201 lines
22 KiB
XML
201 lines
22 KiB
XML
<document id="54C0992BD15C7B09AA81FBA5A5D5C721" ID-CLB-Dataset="8753" ID-DOI="10.11646/zootaxa.4822.1.2" ID-GBIF-Dataset="d073f599-4282-4f4b-a071-5120d1a4af1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401190" ID-ZooBank="F2D5A199-48B1-448A-9B9A-458FBFB5253D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609333769558" checkinUser="plazi" docAuthor="Ferro, Gustavo Borges & Marshall, Stephen A." docDate="2020" docId="014B87AAFFCEFFAD24A6226CFEE6A233" docLanguage="en" docName="zootaxa.4822.1.2.pdf" docOrigin="Zootaxa 4822 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paragrallomyia bifurcata Ferro & Marshall 2020, new species" docType="treatment" docUuid="E3E77F65-E387-4EE5-8145-670A67C9A5D1" docUuidSource="ZooBank" docVersion="7" lastPageNumber="49" masterDocId="FD72FFD2FFC7FFA72431210AFF92A00E" masterDocTitle="A redefinition of Paragrallomyia Hendel (Diptera: Micropezidae, Taeniapterinae) and a revision of the P. albibasis complex" masterLastPageNumber="70" masterPageNumber="39" pageNumber="48" updateTime="1698849606894" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="D4C844B19C5A4F7DC4E95B74117B0725" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BD27534DCDF43B601973DBF0C41B3194">
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<mods:title id="9E27926C3EC223B662E9D78CCE89C5FD">A redefinition of Paragrallomyia Hendel (Diptera: Micropezidae, Taeniapterinae) and a revision of the P. albibasis complex</mods:title>
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</mods:titleInfo>
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<mods:name id="FA5F4EE65C817F39DB34DC48016CDD7E" type="personal">
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<mods:roleTerm id="05A1468BD5375D4896DD82DC9F81D6E1">Author</mods:roleTerm>
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<mods:namePart id="2B100337A70572D385F1AFCA9BBD0E5F">Ferro, Gustavo Borges</mods:namePart>
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<mods:nameIdentifier id="FC9A9890CD97D297A3126E67972295AB" type="ORCID">0000-0003-2091-0872</mods:nameIdentifier>
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<mods:affiliation id="F628B571D269C8E27C8AD4746D2F94DF">School of Environmental Sciences, University of Guelph, Guelph, ON, Canada, N 1 G 2 W 1. & bferrog @ gmail. com; https: // orcid. org / 0000 - 0003 - 2091 - 0872</mods:affiliation>
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<mods:nameIdentifier id="9186661C1A5A4A67FB7498779BEDDEAC" type="email">bferrog@gmail.com</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="79034DE4FAE7301156F502CE3069ADBA">Marshall, Stephen A.</mods:namePart>
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<mods:nameIdentifier id="EB34590A3981110F6EB9C6E4DE7653A4" type="ORCID">0000-0002-5732-9718</mods:nameIdentifier>
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<mods:affiliation id="429EAE2F181D5CDB3B82563013AA148B">School of Environmental Sciences, University of Guelph, Guelph, ON, Canada, N 1 G 2 W 1. & samarsha @ uoguelph. ca; https: // orcid. org / 0000 - 0002 - 5732 - 9718</mods:affiliation>
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<mods:nameIdentifier id="B49EAFED5173F7BCC13419DA65EA8E0F" type="email">samarsha@uoguelph.ca</mods:nameIdentifier>
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<mods:title id="74991DB4E0D198902AA38C5262ECB991">Zootaxa</mods:title>
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<mods:part id="6391B54AB70F72D9D577F47821889776">
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<mods:date id="EC7A59804E05D137D17C1119711948DB">2020</mods:date>
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<mods:number id="E58E6508788A5D0B02236F80652670CA">2020-08-04</mods:number>
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<mods:number id="C7CB173760A902CDD3BB32CD6BBBCEE0">4822</mods:number>
