treatments-xml/data/01/39/87/013987C4FF83FFC9FF61C1F72B8919D9.xml

<document id="AF22589350FEA4A622B4C9B00E554F63" ID-CLB-Dataset="21737" ID-DOI="10.37520/aemnp.2020.019" ID-GBIF-Dataset="172c39ed-46c4-451f-b951-dd2d6d8c95b1" ID-ISSN="1804-6487" ID-Zenodo-Dep="3879941" ID-ZooBank="BF4AF355-51FF-4781-9663-AB855D936263" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591389696428" checkinUser="tatiana" docAuthor="Zhao, Xin &amp; Tang, Liang" docDate="2020" docId="013987C4FF83FFC9FF61C1F72B8919D9" docLanguage="en" docName="ActaEntMusNatPra.60.1.333-341.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Sphaerobulbus Zhao &amp; Tang 2020" docType="key" docVersion="6" lastPageNumber="341" masterDocId="FD00FFBCFF86FFC1FF98C20A297E1D6E" masterDocTitle="Three new species of the genus Sphaerobulbus from China (Coleoptera: Staphylinidae)" masterLastPageNumber="341" masterPageNumber="333" pageNumber="338" updateTime="1698842162098" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="AD51E4F2925ACC14D49ABB7ABD6DAEE7">Three new species of the genus Sphaerobulbus from China (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="135B7D8B22CF112125EAEE38B5EF5169">Zhao, Xin</mods:namePart>
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<mods:namePart id="5D4CEE060A988833DBB43FCCA73C9229">Tang, Liang</mods:namePart>
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<treatment id="013987C4FF83FFC9FF61C1F72B8919D9" ID-DOI="http://doi.org/10.5281/zenodo.3883354" ID-GBIF-Taxon="164481711" ID-Zenodo-Dep="3883354" LSID="urn:lsid:plazi:treatment:013987C4FF83FFC9FF61C1F72B8919D9" httpUri="http://treatment.plazi.org/id/013987C4FF83FFC9FF61C1F72B8919D9" lastPageId="8" lastPageNumber="341" pageId="5" pageNumber="338">
<subSubSection id="C18A6559FF83FFC4FF61C1F728CC19CB" pageId="5" pageNumber="338" type="nomenclature">
<paragraph id="892F36D2FF83FFC4FF61C1F72BE81979" blockId="5.[249,662,1020,1047]" box="[249,662,1020,1047]" pageId="5" pageNumber="338">
<heading id="D26781BEFF83FFC4FF61C1F72BE81979" bold="true" box="[249,662,1020,1047]" centered="true" fontSize="11" level="1" pageId="5" pageNumber="338" reason="1">
<emphasis id="BBE4EAC0FF83FFC4FF61C1F72BE81979" bold="true" box="[249,662,1020,1047]" pageId="5" pageNumber="338">
Key to species of 
<taxonomicName id="4E904D51FF83FFC4FE47C1F62BE81979" ID-CoL="8W3PV" authorityName="Zhao &amp; Tang" authorityYear="2020" box="[479,662,1020,1047]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="BBE4EAC0FF83FFC4FE47C1F62BE81979" bold="true" box="[479,662,1020,1047]" italics="true" pageId="5" pageNumber="338">Sphaerobulbus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="892F36D2FF83FFC4FF2DC62428CC19CB" blockId="5.[151,760,1070,1189]" pageId="5" pageNumber="338">The key is based on species descriptions provided by SMൾŦANA (2003, 2005, 2006, 2010, 2016) and the photographs of all species from Smetana’s collection taken by the second author in 2012.</paragraph>
</subSubSection>
<subSubSection id="C18A6559FF83FFC9FF0FC6B72B8919D9" lastPageId="8" lastPageNumber="341" pageId="5" pageNumber="338" type="key">
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<paragraph id="892F36D2FF83FFC4FF0FC6B72B86187B" blockId="5.[151,760,1213,2069]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FF0FC6B72B86187B" pageId="5" pageNumber="338">1 Elytra about as-long-as or longer than pronotum, EL/ PL=0.99–1.1; abdominal tergites VII with apical palisade fringe. .................................................................... 2</keyLead>
</paragraph>
<paragraph id="892F36D2FF83FFC4FF0FC7172B86181B" blockId="5.[151,760,1213,2069]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FF0FC7172B86181B" pageId="5" pageNumber="338">– Elytra shorter than pronotum, EL/PL=0.77–0.90; abdominal tergites VII without apical palisade fringe. ........................................................................................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF83FFC4FF0FC7772B861B5B" pageId="5" pageNumber="338">
<paragraph id="892F36D2FF83FFC4FF0FC7772B8618BB" blockId="5.[151,760,1213,2069]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FF0FC7772B8618BB" pageId="5" pageNumber="338">2 Abdominal tergites III–V each with deep median impression delimited by two short oblique carinae and posterior basal line; body without bluish tint. ......... 3</keyLead>
</paragraph>
<paragraph id="892F36D2FF83FFC4FF0FC7D72B861B5B" blockId="5.[151,760,1213,2069]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FF0FC7D72B861B5B" pageId="5" pageNumber="338">– Abdominal tergites III–V each without deep median impression and two short oblique carinae; body with bluish tint. .................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF83FFC4FF0FC4372B861A3B" pageId="5" pageNumber="338">
<paragraph id="892F36D2FF83FFC4FF0FC4372B891B9B" blockId="5.[151,760,1213,2069]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FF0FC4372B891B9B" pageId="5" pageNumber="338">
3 Pronotum as long as wide, PL/PW=1.00; subhumeral portion of each elytron with gold tomentose pubescence, abdominal tergites VI and VII each with medi- an golden tomentose patch. Habitus: 
<figureCitation id="11AB2A57FF83FFC4FDCDC4972BDF1BDB" box="[597,673,1693,1717]" captionStart="Figs 32–37" captionStartId="7.[151,188,1692,1711]" captionTargetBox="[182,1416,194,1655]" captionTargetId="figure@7.[164,1424,189,1674]" captionTargetPageId="7" captionText="Figs 32–37. Habitus of species of Sphaerobulbus. 32 – S. pusio Smetana, 2005 (paratype); 33 – S. nagahatai Smetana, 2005 (paratype); 34 – S. rex Smetana, 2005 (paratype); 35 – S. yunnanus Smetana, 2003 (paratype); 36 – S. ornatus Smetana, 2006 (paratype); 37 – S. yulongmontis Smetana, 2003 (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879949" httpUri="https://zenodo.org/record/3879949/files/figure.png" pageId="5" pageNumber="338">Fig. 36</figureCitation>
. Aedeagus: 
<bibRefCitation id="ED014B23FF83FFC4FF6DC4B728DE1BBB" author="SMETANA A." box="[245,416,1725,1749]" pageId="5" pageNumber="338" pagination="41 - 47" refId="ref4824" refString="SMETANA A. 2006: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 13. The genus Sphaerobulbus Smetana 2003. Section 3. Zootaxa 1318: 41 - 47." type="journal article" year="2006">SMETANA (2006</bibRefCitation>
: Figs 8–10). 
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(
<collectingRegion id="4B54F830FF83FFC4FDEBC4B72BB31BBB" box="[627,717,1725,1749]" country="China" name="Shaanxi" pageId="5" pageNumber="338">Shaanxi</collectingRegion>
). .... ............................................. 
