" pageNumber="296">IV 4</date>
; anterior process of coxa I long and tooth-shaped, medial process straight, with one denticle. Tergal chaetotaxy I–X: 6:6:7:7:7:7:7:7:7:8. Sternal chaetotaxy III–X: 4:4:9:9:8:9:8:9; sternites VI and VII each with 2 discal setae (included in sternal formula); discal setae distinctly closer to sternal row than in adults; segment
<date id="FD4BF698F406FF96596AFB7A02FAFB4E" box="[1385,1436,1224,1248]" pageId="13" pageNumber="296">XI 9</date>
setae; stigmata of sternites III and IV with one microseta on each side. Chelicera with five setae on palm, subgaleal seta 0.6 from base, spinneret reduced to a hyaline dome; fixed finger with 12 teeth, movable finger with 10 teeth. Pedipalps smooth, except chelal hand at base of finger; femur with a tiny antiaxial tubercle in basal half; distal opening of patella oblique; chelal hand more oval than in adults, maximum width proximad of middle, antiaxial face with one glandular pore close to base of finger, one dorsal micropore on pedicel; chelal microsetae pattern:
<emphasis id="BB810C4AF406FF965975FAC802FAFA3C" box="[1398,1436,1403,1426]" italics="true" pageId="13" pageNumber="296">Em</emphasis>
2,
<emphasis id="BB810C4AF406FF965CB0FA2C07B9FA18" box="[179,223,1439,1462]" italics="true" pageId="13" pageNumber="296">Mm</emphasis>
1,
<emphasis id="BB810C4AF406FF965D02FA2C0678FA18" box="[257,286,1439,1462]" italics="true" pageId="13" pageNumber="296">Im</emphasis>
0; fixed finger with six trichobothria and 48 teeth, movable finger with two trichobothria and 45 teeth, dental row ending at level of diploid sensillum. Leg IV: telotarsus basally swollen; claws bearing a tiny basal tooth; subterminal setae with tiny distal denticles; tibia TS 0.35, basitarsus TS 0.20, telotarsus TS 0.27.
</paragraph>
<paragraph id="894AD058F406FF965CC4F9BB0232F81F" blockId="13.[151,1437,151,2005]" pageId="13" pageNumber="296">
<emphasis id="BB810C4AF406FF965CC4F9BB0562F98E" bold="true" box="[199,516,1543,1568]" pageId="13" pageNumber="296">Measurements and ratios.</emphasis>
<emphasis id="BB810C4AF406FF965E0DF9BB0799F9ED" italics="true" pageId="13" pageNumber="296">
Female
<typeStatus id="564E6EFAF406FF965E6FF9BB05AAF98E" box="[620,716,1544,1568]" pageId="13" pageNumber="296" type="holotype">holotype</typeStatus>
(followed by female
<typeStatus id="564E6EFAF406FF965FC6F9BA034FF98E" box="[965,1065,1545,1568]" pageId="13" pageNumber="296" type="paratype">paratype</typeStatus>
from Algar do Javali in square brackets)