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<treatment id="014B87AAFFCEFFAD24A6226CFEE6A233" ID-DOI="http://doi.org/10.5281/zenodo.4401196" ID-GBIF-Taxon="177010955" ID-Zenodo-Dep="4401196" LSID="urn:lsid:zoobank.org:act:E3E77F65-E387-4EE5-8145-670A67C9A5D1" httpUri="http://treatment.plazi.org/id/014B87AAFFCEFFAD24A6226CFEE6A233" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
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<subSubSection id="C1F86537FFCEFFAE24A6226CFDF0A38E" box="[151,610,870,897]" pageId="9" pageNumber="48" type="nomenclature">
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<paragraph id="895D36BCFFCEFFAE24A6226CFDF0A38E" blockId="9.[151,610,870,933]" box="[151,610,870,897]" pageId="9" pageNumber="48">
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<heading id="D21581D0FFCEFFAE24A6226CFDF0A38E" bold="true" box="[151,610,870,897]" fontSize="11" level="1" pageId="9" pageNumber="48" reason="1">
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<emphasis id="BB96EAAEFFCEFFAE24A6226CFDF0A38E" bold="true" box="[151,610,870,897]" pageId="9" pageNumber="48">
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<taxonomicName id="4EE24D3FFFCEFFAE24A6226CFE59A38F" ID-CoL="9HVCC" authority="Ferro & Marshall, 2020" authorityName="Ferro & Marshall" authorityYear="2020" box="[151,459,870,897]" class="Insecta" family="Micropezidae" genus="Paragrallomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="bifurcata" status="sp. nov.">
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<emphasis id="BB96EAAEFFCEFFAE24A6226CFE59A38F" bold="true" box="[151,459,870,897]" italics="true" pageId="9" pageNumber="48">Paragrallomyia bifurcata</emphasis>
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<taxonomicNameLabel id="A0A557D5FFCEFFAE25E3226DFDF0A38E" box="[466,610,870,897]" pageId="9" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C1F86537FFCEFFAD24A62281FD6BA113" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="description">
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<paragraph id="895D36BCFFCEFFAE24A62281FEBFA3AB" blockId="9.[151,610,870,933]" box="[151,301,907,933]" pageId="9" pageNumber="48">
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(
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<figureCitation id="11D92A39FFCEFFAE24AE2281FEB6A3AB" box="[159,292,907,933]" captionStart="FIGURES 10–11" captionStartId="9.[151,264,1974,1998]" captionTargetBox="[151,1375,1425,1951]" captionTargetId="figure-494@9.[151,1436,1425,1951]" captionTargetPageId="9" captionText="FIGURES 10–11. Paragrallomyia bifurcata n. sp. 10, female, lateral view;11, female internal reproductive structures, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4401200" httpUri="https://zenodo.org/record/4401200/files/figure.png" pageId="9" pageNumber="48">Figs. 10–11</figureCitation>
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)
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</paragraph>
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<paragraph id="895D36BCFFCEFFAE24A622D9FCA7A473" blockId="9.[151,1437,979,1407]" pageId="9" pageNumber="48">
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Description (female only): Body length
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; wing
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. Head and abdomen black, thorax black. Palpus dark brown except for white outer face. Clypeus brown with sparse pale microsetulae. First flagellomere dark brown. Fronto-orbital plate dark brown. Row of katepisternal setae brownish. Fore tibia dark brown. Fore tarsus mostly white, tarsomere one brown on basal half only. Hind femur mostly brown, white on basal quarter and with a small angled preapical white ring. Tergites black. Oviscape black.