<taxonomicName id="4E904D51FF83FFC4FE70C4D72B891B9B" ID-CoL="4YPPH" authority="Smetana, 2006" authorityName="Smetana" authorityYear="2006" box="[488,759,1757,1781]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="BBE4EAC0FF83FFC4FE70C4D72B291B9B" bold="true" box="[488,599,1757,1781]" italics="true" pageId="5" pageNumber="338">S. ornatus</emphasis>
<bibRefCitation id="ED014B23FF83FFC4FDC5C4D72B891B9B" author="SMETANA A." box="[605,759,1757,1781]" pageId="5" pageNumber="338" pagination="41 - 47" refId="ref4824" refString="SMETANA A. 2006: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 13. The genus Sphaerobulbus Smetana 2003. Section 3. Zootaxa 1318: 41 - 47." type="journal article" year="2006">Smetana, 2006</bibRefCitation>
</taxonomicName>
</keyLead>
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<keyLead id="326195E7FF83FFC4FF0FC4F72B861A3B" pageId="5" pageNumber="338">– Pronotum distinctly longer than wide, PL/PW=1.21– 1.30; elytra and abdominal tergites without gold tomentose pubescence or patch. .................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF83FFC4FF0FC5572CE21C38" pageId="5" pageNumber="338">
<paragraph id="892F36D2FF83FFC4FF0FC5572B89157B" blockId="5.[151,760,1213,2069]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FF0FC5572B89157B" pageId="5" pageNumber="338">
4 Head as wide as long, HW/HL=1.00; paramere asymmetrical, underside of paramere with three degenerated sensory peg setae in apical portion. Habitus: 
<figureCitation id="11AB2A57FF83FFC4FE9FC5B7282D1ABB" box="[263,339,1981,2005]" captionStart="Figs 32–37" captionStartId="7.[151,188,1692,1711]" captionTargetBox="[182,1416,194,1655]" captionTargetId="figure@7.[164,1424,189,1674]" captionTargetPageId="7" captionText="Figs 32–37. Habitus of species of Sphaerobulbus. 32 – S. pusio Smetana, 2005 (paratype); 33 – S. nagahatai Smetana, 2005 (paratype); 34 – S. rex Smetana, 2005 (paratype); 35 – S. yunnanus Smetana, 2003 (paratype); 36 – S. ornatus Smetana, 2006 (paratype); 37 – S. yulongmontis Smetana, 2003 (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879949" httpUri="https://zenodo.org/record/3879949/files/figure.png" pageId="5" pageNumber="338">Fig. 34</figureCitation>
. Aedeagus: 
<bibRefCitation id="ED014B23FF83FFC4FE4CC5B72BFC1ABB" author="SMETANA A." box="[468,642,1981,2005]" pageId="5" pageNumber="338" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">SMETANA (2005</bibRefCitation>
: Figs 5–7). 
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, 
<collectingRegion id="4B54F830FF83FFC4FEEDC5D728C71A9B" box="[373,441,2013,2037]" country="China" name="Hubei" pageId="5" pageNumber="338">Hubei</collectingRegion>
, 
<collectingRegion id="4B54F830FF83FFC4FE5BC5D72B641A9B" box="[451,538,2013,2037]" country="China" name="Sichuan" pageId="5" pageNumber="338">Sichuan</collectingRegion>
, 
<collectingRegion id="4B54F830FF83FFC4FDBCC5D72BFB1A9B" box="[548,645,2013,2037]" country="China" name="Guizhou" pageId="5" pageNumber="338">Guizhou</collectingRegion>
?). .............. ..................................................... 
<taxonomicName id="4E904D51FF83FFC4FD83C5F72B89157B" ID-CoL="4YPPL" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[539,759,2045,2069]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="BBE4EAC0FF83FFC4FD83C5F72B28157B" bold="true" box="[539,598,2045,2069]" italics="true" pageId="5" pageNumber="338">S. rex</emphasis>
<bibRefCitation id="ED014B23FF83FFC4FDC5C5F72B89157B" author="SMETANA A." box="[605,759,2045,2069]" pageId="5" pageNumber="338" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="892F36D2FF83FFC4FCA4C2B52CE21C38" blockId="5.[828,1437,191,2071]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FCA4C2B52CE21C38" pageId="5" pageNumber="338">
– Head a little wider than long, HW/HL=1.08; paramere almost symmetrical, underside of paramere with five degenerated sensory peg setae in subapical portion. Habitus: Figs 3, 4. Aedeagus: 
<figureCitation id="11AB2A57FF83FFC4FB43C3152C2E1C58" box="[1243,1360,286,311]" captionStart="Figs 13–18" captionStartId="3.[150,187,2037,2056]" captionTargetBox="[151,1435,102,2010]" captionTargetPageId="3" captionText="Figs 13–18. Sphaerobulbus guangxiensis sp. nov; 13 – male abdominal sternite IX; 14 – male abdominal tergite X; 15 – aedeagus in ventral view; 16, 17 – underside of apical paramere; 18 – female abdominal tergite X. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879943" httpUri="https://zenodo.org/record/3879943/files/figure.png" pageId="5" pageNumber="338">Figs 15–17</figureCitation>
. 
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(
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). ............................. 
<emphasis id="BBE4EAC0FF83FFC4FB05C3342CE21C38" bold="true" box="[1181,1436,318,342]" pageId="5" pageNumber="338">
<taxonomicName id="4E904D51FF83FFC4FB05C3342C3A1C38" ID-CoL="8Y9Q9" authority="Zhao &amp; Tang, 2020" authorityName="Zhao &amp; Tang" authorityYear="2020" box="[1181,1348,318,342]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="guangxiensis" status="sp. nov.">
<emphasis id="BBE4EAC0FF83FFC4FB05C3342C3A1C38" bold="true" box="[1181,1348,318,342]" italics="true" pageId="5" pageNumber="338">S. guangxiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D757BBFF83FFC4FAD1C3342CE21C38" box="[1353,1436,318,342]" pageId="5" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF83FFC4FCA4C3542CE21F99" pageId="5" pageNumber="338">
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<keyLead id="326195E7FF83FFC4FCA4C3542CE21F78" pageId="5" pageNumber="338">
5 Posterior angles of head rounded; posterior half of head with broad impunctate midline; apical portion of aedeagal median lobe hook-like; underside of paramere with sparse sensory peg setae. Habitus: Figs 5, 6. Aedeagus: Figs 19–23. 
<collectingCountry id="F1877642FF83FFC4FBECC3D52DCB1C99" box="[1140,1205,479,503]" name="China" pageId="5" pageNumber="338">China</collectingCountry>
(
<collectingRegion id="4B54F830FF83FFC4FB5BC3D52C631C99" box="[1219,1309,479,503]" country="China" name="Yunnan" pageId="5" pageNumber="338">Yunnan</collectingRegion>
). .................. ......................................................... 