</emphasis>
. Body 3.74 [1.96, opisthosoma contracted]. Carapace 0.83/0.68 (1.2) [0.78/0.67 (1.2)]. Chelicera: palm 0.58/0.29 (2.0) [0.51/0.25 (2.0)], movable finger 0.36 [0.31]. Pedipalp: trochanter 0.68/0.22 (3.1) [0.61/0.19 (3.1)]; femur 1.28/0.22 (5.8) [1.13/0.20 (5.7)]; patella 1.21/0.26 (4.7) [1.06/0.23 (4.6)], pedicel 0.36 [0.34], club 0.85/0.26 (3.3) [0.72/0.23 (3.1)], ratio club/pedicel 2.4 [2.1]; chela+ 2.28/0.42 (5.5) [2.0/0.38 (5.3)], chela - 2.15/0.42 (5.1) [1.85/0.38 (4.9)]; hand+ 0.99 (2.4) [0.83 (2.2)], hand - 0.86 (2.1) [0.68 (1.8)]; movable finger 1.32 [1.18]; ratio chela+/carapace 2.8 [2.6]; femur/carapace 1.5 [1.5]; finger/femur 1.03 [1.04]; femur/patella 1.06 [1.07]; patella/ hand+ 1.22 [1.3]; finger/hand+ 1.3 [1.4]. Leg I: femur 0.60/0.15 (4.1) [0.55/0.15 (3.8)]; patella 0.48/0.12 (3.9) [0.41/ 0.13 (3.2)]; tibia 0.59/0.09 (6.9) [0.52/0.10 (5.2)]; basitarsus 0.29/0.07 (4.1) [0.26/0.07 (3.6)]; telotarsus 0.48/0.06 (8.0) [0.42/0.06 (6.7)]; ratio femur/patella 1.25 [1.3]; basitarsus/telotarsus 1.7 [1.7]. Leg IV: femur+patella 1.10/ 0.24 (4.7) [0.93/0.21 (4.4)]; tibia 0.99/0.13 (7.8) [0.88/0.12 (7.3)]; basitarsus 0.37/0.10 (3.7) [0.31/0.10 (3.0)]; telotarsus 0.58/0.08 (7.3) [0.53/0.09 (5.9)]; ratio femur+patella/tibia 1.1 [1.1]; telotarsus/basitarsus 1.6 [1.7].
</paragraph>
<paragraph id="894AD058F406FF955CC4F80E06C4FEC2" blockId="13.[151,1437,151,2005]" lastBlockId="14.[151,1437,151,1164]" lastPageId="14" lastPageNumber="297" pageId="13" pageNumber="296">
<emphasis id="BB810C4AF406FF965CC4F80E05C4F87B" box="[199,674,1981,2005]" italics="true" pageId="13" pageNumber="296">
Female
<typeStatus id="564E6EFAF406FF965D25F80D06ECF87B" box="[294,394,1982,2005]" pageId="13" pageNumber="296" type="paratype">paratype</typeStatus>
from Gruta dos Bolhos
</emphasis>
. Body 3.56. Carapace 0.78/0.66 (1.2). Chelicera: palm 0.52/0.26 (2.0); movable finger 0.34. Pedipalp: trochanter 0.60/0.21 (2.8); femur 1.12/0.22 (5.2); patella 1.06/0.27 (3.9), pedicel 0.32, club 0.74/0.27 (2.7), ratio club/pedicel 2.3; chela+ 2.04/0.40 (5.1), chela - 1.89/0.40 (4.7); hand+ 0.94 (2.3), hand - 0.79 (2.0); movable finger 1.12; ratio chela+/carapace 2.6; femur/carapace 1.4; finger/femur 1.0; femur/ patella 1.1; patella/hand+ 1.1; finger/hand+ 1.2. Leg I: femur 0.50/0.14 (3.7); patella 0.38/0.11 (3.4); tibia 0.48/0.08 (5.8); basitarsus 0.22/0.07 (3.4); telotarsus 0.40/0.06 (6.6); ratio femur/patella: 1.3; telotarsus/basitarsus 1.8. Leg IV: both legs IV absent.