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</paragraph>
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<paragraph id="895D36BCFFCEFFAE24F6258DFEF2A503" blockId="9.[151,1437,979,1407]" pageId="9" pageNumber="48">Head: First flagellomere covered by pale microsetulae. Frontal vitta with a darkened spot anterior to ocellar plate. Frontal vitta flat, not swollen anterior to ocellar plate. Epicephala converging, not immediately merging posterior to ocellar plate. Epicephalon somewhat swollen above frontal vitta level. Epicephalon somewhat swollen above frontal vitta level.</paragraph>
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<paragraph id="895D36BCFFCEFFAE24F6241DFC51A577" blockId="9.[151,1437,979,1407]" pageId="9" pageNumber="48">
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Thorax: Mesonotum with lines of thin brownish microsetulae. Anterior half of postpronotal lobe bare. Femur covered with sparse black microsetulae. Mid and hind tibia almost cylindrical, sulcus on outer face weak. Cervical sclerite with a whitish oval swelling. Cell r
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<subScript id="156634F9FFCEFFAE2649247BFD00A571" attach="left" box="[632,658,1393,1407]" fontSize="6" pageId="9" pageNumber="48">4+5</subScript>
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open at the tip of the wing.
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</paragraph>
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<caption id="DD9D6634FFCEFFAE24A626BCFF5CA7FC" ID-DOI="http://doi.org/10.5281/zenodo.4401200" ID-Zenodo-Dep="4401200" httpUri="https://zenodo.org/record/4401200/files/figure.png" pageId="9" pageNumber="48" startId="9.[151,264,1974,1998]" targetBox="[151,1375,1425,1951]" targetPageId="9">
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<paragraph id="895D36BCFFCEFFAE24A626BCFF5CA7FC" blockId="9.[151,1436,1974,2034]" pageId="9" pageNumber="48">
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<emphasis id="BB96EAAEFFCEFFAE24A626BCFEC1A7C0" bold="true" box="[151,339,1974,1998]" pageId="9" pageNumber="48">FIGURES 10–11.</emphasis>
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<taxonomicName id="4EE24D3FFFCEFFAE256826BCFDCCA7C0" authority="Ferro & Marshall, 2020" authorityName="Ferro & Marshall" authorityYear="2020" box="[345,606,1974,1998]" class="Insecta" family="Micropezidae" genus="Paragrallomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="bifurcata" status="sp. nov.">
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<emphasis id="BB96EAAEFFCEFFAE256826BCFDCCA7C0" box="[345,606,1974,1998]" italics="true" pageId="9" pageNumber="48">Paragrallomyia bifurcata</emphasis>
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</taxonomicName>
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<emphasis id="BB96EAAEFFCEFFAE265526BCFD0CA7C0" bold="true" box="[612,670,1974,1998]" pageId="9" pageNumber="48">
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<taxonomicNameLabel id="A0A557D5FFCEFFAE265526BCFD0CA7C0" box="[612,670,1974,1998]" pageId="9" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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10, female, lateral view;11, female internal reproductive structures, ventral view.
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</paragraph>
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</caption>
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<paragraph id="895D36BCFFCDFFAD24F6219DFD6BA113" blockId="10.[151,1437,151,573]" pageId="10" pageNumber="49">
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Abdomen: Single spermathecal duct with an apical small rounded swelling. Single spermatheca cylindrical and forked, with sparse small rounded processes (
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<figureCitation id="11D92A39FFCDFFAD26A121B1FD72A0DB" box="[656,736,187,213]" captionStart="FIGURES 10–11" captionStartId="9.[151,264,1974,1998]" captionTargetBox="[151,1375,1425,1951]" captionTargetId="figure-494@9.[151,1436,1425,1951]" captionTargetPageId="9" captionText="FIGURES 10–11. Paragrallomyia bifurcata n. sp. 10, female, lateral view;11, female internal reproductive structures, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4401200" httpUri="https://zenodo.org/record/4401200/files/figure.png" pageId="10" pageNumber="49">Fig. 11</figureCitation>
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). Paired spermathecal duct uniformly thick, slightly tapered on apical third before splitting into stems. Paired spermathecal duct stems uniformly thick, slightly convoluted. Paired spermathecae oval with an apical nipple-like swelling.