<emphasis id="BBE4EAC0FF83FFC4FB4FC3F42CE21F78" bold="true" box="[1239,1436,510,534]" pageId="5" pageNumber="338">
<taxonomicName id="4E904D51FF83FFC4FB4FC3F42C3A1F78" ID-CoL="8Y9QC" authority="Zhao &amp; Tang, 2020" authorityName="Zhao &amp; Tang" authorityYear="2020" box="[1239,1348,510,534]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="ningliei" status="sp. nov.">
<emphasis id="BBE4EAC0FF83FFC4FB4FC3F42C3A1F78" bold="true" box="[1239,1348,510,534]" italics="true" pageId="5" pageNumber="338">S. ningliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D757BBFF83FFC4FAD1C3F42CE21F78" box="[1353,1436,510,534]" pageId="5" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="892F36D2FF83FFC4FCA4C0152CE21F99" blockId="5.[828,1437,191,2071]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FCA4C0152CE21F99" pageId="5" pageNumber="338">
– Posterior angles of head slightly prominent; posterior half of head with narrow impunctate midline; apical portion of aedeagal median lobe straightly pointed apicad; underside of paramere with dense sensory peg setae. Habitus: 
<figureCitation id="11AB2A57FF83FFC4FBFBC0952DCC1FD9" box="[1123,1202,671,695]" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="5" pageNumber="338">Fig. 26</figureCitation>
. Aedeagus: 
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: Figs 3, 4). 
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, 
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). ................ ......................................... 
<taxonomicName id="4E904D51FF83FFC4FBEDC0D42CE21F99" ID-CoL="4YPPC" authority="Smetana, 2010" authorityName="Smetana" authorityYear="2010" box="[1141,1436,734,759]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="cardinalis">
<emphasis id="BBE4EAC0FF83FFC4FBEDC0D42D851F98" bold="true" box="[1141,1275,734,758]" italics="true" pageId="5" pageNumber="338">S. cardinalis</emphasis>
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6 Forebody metallic blue. Habitus: Figs 1, 2. Aedeagus: Figs 7–12. 
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). ................................. ....................................................... 
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<taxonomicName id="4E904D51FF83FFC4FB5EC1342C3A1E38" ID-CoL="8Y9QD" authority="Zhao &amp; Tang, 2020" authorityName="Zhao &amp; Tang" authorityYear="2020" box="[1222,1348,830,854]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
<emphasis id="BBE4EAC0FF83FFC4FB5EC1342C3A1E38" bold="true" box="[1222,1348,830,854]" italics="true" pageId="5" pageNumber="338">S. smetanai</emphasis>
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</emphasis>
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– Forebody blackish (head and pronotum with dark blue hue in 
<taxonomicName id="4E904D51FF83FFC4FC7DC1742D3E1EF8" ID-CoL="4YPPD" authorityName="Smetana" authorityYear="2016" box="[997,1088,894,918]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="BBE4EAC0FF83FFC4FC7DC1742D3E1EF8" box="[997,1088,894,918]" italics="true" pageId="5" pageNumber="338">S. davidi</emphasis>
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). .................................................. 7
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<keyLead id="326195E7FF83FFC4FCA4C1952CE21EB9" pageId="5" pageNumber="338">7 Abdominal tergites VI and VII each with median golden tomentose patch. ................................................... 8</keyLead>
</paragraph>
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<keyLead id="326195E7FF83FFC4FCA4C1D52CE31979" pageId="5" pageNumber="338">– Abdominal tergites VI and VII each without median golden tomentose patch. .......................................... 10</keyLead>
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8 Larger species, BL= 
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. Aedeagus: 
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: Figs 7, 8). 
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). ............................... ............................................ 
<taxonomicName id="4E904D51FF83FFC4FB12C6542CE21919" ID-CoL="4YPPM" authority="Smetana, 2016" authorityName="Smetana" authorityYear="2016" box="[1162,1436,1118,1143]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="thomasi">
<emphasis id="BBE4EAC0FF83FFC4FB12C6542D851918" bold="true" box="[1162,1275,1118,1142]" italics="true" pageId="5" pageNumber="338">S. thomasi</emphasis>
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– Smaller species, BL= 
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. ................................ 9
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9 Elytra much shorter than wide, EL/EW=0.75; elytra reddish brown. Habitus: 
<figureCitation id="11AB2A57FF83FFC4FBE8C6B52DC419B9" box="[1136,1210,1215,1239]" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="5" pageNumber="338">Fig. 29</figureCitation>
. Aedeagus: 
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, Figs 3–5). 
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). ............................... ....................................... 
<taxonomicName id="4E904D51FF83FFC4FBFFC6F42CE21879" ID-CoL="4YPP8" authority="Smetana, 2006" authorityName="Smetana" authorityYear="2006" box="[1127,1436,1278,1303]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="biplagiatus">
<emphasis id="BBE4EAC0FF83FFC4FBFFC6F42D851878" bold="true" box="[1127,1275,1278,1302]" italics="true" pageId="5" pageNumber="338">S. biplagiatus</emphasis>
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– Elytra shorter than wide, EL/EW=0.86; elytra blackish. Habitus: 
<figureCitation id="11AB2A57FF83FFC4FC6CC7352D411839" box="[1012,1087,1343,1367]" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="5" pageNumber="338">Fig. 30</figureCitation>
. Aedeagus: 
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: 
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). 
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(
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). ............................................... ............................................ 
<taxonomicName id="4E904D51FF83FFC4FB10C7742CE218F9" ID-CoL="6Z8C9" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[1160,1436,1406,1431]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="brezinai">
<emphasis id="BBE4EAC0FF83FFC4FB10C7742D8518F8" bold="true" box="[1160,1275,1406,1430]" italics="true" pageId="5" pageNumber="338">S. brezinai</emphasis>
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10 Smaller species, BL= 
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; posterior margin of abdominal segment VII and entire abdominal segments VIII–X reddish yellow. Habitus: 
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. Aedeagus: 
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: Figs 10–12). 
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). ................................................. 
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<emphasis id="BBE4EAC0FF83FFC4FB3FC4142D821B58" bold="true" box="[1191,1276,1566,1590]" italics="true" pageId="5" pageNumber="338">S. pusio</emphasis>
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– Larger species, BL= 
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; entire abdominal segments VII–X blackish. ........................................ 11
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11 Median impression of abdominal tergites III–V relatively deeper, laterally delimited by two short oblique carinae. Habitus: 
<figureCitation id="11AB2A57FF83FFC4FBBCC4B52D111BB8" box="[1060,1135,1726,1751]" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="5" pageNumber="338">Fig. 25</figureCitation>
. Aedeagus: 
<bibRefCitation id="ED014B23FF83FFC4FB75C4B52CE71BB9" author="SMETANA A." box="[1261,1433,1727,1751]" pageId="5" pageNumber="338" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">SMETANA (2003</bibRefCitation>
: Figs 22–25). 
<collectingCountry id="F1877642FF83FFC4FC69C4D52D4C1B99" box="[1009,1074,1759,1783]" name="China" pageId="5" pageNumber="338">China</collectingCountry>
(
<collectingRegion id="4B54F830FF83FFC4FBD8C4D52DE61B99" box="[1088,1176,1759,1783]" country="China" name="Yunnan" pageId="5" pageNumber="338">Yunnan</collectingRegion>
, 
<collectingRegion id="4B54F830FF83FFC4FB3AC4D52D851B99" box="[1186,1275,1759,1783]" country="China" name="Sichuan" pageId="5" pageNumber="338">Sichuan</collectingRegion>
). ....................... ........................................ 