</paragraph>
<paragraph id="894AD058F405FF955CC4FECB037DFDE2" blockId="14.[151,1437,151,1164]" pageId="14" pageNumber="297">
<emphasis id="BB810C4AF405FF955CC4FECB0630FE3E" box="[199,342,376,400]" italics="true" pageId="14" pageNumber="297">Deutonymph</emphasis>
. Body 1.52. Carapace 0.46/0.37 (1.2). Chelicera: palm 0.27/0.14 (2.0), movable finger 0.18. Pedipalp: trochanter 0.27/0.11 (2.4); femur 0.44/.012 (3.8); patella 0.37/0.14 (2.7), pedicel 0.12, club 0.25/0.14 (1.9), ratio club/pedicel 2.1; chela+ 0.86/0.21 (4.2), chela - 0.81/0.21 (3.9); hand+ 0.38 (1.8), hand - 0.33 (1.6); movable finger 0.48; ratio chela+/carapace 1.9; femur/carapace 1.0; finger/femur 1.1; femur/patella 1.2; patella/ hand+ 1.0; finger/hand+ 1.3. Leg IV: femur+patella 41/14 (3.0); tibia 0.32/0.7 (4.6); basitarsus 0.14/0.06 (2.4); telotarsus 0.22/0.06 (4.0); ratio femur+patella/tibia 1.3; telotarsus/basitarsus 1.6.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F405FF955CC4FDEA039BFD16" pageId="14" pageNumber="297" type="discussion">
<paragraph id="894AD058F405FF955CC4FDEA039BFD16" blockId="14.[151,1437,151,1164]" pageId="14" pageNumber="297">
<emphasis id="BB810C4AF405FF955CC4FDEA065DFDDF" bold="true" box="[199,315,601,625]" pageId="14" pageNumber="297">Remarks.</emphasis>
The main diagnostic characters of the
<typeStatus id="564E6EFAF405FF955EFEFDEA0438FDDF" box="[765,862,601,625]" pageId="14" pageNumber="297" type="paratype">paratype</typeStatus>
from Gruta dos Bolhos generally coinciding with those of the
<typeStatus id="564E6EFAF405FF955D24FDCF06EFFD3A" box="[295,393,636,660]" pageId="14" pageNumber="297" type="holotype">holotype</typeStatus>
and
<typeStatus id="564E6EFAF405FF955DC7FDCE0542FD3B" box="[452,548,637,661]" pageId="14" pageNumber="297" type="paratype">paratype</typeStatus>
from Algar do Javali, but it has a less slender chela. We prefer to assign the specimen from Gruta dos Bolhos to
<taxonomicName id="4EF5ABDBF405FF955E2EFD1105B5FD17" box="[557,723,673,697]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="BB810C4AF405FF955E2EFD1105B5FD17" box="[557,723,673,697]" italics="true" pageId="14" pageNumber="297">R. occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F405FF955ED9FD120455FD17" box="[730,819,673,697]" pageId="14" pageNumber="297" rank="species">
<emphasis id="BB810C4AF405FF955ED9FD120455FD17" bold="true" box="[730,819,673,697]" pageId="14" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
until new material is available for study.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F405FF955CC4FD7504BFFCC2" pageId="14" pageNumber="297" type="reference_group">
<paragraph id="894AD058F405FF955CC4FD7504BFFCC2" blockId="14.[151,1437,151,1164]" pageId="14" pageNumber="297">
<taxonomicName id="4EF5ABDBF405FF955CC4FD75069BFD73" ID-CoL="4TB4G" box="[199,509,709,733]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="BB810C4AF405FF955CC4FD75069BFD73" box="[199,509,709,733]" italics="true" pageId="14" pageNumber="297">Roncocreagris occidentalis</emphasis>
</taxonomicName>
represents the second hypogean
<taxonomicName id="4EF5ABDBF405FF955F82FD750343FD73" ID-CoL="63S5G" box="[897,1061,710,733]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="BB810C4AF405FF955F82FD750343FD73" box="[897,1061,710,733]" italics="true" pageId="14" pageNumber="297">Roncocreagris</emphasis>
</taxonomicName>
with the spinneret reduced to a hyaline dome, the other being
<taxonomicName id="4EF5ABDBF405FF955E00FD590433FCAE" ID-CoL="4TB4D" authority="Zaragoza, 2003" authorityName="Zaragoza" authorityYear="2003" box="[515,853,744,769]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="iglesiasae">
<emphasis id="BB810C4AF405FF955E00FD5905F1FCAF" box="[515,663,745,769]" italics="true" pageId="14" pageNumber="297">R. iglesiasae</emphasis>
<bibRefCitation id="ED64ADA9F405FF955EA0FD5A0433FCAE" author="Zaragoza" box="[675,853,744,769]" pageId="14" pageNumber="297" refString="Zaragoza, J. A. (2003) Roncocreagris iglesiasae, nueva especie cavernicola de Asturias (Arachnida, Pseudoscorpiones, Neobisiidae). Revista Iberica de Aracnologia, 7, 89 - 94." type="journal article" year="2003">Zaragoza, 2003</bibRefCitation>
</taxonomicName>
(Asturias,
<collectingCountry id="F1E290C8F405FF955FD8FD5B0379FCAE" box="[987,1055,744,768]" name="Spain" pageId="14" pageNumber="297">Spain</collectingCountry>
).
<taxonomicName id="4EF5ABDBF405FF955835FD590387FCAF" ID-CoL="4TB4G" box="[1078,1249,745,769]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="BB810C4AF405FF955835FD590387FCAF" box="[1078,1249,745,769]" italics="true" pageId="14" pageNumber="297">R. occidentalis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4EF5ABDBF405FF955987FD590661FC8B" ID-CoL="4TB4D" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="iglesiasae">
<emphasis id="BB810C4AF405FF955987FD590661FC8B" italics="true" pageId="14" pageNumber="297">R. iglesiasae</emphasis>
</taxonomicName>
in the position of trichobothrium
<emphasis id="BB810C4AF405FF955EA3FCBD05DCFC8B" box="[672,698,782,805]" italics="true" pageId="14" pageNumber="297">ist</emphasis>
, which is distinctly nearer to the finger base than to
<emphasis id="BB810C4AF405FF95593FFCBD022AFC8B" box="[1340,1356,782,805]" italics="true" pageId="14" pageNumber="297">it</emphasis>
in the Portuguese species, as opposed to being closer to
<emphasis id="BB810C4AF405FF955ED5FC810583FCE7" box="[726,741,818,841]" italics="true" pageId="14" pageNumber="297">it</emphasis>
than to the finger base in the Spanish species. Moreover,
<taxonomicName id="4EF5ABDBF405FF955987FC810678FCC3" ID-CoL="4TB4G" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="BB810C4AF405FF955987FC810678FCC3" italics="true" pageId="14" pageNumber="297">R. occidentalis</emphasis>
</taxonomicName>
has the pedipalps longer but less slender than in
<taxonomicName id="4EF5ABDBF405FF955F40FCE504B4FCC3" ID-CoL="4TB4D" box="[835,978,853,877]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="iglesiasae">
<emphasis id="BB810C4AF405FF955F40FCE504B4FCC3" box="[835,978,853,877]" italics="true" pageId="14" pageNumber="297">R. iglesiasae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F405FF955CC4FCCB0463FB22" pageId="14" pageNumber="297" type="distribution">
<paragraph id="894AD058F405FF955CC4FCCB0378FC1A" blockId="14.[151,1437,151,1164]" pageId="14" pageNumber="297">
<emphasis id="BB810C4AF405FF955CC4FCCB0694FC3F" bold="true" box="[199,498,888,913]" pageId="14" pageNumber="297">Distribution and habitat.</emphasis>
<taxonomicName id="4EF5ABDBF405FF955DF9FCC9044AFC3F" box="[506,812,889,913]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="BB810C4AF405FF955DF9FCC9044AFC3F" box="[506,812,889,913]" italics="true" pageId="14" pageNumber="297">Roncocreagris occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F405FF955F37FCCA04E9FC3F" box="[820,911,889,913]" pageId="14" pageNumber="297" rank="species">
<emphasis id="BB810C4AF405FF955F37FCCA04E9FC3F" bold="true" box="[820,911,889,913]" pageId="14" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
was collected in two caves from two different karst units, Javali in the Montejunto massif and Bolhos in the Cesaredas plateau.
</paragraph>
<paragraph id="894AD058F405FF955CC4FC720463FB22" blockId="14.[151,1437,151,1164]" pageId="14" pageNumber="297">
In Javali Cave, which has a year-round temperature of 15.1 + 0.5ºC at soil level,
<taxonomicName id="4EF5ABDBF405FF95585BFC710399FC77" box="[1112,1279,961,985]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="BB810C4AF405FF95585BFC710399FC77" box="[1112,1279,961,985]" italics="true" pageId="14" pageNumber="297">R. occidentalis</emphasis>
</taxonomicName>
is found with other interesting cave-adapted species, such as the ground beetle
<taxonomicName id="4EF5ABDBF405FF955FB5FC5602FAFC52" authority="Reboleira &amp; Ortuno, 2010" authorityName="Reboleira &amp; Ortuno" authorityYear="2010" box="[950,1436,996,1021]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="tatai">
<emphasis id="BB810C4AF405FF955FB5FC560329FC53" box="[950,1103,997,1021]" italics="true" pageId="14" pageNumber="297">Trechus tatai</emphasis>
<bibRefCitation id="ED64ADA9F405FF95585CFC5702FAFC52" author="Reboleira" box="[1119,1436,996,1020]" pageId="14" pageNumber="297" refString="Reboleira, A. S. P. S., Ortuno, V. M., Goncalves, F. &amp; Oromi, P. (2010 b) A hypogean new species of Trechus Clairville, 1806 (Coleoptera, Carabidae) from Portugal and considerations about the T. fulvus species group. Zootaxa, 2689, 15 - 26." type="journal article" year="2010">Reboleira &amp; Ortuño, 2010</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="ED64ADA9F405FF955C9DFBBB06C6FB8E" author="Reboleira" box="[158,416,1032,1057]" pageId="14" pageNumber="297" refString="Reboleira, A. S. P. S., Ortuno, V. M., Goncalves, F. &amp; Oromi, P. (2010 b) A hypogean new species of Trechus Clairville, 1806 (Coleoptera, Carabidae) from Portugal and considerations about the T. fulvus species group. Zootaxa, 2689, 15 - 26." type="journal article" year="2010" yearSuffix="b">
Reboleira
<emphasis id="BB810C4AF405FF955D16FBB90628FB8F" box="[277,334,1033,1057]" italics="true" pageId="14" pageNumber="297">et al.</emphasis>
2010b
</bibRefCitation>
), while in Bolhos Cave the biocoenosis is composed of a new trichoniscid woodlouse, the dipluran
<taxonomicName id="4EF5ABDBF405FF955CFDFB9E06E5FBEB" box="[254,387,1069,1093]" class="Entognatha" family="Campodeidae" genus="Podocampa" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="BB810C4AF405FF955CFDFB9E06E5FBEB" box="[254,387,1069,1093]" italics="true" pageId="14" pageNumber="297">Podocampa</emphasis>
</taxonomicName>
cf.
<emphasis id="BB810C4AF405FF955DB7FB9E054BFBEB" box="[436,557,1069,1093]" italics="true" pageId="14" pageNumber="297">fragiloides</emphasis>
and the thysanuran
<taxonomicName id="4EF5ABDBF405FF955F1FFB9E04E2FBEB" box="[796,900,1069,1093]" class="Insecta" family="Nicoletiidae" genus="Coletinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="14" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="BB810C4AF405FF955F1FFB9E04E2FBEB" box="[796,900,1069,1093]" italics="true" pageId="14" pageNumber="297">Coletinia</emphasis>
</taxonomicName>
sp. and the mean temperature is 16.3ºC. Also found in both caves was the pseudoscorpion
<taxonomicName id="4EF5ABDBF405FF955E97FBE2027CFBC6" authority="Zaragoza, 2012" authorityName="Zaragoza" authorityYear="2012" box="[660,1306,1104,1129]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="cardosoi" subGenus="Ephippiochthonius">
<emphasis id="BB810C4AF405FF955E97FBE20339FBC7" box="[660,1119,1105,1129]" italics="true" pageId="14" pageNumber="297">Chthonius (Ephippiochthonius) cardosoi</emphasis>
Zaragoza, 2012
</taxonomicName>
, previously known only from the Arrábida massif (Zaragoza 2012).
</paragraph>
</subSubSection>
</treatment>
</document>
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