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</paragraph>
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</subSubSection>
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<subSubSection id="C1F86537FFCDFFAD24F6202DFD6BA1A3" pageId="10" pageNumber="49" type="materials_examined">
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<emphasis id="BB96EAAEFFCDFFAD24F6202DFEE8A14E" bold="true" box="[199,378,294,321]" pageId="10" pageNumber="49">Type material:</emphasis>
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<collectingCountry id="F1F5762CFFCDFFAD25CC202DFD37A14F" box="[509,677,295,321]" name="Costa Rica" pageId="10" pageNumber="49">COSTA RICA</collectingCountry>
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.
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<collectingRegion id="4B26F85EFFCDFFAD2683202DFC9CA14F" box="[690,782,295,321]" country="Costa Rica" name="Heredia" pageId="10" pageNumber="49">Heredia</collectingRegion>
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, Rara Avis Nat. Res.
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S
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(
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,
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<collectionCode id="EFF3AE79FFCDFFAD20262041FBFDA16B" box="[1047,1135,331,357]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="10" pageNumber="49">MNCR</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="398A3CE1FFCDFFAD20B32041FD67A1A3" ID-GBIF-Occurrence="3020124395" collectingDate="2005-02-22" collectingDateMax="2005-02-27" collectingDateMin="2005-02-22" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" elevation="700" latitude="10.283334" location="Las Horquetas" longLatPrecision="21" longitude="-84.04722" pageId="10" pageNumber="49" specimenCode="DEBU00241632, DEBU00241634, DEBU00241650" specimenCount="3" specimenCount-female="3" stateProvince="Heredia" typeStatus="paratype">
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<typeStatus id="5659881EFFCDFFAD20B32041FB61A16B" box="[1154,1267,331,357]" pageId="10" pageNumber="49" type="paratype">Paratypes</typeStatus>
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:
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<collectingRegion id="4B26F85EFFCDFFAD20CE2041FAC9A16B" box="[1279,1371,331,357]" country="Costa Rica" name="Heredia" pageId="10" pageNumber="49">Heredia</collectingRegion>
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, Rara Avis Nat. Res.
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<quantity id="4E1A9B59FFCDFFAD25782065FE01A187" box="[329,403,367,393]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="49" unit="km" value="12.0">12 km</quantity>
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S
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<location id="8C3D6067FFCDFFAD25872065FDCDA187" LSID="urn:lsid:plazi:treatment:014B87AAFFCEFFAD24A6226CFEE6A233:8C3D6067FFCDFFAD25872065FDCDA187" box="[438,607,367,393]" country="Costa Rica" latitude="10.283334" longLatPrecision="21" longitude="-84.04722" name="Las Horquetas" pageId="10" pageNumber="49" stateProvince="Heredia">Las Horquetas</location>
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,
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||
<geoCoordinate id="ECD6507BFFCDFFAD265C2065FD6FA187" box="[621,765,367,393]" degrees="10" direction="north" minutes="17" orientation="latitude" pageId="10" pageNumber="49" precision="15" seconds="00" value="10.283334">10º17’00” N</geoCoordinate>
|
||
<geoCoordinate id="ECD6507BFFCDFFAD27362064FC0CA187" box="[775,926,366,393]" degrees="84" direction="west" minutes="02" orientation="longitude" pageId="10" pageNumber="49" precision="15" seconds="50" value="-84.04722">84º02’50” W</geoCoordinate>
|
||
,
|
||
<quantity id="4E1A9B59FFCDFFAD279D2065FC6AA187" box="[940,1016,367,393]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="49" unit="m" value="700.0">