<taxonomicName id="4E904D51FF83FFC4FBE8C4F42CE21A79" ID-CoL="4YPP9" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[1136,1436,1790,1815]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="bisinuatus">
<emphasis id="BBE4EAC0FF83FFC4FBE8C4F42D821A78" bold="true" box="[1136,1276,1790,1814]" italics="true" pageId="5" pageNumber="338">S. bisinuatus</emphasis>
<bibRefCitation id="ED014B23FF83FFC4FA99C4F52CE21A79" author="SMETANA A." box="[1281,1436,1791,1815]" pageId="5" pageNumber="338" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="892F36D2FF83FFC4FCA4C5152CE31A19" blockId="5.[828,1437,191,2071]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FCA4C5152CE31A19" pageId="5" pageNumber="338">– Median impression of abdominal tergites III–V shallower or indistinct, laterally without two short oblique carinae. ......................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF83FFC4FCA4C5752CE31AD9" pageId="5" pageNumber="338">
<paragraph id="892F36D2FF83FFC4FCA4C5752CE31AF9" blockId="5.[828,1437,191,2071]" box="[828,1437,1919,1943]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FCA4C5752CE31AF9" box="[828,1437,1919,1943]" pageId="5" pageNumber="338">12 Legs reddish. .............................................................. 13</keyLead>
</paragraph>
<paragraph id="892F36D2FF83FFC4FCA4C5952CE31AD9" blockId="5.[828,1437,191,2071]" box="[828,1437,1951,1975]" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC4FCA4C5952CE31AD9" box="[828,1437,1951,1975]" pageId="5" pageNumber="338">– Legs blackish. ............................................................. 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF83FFC6FCA4C5B52B861AAE" lastPageId="7" lastPageNumber="340" pageId="5" pageNumber="338">
<paragraph id="892F36D2FF83FFC6FCA4C5B52B891A2E" blockId="5.[828,1437,191,2071]" lastBlockId="7.[151,761,1800,2080]" lastPageId="7" lastPageNumber="340" pageId="5" pageNumber="338">
<keyLead id="326195E7FF83FFC6FCA4C5B52B891A2E" lastPageId="7" lastPageNumber="340" pageId="5" pageNumber="338">
13 Posterior half of head with impunctate midline; punctation of head and pronotum mostly very dense and well delimited, interstices smooth and ridge-like. Ha- bitus: 
<figureCitation id="11AB2A57FF81FFC6FE9BC50228321A4E" box="[259,332,1800,1824]" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="7" pageNumber="340">Fig. 28</figureCitation>
. Aedeagus: 
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: Figs 20–22). 
<collectingCountry id="F1877642FF81FFC6FF5CC522287A1A2E" box="[196,260,1832,1856]" name="China" pageId="7" pageNumber="340">China</collectingCountry>
(
<collectingRegion id="4B54F830FF81FFC6FE8AC52228151A2E" box="[274,363,1832,1856]" country="China" name="Sichuan" pageId="7" pageNumber="340">Sichuan</collectingRegion>
). ................. 
<taxonomicName id="4E904D51FF81FFC6FE69C5222B891A2E" ID-CoL="4YPP7" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[497,759,1832,1856]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="BBE4EAC0FF81FFC6FE69C5222B291A2E" bold="true" box="[497,599,1832,1856]" italics="true" pageId="7" pageNumber="340">S. bicolor</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FDC5C5222B891A2E" author="SMETANA A." box="[605,759,1832,1856]" pageId="7" pageNumber="340" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<caption id="DDEF665AFF80FFC7FF0FC5D62DD0154A" ID-DOI="http://doi.org/10.5281/zenodo.3879947" ID-Zenodo-Dep="3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="6" pageNumber="339" startId="6.[151,188,2012,2031]" targetBox="[207,1381,193,1988]" targetPageId="6">
<paragraph id="892F36D2FF80FFC7FF0FC5D62DD0154A" blockId="6.[151,1437,2012,2085]" pageId="6" pageNumber="339">
Figs 24–31. Habitus of species of 
<taxonomicName id="4E904D51FF80FFC7FE50C5D62B391A81" ID-CoL="8W3PV" authority="Zhao &amp; Tang, 2020" authorityName="Zhao &amp; Tang" authorityYear="2020" box="[456,583,2012,2031]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBE4EAC0FF80FFC7FE50C5D62B391A81" box="[456,583,2012,2031]" italics="true" pageId="6" pageNumber="339">Sphaerobulbus</emphasis>
</taxonomicName>
. 24 – 
<taxonomicName id="4E904D51FF80FFC7FD1BC5D62A0F1A81" ID-CoL="4YPP6" authority="(Smetana, 2003)" baseAuthorityName="Smetana" baseAuthorityYear="2003" box="[643,881,2012,2031]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="339" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="BBE4EAC0FF80FFC7FD1BC5D62BA41A81" box="[643,730,2012,2031]" italics="true" pageId="6" pageNumber="339">S. alpinus</emphasis>
(
<bibRefCitation id="ED014B23FF80FFC7FD7FC5D62A151A81" author="SMETANA A." box="[743,875,2012,2031]" pageId="6" pageNumber="339" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
)
</taxonomicName>
(holotype); 25 – 
<taxonomicName id="4E904D51FF80FFC7FB96C5D62C791A81" ID-CoL="4YPP9" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[1038,1287,2012,2031]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="339" phylum="Arthropoda" rank="species" species="bisinuatus">
<emphasis id="BBE4EAC0FF80FFC7FB96C5D62D001A81" box="[1038,1150,2012,2031]" italics="true" pageId="6" pageNumber="339">S. bisinuatus</emphasis>
<bibRefCitation id="ED014B23FF80FFC7FB1DC5D62C791A81" author="SMETANA A." box="[1157,1287,2012,2031]" pageId="6" pageNumber="339" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
(holotype); 26 – 
<taxonomicName id="4E904D51FF80FFC7FF0FC5FD28FD1567" ID-CoL="4YPPC" authority="Smetana, 2010" authorityName="Smetana" authorityYear="2010" box="[151,387,2038,2058]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="339" phylum="Arthropoda" rank="species" species="cardinalis">
<emphasis id="BBE4EAC0FF80FFC7FF0FC5FD287F1567" box="[151,257,2038,2058]" italics="true" pageId="6" pageNumber="339">S. cardinalis</emphasis>
<bibRefCitation id="ED014B23FF80FFC7FE9DC5FC28FD1567" author="SMETANA A." box="[261,387,2038,2057]" pageId="6" pageNumber="339" pagination="241 - 246" refId="ref4867" refString="SMETANA A. 2010: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 24. The genus Sphaerobulbus Smetana 2003. Section 4. Studies and Reports, Taxonomical Series 6: 241 - 246." type="book chapter" year="2010">Smetana, 2010</bibRefCitation>
</taxonomicName>
(allotype); 27 – 
<taxonomicName id="4E904D51FF80FFC7FD94C5FD2B9A1567" ID-CoL="6YWBY" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[524,740,2038,2058]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="339" phylum="Arthropoda" rank="species" species="murzini">
<emphasis id="BBE4EAC0FF80FFC7FD94C5FD2B1C1567" box="[524,610,2039,2058]" italics="true" pageId="6" pageNumber="339">S. murzini</emphasis>
<bibRefCitation id="ED014B23FF80FFC7FDFEC5FC2B9A1567" author="SMETANA A." box="[614,740,2038,2057]" pageId="6" pageNumber="339" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
(holotype); 28 – 
<taxonomicName id="4E904D51FF80FFC7FCEBC5FD2D391567" ID-CoL="4YPP7" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[883,1095,2038,2058]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="339" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="BBE4EAC0FF80FFC7FCEBC5FD2ABB1567" box="[883,965,2038,2058]" italics="true" pageId="6" pageNumber="339">S. bicolor</emphasis>
<bibRefCitation id="ED014B23FF80FFC7FC51C5FC2D391567" author="SMETANA A." box="[969,1095,2038,2057]" pageId="6" pageNumber="339" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
(paratype); 29 – 
<taxonomicName id="4E904D51FF80FFC7FB4DC5FD29BC154A" ID-CoL="4YPP8" authority="Smetana, 2006" authorityName="Smetana" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="339" phylum="Arthropoda" rank="species" species="biplagiatus">
<emphasis id="BBE4EAC0FF80FFC7FB4DC5FD2C371567" box="[1237,1353,2038,2058]" italics="true" pageId="6" pageNumber="339">S. biplagiatus</emphasis>
<bibRefCitation id="ED014B23FF80FFC7FAD5C5FC29BC154A" author="SMETANA A." pageId="6" pageNumber="339" pagination="41 - 47" refId="ref4824" refString="SMETANA A. 2006: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 13. The genus Sphaerobulbus Smetana 2003. Section 3. Zootaxa 1318: 41 - 47." type="journal article" year="2006">Smetana, 2006</bibRefCitation>
</taxonomicName>
(holotype); 30 – 
<taxonomicName id="4E904D51FF80FFC7FECECA182B4B154A" ID-CoL="6Z8C9" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[342,565,2065,2085]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="339" phylum="Arthropoda" rank="species" species="brezinai">
<emphasis id="BBE4EAC0FF80FFC7FECECA1828CE154A" box="[342,432,2065,2085]" italics="true" pageId="6" pageNumber="339">S. brezinai</emphasis>
<bibRefCitation id="ED014B23FF80FFC7FE2ECA1B2B4B154A" author="SMETANA A." box="[438,565,2065,2084]" pageId="6" pageNumber="339" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
(holotype); 31 – 
<taxonomicName id="4E904D51FF80FFC7FD51CA182AD4154A" ID-CoL="4YPPG" authority="(Smetana, 2003)" baseAuthorityName="Smetana" baseAuthorityYear="2003" box="[713,938,2065,2085]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="339" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="BBE4EAC0FF80FFC7FD51CA182A66154A" box="[713,792,2066,2085]" italics="true" pageId="6" pageNumber="339">S. nigrita</emphasis>
(
<bibRefCitation id="ED014B23FF80FFC7FCBBCA1B2ADB154A" author="SMETANA A." box="[803,933,2065,2084]" pageId="6" pageNumber="339" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
)
</taxonomicName>
(holotype). Scale bars: 2 mm.
</paragraph>
</caption>
<caption id="DDEF665AFF81FFC6FF0FC49628EB1B8B" ID-DOI="http://doi.org/10.5281/zenodo.3879949" ID-Zenodo-Dep="3879949" httpUri="https://zenodo.org/record/3879949/files/figure.png" pageId="7" pageNumber="340" startId="7.[151,188,1692,1711]" targetBox="[182,1416,194,1655]" targetPageId="7">
<paragraph id="892F36D2FF81FFC6FF0FC49628EB1B8B" blockId="7.[151,1436,1692,1765]" pageId="7" pageNumber="340">
Figs 32–37. Habitus of species of 
<taxonomicName id="4E904D51FF81FFC6FE5AC4962B3F1BC1" ID-CoL="8W3PV" authority="Zhao &amp; Tang, 2020" authorityName="Zhao &amp; Tang" authorityYear="2020" box="[450,577,1692,1711]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBE4EAC0FF81FFC6FE5AC4962B3F1BC1" box="[450,577,1692,1711]" italics="true" pageId="7" pageNumber="340">Sphaerobulbus</emphasis>
</taxonomicName>
. 32 – 
<taxonomicName id="4E904D51FF81FFC6FDE3C4972A391BC1" ID-CoL="4YPPJ" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[635,839,1692,1712]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="BBE4EAC0FF81FFC6FDE3C4972BF21BDE" box="[635,652,1693,1712]" italics="true" pageId="7" pageNumber="340">S.</emphasis>
<emphasis id="BBE4EAC0FF81FFC6FD09C4972BC11BC1" box="[657,703,1693,1711]" italics="true" pageId="7" pageNumber="340">pusio</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FD5EC4962A391BC1" author="SMETANA A." box="[710,839,1692,1711]" pageId="7" pageNumber="340" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
(paratype); 33 – 
<taxonomicName id="4E904D51FF81FFC6FC47C4972DAD1BC1" ID-CoL="9KTZ3" authority="Smetana" authorityName="Smetana" authorityYear="2005" box="[991,1235,1692,1712]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="nagahatai">
<emphasis id="BBE4EAC0FF81FFC6FC47C4972A8E1BDE" box="[991,1008,1693,1712]" italics="true" pageId="7" pageNumber="340">S.</emphasis>
<emphasis id="BBE4EAC0FF81FFC6FC6DC4962D321BC1" box="[1013,1100,1692,1711]" italics="true" pageId="7" pageNumber="340">nagahatai</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FBCBC4962DAD1BC1" author="SMETANA A." box="[1107,1235,1692,1711]" pageId="7" pageNumber="340" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
(paratype); 34 – 
<taxonomicName id="4E904D51FF81FFC6FAF3C49728681BA4" ID-CoL="4YPPL" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="BBE4EAC0FF81FFC6FAF3C4972CE21BC1" box="[1387,1436,1693,1712]" italics="true" pageId="7" pageNumber="340">S. rex</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FF0FC4BD28681BA4" author="SMETANA A." box="[151,278,1719,1738]" pageId="7" pageNumber="340" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
(paratype); 35 – 
<taxonomicName id="4E904D51FF81FFC6FE30C4B22BED1BA4" ID-CoL="9KTZ7" authority="Smetana" authorityName="Smetana" authorityYear="2003" box="[424,659,1719,1739]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="BBE4EAC0FF81FFC6FE30C4B22B711BA4" box="[424,527,1720,1739]" italics="true" pageId="7" pageNumber="340">S. yunnanus</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FD8CC4BD2BED1BA4" author="SMETANA A." box="[532,659,1719,1738]" pageId="7" pageNumber="340" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
(paratype); 36 – 
<taxonomicName id="4E904D51FF81FFC6FCBCC4B22A811BA4" ID-CoL="4YPPH" authority="Smetana, 2006" authorityName="Smetana" authorityYear="2006" box="[804,1023,1719,1739]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="BBE4EAC0FF81FFC6FCBCC4B22A041BA4" box="[804,890,1720,1739]" italics="true" pageId="7" pageNumber="340">S. ornatus</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FC18C4BD2A811BA4" author="SMETANA A." box="[896,1023,1719,1738]" pageId="7" pageNumber="340" pagination="41 - 47" refId="ref4824" refString="SMETANA A. 2006: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 13. The genus Sphaerobulbus Smetana 2003. Section 3. Zootaxa 1318: 41 - 47." type="journal article" year="2006">Smetana, 2006</bibRefCitation>
</taxonomicName>
(paratype); 37 – 
<taxonomicName id="4E904D51FF81FFC6FB08C4B22CE21BA4" ID-CoL="4YPPN" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[1168,1436,1719,1739]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="yulongmontis">
<emphasis id="BBE4EAC0FF81FFC6FB08C4B22C661BA4" box="[1168,1304,1719,1739]" italics="true" pageId="7" pageNumber="340">S. yulongmontis</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FA86C4BD2CE21BA4" author="SMETANA A." box="[1310,1436,1719,1738]" pageId="7" pageNumber="340" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
(holotype). Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="892F36D2FF81FFC6FF0FC5432B861AAE" blockId="7.[151,761,1800,2080]" pageId="7" pageNumber="340">
<keyLead id="326195E7FF81FFC6FF0FC5432B861AAE" pageId="7" pageNumber="340">– Posterior half of head without impunctate midline; punctation of head and pronotum mostly less dense and slightly confluent, interstices microsculptured and not ridge-like. .................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF81FFC6FF0FC5C22CE31A4E" pageId="7" pageNumber="340">
<paragraph id="892F36D2FF81FFC6FF0FC5C22B89154E" blockId="7.[151,761,1800,2080]" pageId="7" pageNumber="340">
<keyLead id="326195E7FF81FFC6FF0FC5C22B89154E" pageId="7" pageNumber="340">
14 Head wider than long, HW/HL=1.11. Aedeagus: 
<bibRefCitation id="ED014B23FF81FFC6FF5CC5E22811156E" author="SMETANA A." box="[196,367,2024,2048]" pageId="7" pageNumber="340" pagination="235 - 243" refId="ref4915" refString="SMETANA A. 2016: Contributions to the knowledge of the Staphylinus-complex of China. (Coleoptera: Staphylinidae: Staphylinini). Part 27. The genus Sphaerobulbus Smetana, 2003, section 5. Studies and Reports, Taxonomical Series 12 (1): 235 - 243." type="journal article" year="2016">SMETANA (2016</bibRefCitation>
: Figs 3, 4). 
<collectingCountry id="F1877642FF81FFC6FE7DC5E22B58156E" box="[485,550,2024,2048]" name="China" pageId="7" pageNumber="340">China</collectingCountry>
(
<collectingRegion id="4B54F830FF81FFC6FDACC5E22BF0156E" box="[564,654,2024,2048]" country="China" name="Yunnan" pageId="7" pageNumber="340">Yunnan</collectingRegion>
). .............. ............................................... 
<taxonomicName id="4E904D51FF81FFC6FE6BCA022B89154E" ID-CoL="4YPPK" authority="Smetana, 2016" authorityName="Smetana" authorityYear="2016" box="[499,759,2056,2080]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="radani">
<emphasis id="BBE4EAC0FF81FFC6FE6BCA022B29154E" bold="true" box="[499,599,2056,2080]" italics="true" pageId="7" pageNumber="340">S. radani</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FDC5CA022B89154E" author="SMETANA A." box="[605,759,2056,2080]" pageId="7" pageNumber="340" pagination="235 - 243" refId="ref4915" refString="SMETANA A. 2016: Contributions to the knowledge of the Staphylinus-complex of China. (Coleoptera: Staphylinidae: Staphylinini). Part 27. The genus Sphaerobulbus Smetana, 2003, section 5. Studies and Reports, Taxonomical Series 12 (1): 235 - 243." type="journal article" year="2016">Smetana, 2016</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="892F36D2FF81FFC6FCA4C5032CE31A4E" blockId="7.[828,1437,1800,2080]" box="[828,1437,1800,1824]" pageId="7" pageNumber="340">
<keyLead id="326195E7FF81FFC6FCA4C5032CE31A4E" box="[828,1437,1800,1824]" pageId="7" pageNumber="340">– Head much wider than long, HW/HL=1.31. ........ 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF81FFC9FCA4C5222B891C59" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340">
<paragraph id="892F36D2FF81FFC6FCA4C5222CE21A8E" blockId="7.[828,1437,1800,2080]" pageId="7" pageNumber="340">
<keyLead id="326195E7FF81FFC6FCA4C5222CE21A8E" pageId="7" pageNumber="340">
15 Midanterior portion of head with sparse punctation; pronotum with broad impunctate midline; posterior basal line of abdominal tergites IV and V bisinuate. Habitus: 
<figureCitation id="11AB2A57FF81FFC6FC74C5822D461ACE" box="[1004,1080,1928,1952]" captionStart="Figs 32–37" captionStartId="7.[151,188,1692,1711]" captionTargetBox="[182,1416,194,1655]" captionTargetId="figure@7.[164,1424,189,1674]" captionTargetPageId="7" captionText="Figs 32–37. Habitus of species of Sphaerobulbus. 32 – S. pusio Smetana, 2005 (paratype); 33 – S. nagahatai Smetana, 2005 (paratype); 34 – S. rex Smetana, 2005 (paratype); 35 – S. yunnanus Smetana, 2003 (paratype); 36 – S. ornatus Smetana, 2006 (paratype); 37 – S. yulongmontis Smetana, 2003 (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879949" httpUri="https://zenodo.org/record/3879949/files/figure.png" pageId="7" pageNumber="340">Fig. 33</figureCitation>
. Aedeagus: 
<bibRefCitation id="ED014B23FF81FFC6FB21C5822C181ACE" author="SMETANA A." box="[1209,1382,1928,1952]" pageId="7" pageNumber="340" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">SMETANA (2005</bibRefCitation>
: Figs 3–5). 
<collectingCountry id="F1877642FF81FFC6FC3EC5A22A991AAE" box="[934,999,1960,1984]" name="China" pageId="7" pageNumber="340">China</collectingCountry>
(
<collectingRegion id="4B54F830FF81FFC6FC6DC5A22D2E1AAE" box="[1013,1104,1960,1984]" country="China" name="Sichuan" pageId="7" pageNumber="340">Sichuan</collectingRegion>
). ................................................... ......................................... 
<taxonomicName id="4E904D51FF81FFC6FBEBC5C22CE21A8E" ID-CoL="4YPPF" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[1139,1436,1992,2016]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="nagahatai">
<emphasis id="BBE4EAC0FF81FFC6FBEBC5C22D821A8E" bold="true" box="[1139,1276,1992,2016]" italics="true" pageId="7" pageNumber="340">S. nagahatai</emphasis>
<bibRefCitation id="ED014B23FF81FFC6FA99C5C22CE21A8E" author="SMETANA A." box="[1281,1436,1992,2016]" pageId="7" pageNumber="340" pagination="53 - 64" refId="ref4781" refString="SMETANA A. 2005: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa 1006: 53 - 64." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="892F36D2FF81FFC9FCA4C5E32B891C59" blockId="7.[828,1437,1800,2080]" lastBlockId="8.[151,760,191,1207]" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340">
<keyLead id="326195E7FF81FFC9FCA4C5E32B891C59" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340">
– Midanterior portion of head with dense punctation; pronotum with narrow impunctate midline; posteri- or basal line of abdominal tergites IV and V simple. Habitus: 
<figureCitation id="11AB2A57FF8EFFC9FEBDC2D5280E1D99" box="[293,368,223,247]" captionStart="Figs 32–37" captionStartId="7.[151,188,1692,1711]" captionTargetBox="[182,1416,194,1655]" captionTargetId="figure@7.[164,1424,189,1674]" captionTargetPageId="7" captionText="Figs 32–37. Habitus of species of Sphaerobulbus. 32 – S. pusio Smetana, 2005 (paratype); 33 – S. nagahatai Smetana, 2005 (paratype); 34 – S. rex Smetana, 2005 (paratype); 35 – S. yunnanus Smetana, 2003 (paratype); 36 – S. ornatus Smetana, 2006 (paratype); 37 – S. yulongmontis Smetana, 2003 (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879949" httpUri="https://zenodo.org/record/3879949/files/figure.png" pageId="8" pageNumber="341">Fig. 37</figureCitation>
. Aedeagus: 
<bibRefCitation id="ED014B23FF8EFFC9FE77C2D52BE51D99" author="SMETANA A." box="[495,667,223,247]" pageId="8" pageNumber="341" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">SMETANA (2003</bibRefCitation>
: 
<figureCitation id="11AB2A57FF8EFFC9FD3DC2D5299F1C79" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="8" pageNumber="341">Figs 27, 28</figureCitation>
). 
<collectingCountry id="F1877642FF8EFFC9FF6BC2F5284A1C79" box="[243,308,255,279]" name="China" pageId="8" pageNumber="341">China</collectingCountry>
(
<collectingRegion id="4B54F830FF8EFFC9FEDAC2F528E21C79" box="[322,412,255,279]" country="China" name="Yunnan" pageId="8" pageNumber="341">Yunnan</collectingRegion>
). ...................................................... .................................. 
<taxonomicName id="4E904D51FF8EFFC9FE30C3142B891C59" ID-CoL="4YPPN" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[424,759,286,311]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="yulongmontis">
<emphasis id="BBE4EAC0FF8EFFC9FE30C3142B291C58" bold="true" box="[424,599,286,310]" italics="true" pageId="8" pageNumber="341">S. yulongmontis</emphasis>
<bibRefCitation id="ED014B23FF8EFFC9FDC5C3152B891C59" author="SMETANA A." box="[605,759,287,311]" pageId="8" pageNumber="341" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF8EFFC9FF0FC3352B861C99" pageId="8" pageNumber="341">
<paragraph id="892F36D2FF8EFFC9FF0FC3352B891CD9" blockId="8.[151,760,191,1207]" pageId="8" pageNumber="341">
<keyLead id="326195E7FF8EFFC9FF0FC3352B891CD9" pageId="8" pageNumber="341">
16 Head and pronotum with dark blue hue; interstices of head and pronotum smooth. Aedeagus: 
<bibRefCitation id="ED014B23FF8EFFC9FD0AC355287D1CF9" author="SMETANA A." pageId="8" pageNumber="341" pagination="235 - 243" refId="ref4915" refString="SMETANA A. 2016: Contributions to the knowledge of the Staphylinus-complex of China. (Coleoptera: Staphylinidae: Staphylinini). Part 27. The genus Sphaerobulbus Smetana, 2003, section 5. Studies and Reports, Taxonomical Series 12 (1): 235 - 243." type="journal article" year="2016">SMETANA (2016</bibRefCitation>
, Figs 11–13). 
<collectingCountry id="F1877642FF8EFFC9FE0DC37528A81CF9" box="[405,470,383,407]" name="China" pageId="8" pageNumber="341">China</collectingCountry>
(
<collectingRegion id="4B54F830FF8EFFC9FE7CC3752B401CF9" box="[484,574,383,407]" country="China" name="Yunnan" pageId="8" pageNumber="341">Yunnan</collectingRegion>
). ........................... ............................................... 
<taxonomicName id="4E904D51FF8EFFC9FE6EC3942B891CD9" ID-CoL="4YPPD" authority="Smetana, 2016" authorityName="Smetana" authorityYear="2016" box="[502,759,414,439]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="BBE4EAC0FF8EFFC9FE6EC3942B291CD8" bold="true" box="[502,599,414,438]" italics="true" pageId="8" pageNumber="341">S. davidi</emphasis>
<bibRefCitation id="ED014B23FF8EFFC9FDC5C3952B891CD9" author="SMETANA A." box="[605,759,415,439]" pageId="8" pageNumber="341" pagination="235 - 243" refId="ref4915" refString="SMETANA A. 2016: Contributions to the knowledge of the Staphylinus-complex of China. (Coleoptera: Staphylinidae: Staphylinini). Part 27. The genus Sphaerobulbus Smetana, 2003, section 5. Studies and Reports, Taxonomical Series 12 (1): 235 - 243." type="journal article" year="2016">Smetana, 2016</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="892F36D2FF8EFFC9FF0FC3B52B861C99" blockId="8.[151,760,191,1207]" pageId="8" pageNumber="341">
<keyLead id="326195E7FF8EFFC9FF0FC3B52B861C99" pageId="8" pageNumber="341">– Head and pronotum without dark blue hue; interstices of head and pronotum microsculptured. ........ 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF8EFFC9FF0FC3F52B861F19" pageId="8" pageNumber="341">
<paragraph id="892F36D2FF8EFFC9FF0FC3F52B861F59" blockId="8.[151,760,191,1207]" pageId="8" pageNumber="341">
<keyLead id="326195E7FF8EFFC9FF0FC3F52B861F59" pageId="8" pageNumber="341">17 Superior and inferior lines of pronotal hypomeron widely separated at anterior angle. ......................... 18</keyLead>
</paragraph>
<paragraph id="892F36D2FF8EFFC9FF0FC0352B861F19" blockId="8.[151,760,191,1207]" pageId="8" pageNumber="341">
<keyLead id="326195E7FF8EFFC9FF0FC0352B861F19" pageId="8" pageNumber="341">– Superior and inferior lines of pronotal hypomeron joined at anterior angle. ............................................ 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="32642E77FF8EFFC9FF0FC0752B891E19" pageId="8" pageNumber="341">
<paragraph id="892F36D2FF8EFFC9FF0FC0752B891F99" blockId="8.[151,760,191,1207]" pageId="8" pageNumber="341">
<keyLead id="326195E7FF8EFFC9FF0FC0752B891F99" pageId="8" pageNumber="341">
18 Pronotum with punctation mostly well delimited and interstices even. Habitus: 
<figureCitation id="11AB2A57FF8EFFC9FE42C0952B581FD9" box="[474,550,671,695]" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="8" pageNumber="341">Fig. 31</figureCitation>
. Aedeagus: SMETA- NA (2003: Figs 82–84). 
<collectingCountry id="F1877642FF8EFFC9FE26C0B528811FB9" box="[446,511,703,727]" name="China" pageId="8" pageNumber="341">China</collectingCountry>
(
<collectingRegion id="4B54F830FF8EFFC9FD96C0B52B161FB9" box="[526,616,703,727]" country="China" name="Qinghai" pageId="8" pageNumber="341">Qinghai</collectingRegion>
, 
<collectingRegion id="4B54F830FF8EFFC9FDECC0B52BB41FB8" box="[628,714,703,726]" country="China" name="Yunnan" pageId="8" pageNumber="341">Yunnan</collectingRegion>
, 
<collectingRegion id="4B54F830FF8EFFC9FD4DC0B529841F99" country="China" name="Xizang" pageId="8" pageNumber="341">Xizang</collectingRegion>
). ................................. 
<taxonomicName id="4E904D51FF8EFFC9FE78C0D42B891F99" ID-CoL="4YPPG" authority="(Smetana, 2003)" baseAuthorityName="Smetana" baseAuthorityYear="2003" box="[480,759,734,759]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="BBE4EAC0FF8EFFC9FE78C0D42B3B1F98" bold="true" box="[480,581,734,758]" italics="true" pageId="8" pageNumber="341">S. nigrita</emphasis>
(
<bibRefCitation id="ED014B23FF8EFFC9FDCAC0D52B8E1F99" author="SMETANA A." box="[594,752,735,759]" pageId="8" pageNumber="341" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="892F36D2FF8EFFC9FF0FC0F52B891E19" blockId="8.[151,760,191,1207]" pageId="8" pageNumber="341">
<keyLead id="326195E7FF8EFFC9FF0FC0F52B891E19" pageId="8" pageNumber="341">
– Pronotum with punctation mostly confluent and interstices bumpy. Habitus: 
<figureCitation id="11AB2A57FF8EFFC9FE43C1152B581E59" box="[475,550,799,823]" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="8" pageNumber="341">Fig. 24</figureCitation>
. Aedeagus: SMETA- NA (2003: Figs 90, 91). 
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(
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). ....................... ........................................... 
<taxonomicName id="4E904D51FF8EFFC9FE42C1542B891E19" ID-CoL="4YPP6" authority="(Smetana, 2003)" baseAuthorityName="Smetana" baseAuthorityYear="2003" box="[474,759,862,887]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="BBE4EAC0FF8EFFC9FE42C1542B3B1E18" bold="true" box="[474,581,862,886]" italics="true" pageId="8" pageNumber="341">S. alpinus</emphasis>
(
<bibRefCitation id="ED014B23FF8EFFC9FDCAC1552B8E1E19" author="SMETANA A." box="[594,752,863,887]" pageId="8" pageNumber="341" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
)
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<paragraph id="892F36D2FF8EFFC9FF0FC1752B891979" blockId="8.[151,760,191,1207]" pageId="8" pageNumber="341">
<keyLead id="326195E7FF8EFFC9FF0FC1752B891979" pageId="8" pageNumber="341">
19 Sternite 9 with posterior margin deeply emarginate; median lobe of aedeagus less acute at apex, paramere wider. Habitus: 
<figureCitation id="11AB2A57FF8EFFC9FEEBC1B528BF1EB8" box="[371,449,958,983]" captionStart="Figs 32–37" captionStartId="7.[151,188,1692,1711]" captionTargetBox="[182,1416,194,1655]" captionTargetId="figure@7.[164,1424,189,1674]" captionTargetPageId="7" captionText="Figs 32–37. Habitus of species of Sphaerobulbus. 32 – S. pusio Smetana, 2005 (paratype); 33 – S. nagahatai Smetana, 2005 (paratype); 34 – S. rex Smetana, 2005 (paratype); 35 – S. yunnanus Smetana, 2003 (paratype); 36 – S. ornatus Smetana, 2006 (paratype); 37 – S. yulongmontis Smetana, 2003 (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879949" httpUri="https://zenodo.org/record/3879949/files/figure.png" pageId="8" pageNumber="341">Fig. 35</figureCitation>
. Aedeagus: 
<bibRefCitation id="ED014B23FF8EFFC9FDDEC1B52B8B1EB9" author="SMETANA A." box="[582,757,959,983]" pageId="8" pageNumber="341" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">SMETANA (2003</bibRefCitation>
: 
<figureCitation id="11AB2A57FF8EFFC9FF5CC1D5286F1E99" box="[196,273,991,1015]" captionStart="Figs 32–37" captionStartId="7.[151,188,1692,1711]" captionTargetBox="[182,1416,194,1655]" captionTargetId="figure@7.[164,1424,189,1674]" captionTargetPageId="7" captionText="Figs 32–37. Habitus of species of Sphaerobulbus. 32 – S. pusio Smetana, 2005 (paratype); 33 – S. nagahatai Smetana, 2005 (paratype); 34 – S. rex Smetana, 2005 (paratype); 35 – S. yunnanus Smetana, 2003 (paratype); 36 – S. ornatus Smetana, 2006 (paratype); 37 – S. yulongmontis Smetana, 2003 (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879949" httpUri="https://zenodo.org/record/3879949/files/figure.png" pageId="8" pageNumber="341">Figs 37</figureCitation>
–39). 
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<collectingRegion id="4B54F830FF8EFFC9FE03C1D5288B1E99" box="[411,501,991,1015]" country="China" name="Yunnan" pageId="8" pageNumber="341">Yunnan</collectingRegion>
). ....................................... ......................................... 
<taxonomicName id="4E904D51FF8EFFC9FE48C1F42B891979" ID-CoL="4YPPP" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[464,759,1022,1047]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="BBE4EAC0FF8EFFC9FE48C1F42B281978" bold="true" box="[464,598,1022,1046]" italics="true" pageId="8" pageNumber="341">S. yunnanus</emphasis>
<bibRefCitation id="ED014B23FF8EFFC9FDC5C1F52B891979" author="SMETANA A." box="[605,759,1023,1047]" pageId="8" pageNumber="341" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
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</paragraph>
<paragraph id="892F36D2FF8EFFC9FF0FC6152B8919D9" blockId="8.[151,760,191,1207]" pageId="8" pageNumber="341">
<keyLead id="326195E7FF8EFFC9FF0FC6152B8919D9" pageId="8" pageNumber="341">
– Sternite 9 with posterior margin shallowly emarginate; median lobe of aedeagus acute at apex, paramere narrower. Habitus: 
<figureCitation id="11AB2A57FF8EFFC9FE24C6552B751919" box="[444,523,1119,1143]" captionStart="Figs 24–31" captionStartId="6.[151,188,2012,2031]" captionTargetBox="[207,1381,193,1988]" captionTargetId="figure@6.[203,1385,189,1988]" captionTargetPageId="6" captionText="Figs 24–31. Habitus of species of Sphaerobulbus. 24 – S. alpinus (Smetana, 2003) (holotype); 25 – S. bisinuatus Smetana, 2003 (holotype); 26 – S. cardinalis Smetana, 2010 (allotype); 27 – S. murzini Smetana, 2003 (holotype); 28 – S. bicolor Smetana, 2005 (paratype); 29 – S. biplagiatus Smetana, 2006 (holotype); 30 – S. brezinai Smetana, 2003 (holotype); 31 – S. nigrita (Smetana, 2003) (holotype). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3879947" httpUri="https://zenodo.org/record/3879947/files/figure.png" pageId="8" pageNumber="341">Fig. 27</figureCitation>
. Aedeagus: 
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: Figs 41, 42). 
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(
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). ............................ ............................................. 
<taxonomicName id="4E904D51FF8EFFC9FE7EC6942B8919D9" ID-CoL="6YWBY" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[486,759,1182,1207]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="murzini">
<emphasis id="BBE4EAC0FF8EFFC9FE7EC6942B2919D8" bold="true" box="[486,599,1182,1206]" italics="true" pageId="8" pageNumber="341">S. murzini</emphasis>
<bibRefCitation id="ED014B23FF8EFFC9FDC5C6952B8919D9" author="SMETANA A." box="[605,759,1183,1207]" pageId="8" pageNumber="341" pagination="57 - 135" refId="ref4709" refString="SMETANA A. 2003: Contributions to the knowledge of the genera of the &quot; Staphylinus-complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n. Aulacocypus and Apecholinus, and comment on the genus Protocypus. Folia Heyrovskyana 11: 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
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