|
||
<elevation id="02CFD18FFFCDFFAD279D2065FC6AA187" box="[940,1016,367,393]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="49" unit="m" value="700.0">700 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FD5C107CFFCDFFAD20362064FB3AA187" box="[1031,1192,366,393]" pageId="10" pageNumber="49" value="2005-02-22" valueMax="2005-02-27" valueMin="2005-02-22">
|
||
<collectingDate id="ED18E994FFCDFFAD20362064FB3AA187" box="[1031,1192,366,393]" pageId="10" pageNumber="49" value="2005-02-22" valueMax="2005-02-27" valueMin="2005-02-22">22–27.II.2005</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2417536AFFCDFFAD20862065FACDA187" box="[1207,1375,367,393]" pageId="10" pageNumber="49">S. A. Marshall</collectorName>
|
||
(
|
||
<specimenCount id="9FE4FD35FFCDFFAD21452065FA04A187" box="[1396,1430,367,393]" pageId="10" pageNumber="49" type="female">3♀</specimenCount>
|
||
,
|
||
<specimenCode id="D9449EC7FFCDFFAD24A62099FEC4A1A3" box="[151,342,403,429]" collectionCode="DEBU" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="10" pageNumber="49">DEBU00241632</specimenCode>
|
||
,
|
||
<specimenCode id="D9449EC7FFCDFFAD25502099FDB2A1A3" box="[353,544,403,429]" collectionCode="DEBU" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="10" pageNumber="49">DEBU00241634</specimenCode>
|
||
,
|
||
<specimenCode id="D9449EC7FFCDFFAD261D2098FD7FA1A3" box="[556,749,402,429]" collectionCode="DEBU" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="10" pageNumber="49">DEBU00241650</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1F86537FFCDFFAD24F620BCFE79A1DF" box="[199,491,438,465]" pageId="10" pageNumber="49" type="distribution">
|
||
<paragraph id="895D36BCFFCDFFAD24F620BCFE79A1DF" blockId="10.[151,1437,151,573]" box="[199,491,438,465]" pageId="10" pageNumber="49">
|
||
<emphasis id="BB96EAAEFFCDFFAD24F620BCFEF1A1DE" bold="true" box="[199,355,438,464]" pageId="10" pageNumber="49">Distribution:</emphasis>
|
||
<collectingCountry id="F1F5762CFFCDFFAD255B20BDFE7AA1DF" box="[362,488,439,465]" name="Costa Rica" pageId="10" pageNumber="49">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1F86537FFCDFFAD24F620D1FAABA1FB" box="[199,1337,475,501]" pageId="10" pageNumber="49" type="etymology">
|
||
<paragraph id="895D36BCFFCDFFAD24F620D1FAABA1FB" blockId="10.[151,1437,151,573]" box="[199,1337,475,501]" pageId="10" pageNumber="49">
|
||
<emphasis id="BB96EAAEFFCDFFAD24F620D1FEC0A1FB" bold="true" box="[199,338,475,501]" pageId="10" pageNumber="49">Etymology:</emphasis>
|
||
The specific name refers to the remarkable bifurcate single spermathecae of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1F86537FFCDFFAD24F620F4FEE6A233" pageId="10" pageNumber="49" type="discussion">
|
||
<paragraph id="895D36BCFFCDFFAD24F620F4FEE6A233" blockId="10.[151,1437,151,573]" pageId="10" pageNumber="49">
|
||
<emphasis id="BB96EAAEFFCDFFAD24F620F4FEC0A216" bold="true" box="[199,338,510,536]" pageId="10" pageNumber="49">Comments:</emphasis>
|
||
The conspicuously bifurcate single spermathecae confirm that
|
||
<taxonomicName id="4EE24D3FFFCDFFAD203E20F5FB1DA217" authorityName="Ferro & Marshall" authorityYear="2020" box="[1039,1167,511,537]" class="Insecta" family="Micropezidae" genus="Paragrallomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="bifurcata">
|
||
<emphasis id="BB96EAAEFFCDFFAD203E20F5FB1DA217" box="[1039,1167,511,537]" italics="true" pageId="10" pageNumber="49">P. bifurcata</emphasis>
|
||
</taxonomicName>
|
||
is distinct from the very similar
|
||
<taxonomicName id="4EE24D3FFFCDFFAD24DD2329FEFEA233" authorityName="Enderlein" authorityYear="1922" box="[236,364,547,573]" class="Insecta" family="Micropezidae" genus="Grallomya" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="albibasis">
|
||
<emphasis id="BB96EAAEFFCDFFAD24DD2329FEFEA233" box="[236,364,547,573]" italics="true" pageId="10" pageNumber="49">P. albibasis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |