treatments-xml/data/01/23/56/0123566FFFEA2B3AFF1640B5FA8FF869.xml

<document id="F35C8BCCCC88DB9D7401F169A321A77E" ID-DOI="10.5281/zenodo.182422" ID-GBIF-Dataset="b7d81382-916a-4660-8a8f-4d5374fc0559" ID-ISSN="1175-5326" ID-Zenodo-Dep="182422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459861597832" checkinUser="plazi" docAuthor="Elmer, Kathryn R. &amp; Cannatella, David C." docDate="2008" docId="0123566FFFEA2B3AFF1640B5FA8FF869" docLanguage="en" docName="zt01784p038.pdf" docOrigin="Zootaxa 1784" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pristimantis kichwarum Elmer &amp; Cannatella, 2008, sp. nov." docType="treatment" docVersion="10" lastPageNumber="22" masterDocId="FD1A2E17FFED2B36FF814110FFC4FFE1" masterDocTitle="Three new species of leaflitter frogs from the upper Amazon forests: cryptic diversity within Pristimantis “ ockendeni ” (Anura: Strabomantidae) in Ecuador" masterLastPageNumber="38" masterPageNumber="11" pageNumber="18" updateTime="1698226162116" updateUser="plazi">
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<mods:title id="5A540007606328E187F8FAFE87321C24">Three new species of leaflitter frogs from the upper Amazon forests: cryptic diversity within Pristimantis “ ockendeni ” (Anura: Strabomantidae) in Ecuador</mods:title>
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<mods:namePart id="F52598D63F539AEEFF0B5A382A8253FC">Elmer, Kathryn R.</mods:namePart>
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<mods:namePart id="58EFBEABF737DF09CB64CB6D15B2FE2E">Cannatella, David C.</mods:namePart>
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<mods:date id="C66305EE6D60A37F03507D41F0B09329">2008</mods:date>
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<subSubSection id="C190B4F2FFEA2B31FF1640B5FE87FE03" pageId="7" pageNumber="18" type="nomenclature">
<paragraph id="8935E779FFEA2B31FF1640B5FDDAFE5E" blockId="7.[151,542,421,482]" box="[151,542,421,447]" pageId="7" pageNumber="18">
<heading id="D27D5015FFEA2B31FF1640B5FDDAFE5E" bold="true" box="[151,542,421,447]" fontSize="11" level="1" pageId="7" pageNumber="18" reason="1">
<emphasis id="BBFE3B6BFFEA2B31FF1640B5FDDAFE5E" bold="true" box="[151,542,421,447]" pageId="7" pageNumber="18">
<taxonomicName id="4E8A9CFAFFEA2B31FF1640B5FE7DFE5E" box="[151,441,421,447]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="18" phylum="Chordata" rank="species" species="kichwarum" status="sp. nov.">
<emphasis id="BBFE3B6BFFEA2B31FF1640B5FE7DFE5E" bold="true" box="[151,441,421,447]" italics="true" pageId="7" pageNumber="18">Pristimantis kichwarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0CD8610FFEA2B31FE4040B5FDDAFE5E" box="[449,542,421,447]" pageId="7" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8935E779FFEA2B31FF1640D8FE87FE03" blockId="7.[151,542,421,482]" box="[151,323,456,482]" pageId="7" pageNumber="18">
<figureCitation id="11B1FBFCFFEA2B31FF1640D8FEE6FE03" box="[151,290,456,482]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="23.[151,255,1686,1710]" captionStartId-1="24.[151,255,1718,1742]" captionStartId-2="22.[151,255,1371,1395]" captionTargetBox-0="[228,1364,196,1643]" captionTargetBox-1="[210,1406,207,1659]" captionTargetBox-2="[862,1407,212,1272]" captionTargetId-0="figure@23.[151,1436,94,1730]" captionTargetId-1="figure@24.[151,1436,116,1752]" captionTargetId-2="figure@22.[840,1407,181,1314]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="22" captionText-0="FIGURE 2. Dorsal view of three paratypes of P. altamnis (top row; QCAZ 25313, 25302, 25304), P. kichwarum (middle row; QCAZ 25287, 27762, 25782), and P. achuar (bottom row; QCAZ 25538, 25490, 25491). Note the presence of a canthal stripe in P. kichwarum and its absence in P. altamnis and P. achuar." captionText-1="FIGURE 3. Ventral view of three paratypes of P. altamnis (top row; QCAZ 25445, 25442, 25447), P. kichwarum (middle row; QCAZ 25287, 25558, 25759), and P. achuar (bottom row; QCAZ 25461, 25530, 25534). Note the difference in ventral colouration between the three species, with P. altamnis having the most dense flecking of melanophores and P. achuar having the palest venter." captionText-2="FIGURE 4. Lateral view of specimens of P. kichwarum (QCAZ 18033), P. achuar (QCAZ 25463), and P. altamnis (QCAZ 25311). Note the presence of a canthal stripe and darkened supratympanic ridge in P. kichwarum and their absence in P. achuar and P. altamnis." httpUri-0="https://zenodo.org/record/182426/files/figure.png" httpUri-1="https://zenodo.org/record/182427/files/figure.png" httpUri-2="https://zenodo.org/record/182425/files/figure.png" pageId="7" pageNumber="18">Figures 2–4</figureCitation>
, 
<figureCitation id="11B1FBFCFFEA2B31FEAF40D8FEFBFE03" box="[302,319,456,482]" captionStart="FIGURE 6" captionStartId="22.[151,255,1475,1499]" captionText="FIGURE 6. Eleutherodactylus kichwarum holotype (QCAZ 18128) in life." pageId="7" pageNumber="18">6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C190B4F2FFEA2B31FF164302FA43FC8D" pageId="7" pageNumber="18" type="materials_examined">
<paragraph id="8935E779FFEA2B31FF164302FAD4FD9D" blockId="7.[151,1436,530,956]" pageId="7" pageNumber="18">
<emphasis id="BBFE3B6BFFEA2B31FF164302FED4FDCD" bold="true" box="[151,272,530,556]" pageId="7" pageNumber="18">
<typeStatus id="563159DBFFEA2B31FF164302FECFFDCD" box="[151,267,530,556]" pageId="7" pageNumber="18" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="EF9B7FBCFFEA2B31FE9B4302FEAEFDCD" box="[282,362,530,556]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="7" pageNumber="18">QCAZ</collectionCode>
18128 (field number SC7783), an adult male collected at Jatun Sacha Biological Station, parroquía Ahuano, Napo province, 
<collectingCountry id="F19DA7E9FFEA2B31FDC3432AFD67FDB5" box="[578,675,570,596]" name="Ecuador" pageId="7" pageNumber="18">Ecuador</collectingCountry>
(
<geoCoordinate id="ECBE81BEFFEA2B31FD36432AFC8EFDB5" box="[695,842,570,596]" direction="south" orientation="latitude" pageId="7" pageNumber="18" precision="9" value="-1.0741667">S 01°04.45’</geoCoordinate>
, 
<geoCoordinate id="ECBE81BEFFEA2B31FCD8432AFC33FDB5" box="[857,1015,570,596]" direction="west" orientation="longitude" pageId="7" pageNumber="18" precision="9" value="-77.6155">W 77°36.93’</geoCoordinate>
, altitude 
<quantity id="4E724A9CFFEA2B31FBE9432AFB7CFDB5" box="[1128,1208,570,596]" metricMagnitude="2" metricUnit="m" metricValue="4.72" pageId="7" pageNumber="18" unit="m" value="472.0">472 m</quantity>
), on 
<date id="FD34C1B9FFEA2B31FB79432AFF15FD9D" pageId="7" pageNumber="18" value="2001-11-14">14 November 2001</date>
by KRE and F. Ayala-Varela. MtDNA cytochrome 
<emphasis id="BBFE3B6BFFEA2B31FCAF4372FCF9FD9A" box="[814,829,610,635]" italics="true" pageId="7" pageNumber="18">b</emphasis>
Genbank accession number 
<accessionNumber id="96D97A9AFFEA2B31FB0F4372FAC9FD9D" box="[1166,1293,610,636]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF581033" pageId="7" pageNumber="18">EF581033</accessionNumber>
.
</paragraph>
<paragraph id="8935E779FFEA2B31FF47439AFA43FC8D" blockId="7.[151,1436,530,956]" pageId="7" pageNumber="18">
<emphasis id="BBFE3B6BFFEA2B31FF47439AFE8FFD45" bold="true" box="[198,331,650,676]" pageId="7" pageNumber="18">
<typeStatus id="563159DBFFEA2B31FF47439AFE82FD45" box="[198,326,650,676]" pageId="7" pageNumber="18" type="paratype">Paratypes</typeStatus>
.
</emphasis>
(
<specimenCount id="9F8C2CF0FFEA2B31FEE1439AFDCFFD45" box="[352,523,650,676]" pageId="7" pageNumber="18" type="generic">45 specimens</specimenCount>
: 
<specimenCount id="9F8C2CF0FFEA2B31FD9A439AFD49FD45" box="[539,653,650,676]" pageId="7" pageNumber="18" type="male">29 males</specimenCount>
, 
<specimenCount id="9F8C2CF0FFEA2B31FD1E439AFCE3FD45" box="[671,807,650,676]" pageId="7" pageNumber="18" type="female">13 females</specimenCount>
, 
<specimenCount id="9F8C2CF0FFEA2B31FCB8439AFC04FD45" box="[825,960,650,676]" pageId="7" pageNumber="18" type="juvenile">3 juveniles</specimenCount>
) 
<collectionCode id="EF9B7FBCFFEA2B31FC54439AFBE1FD45" box="[981,1061,650,676]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="7" pageNumber="18">QCAZ</collectionCode>
18023, 18027, 18030, 18033, 18039, 18041, 18045, 18053, 18069, 18071, 18077, 18085, 18089, 18092, 18096, 18098, 18110, 18112, 18117, 18126, 18132, 18137, 18139, 18153, 18165, 25287, 25288, 25290, 25291, 25295, 25552, 25554, 25555, 25558, 25561, 25564, 25573, 25575, 25579, 25580, 25751, 25757, 25759, 25773, and 25782 were collected between 
<date id="FD34C1B9FFEA2B31FECE423AFDE2FCA5" box="[335,550,810,836]" pageId="7" pageNumber="18" value="2001-11-04">4 November 2001</date>
and 
<date id="FD34C1B9FFEA2B31FDE1423AFCC3FCA5" box="[608,775,810,836]" pageId="7" pageNumber="18" value="2003-04-14">14 April 2003</date>
from Jatun Sacha Biological Station and adjacent areas (
<taxonomicName id="4E8A9CFAFFEA2B31FF1F4242FECFFC8D" box="[158,267,850,876]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="31" phylum="Chordata" rank="species" species="librarius">Librarius</taxonomicName>
Study Center and Inner Vision Lodge [currently known as Arajuno Jungle Lodge]) (Appendix 1).
</paragraph>
</subSubSection>
<subSubSection id="C190B4F2FFEA2B3AFF47426AFA8FF869" lastPageId="12" lastPageNumber="23" pageId="7" pageNumber="18" type="description">
<paragraph id="8935E779FFEA2B31FF47426AFAE1FC5D" blockId="7.[151,1436,530,956]" pageId="7" pageNumber="18">
<emphasis id="BBFE3B6BFFEA2B31FF47426AFEFAFC75" bold="true" box="[198,318,890,916]" pageId="7" pageNumber="18">Referred.</emphasis>
Specimens collected adjacent to Jatun Sacha Biological Station (at Inner Vision Lodge) by KRE and T. Sugahara, QCAZ 25752 (male) 
<date id="FD34C1B9FFEA2B31FD2142B2FC81FC5D" box="[672,837,930,956]" pageId="7" pageNumber="18" value="2003-04-13">13 April 2003</date>
and QCAZ 25783 (male) 
<date id="FD34C1B9FFEA2B31FBF842B2FAE5FC5D" box="[1145,1313,930,956]" pageId="7" pageNumber="18" value="2003-04-17">17 April 2003</date>
.
</paragraph>
<paragraph id="8935E779FFEA2B31FF1642FDFEDDFBE6" blockId="7.[151,1436,1005,1191]" box="[151,281,1005,1031]" pageId="7" pageNumber="18">Etymology</paragraph>
<paragraph id="8935E779FFEA2B31FF474505FE1BFB46" blockId="7.[151,1436,1005,1191]" pageId="7" pageNumber="18">
This species is named for the indigenous Kichwa communities in the Oriente of 
<collectingCountry id="F19DA7E9FFEA2B31FBF54505FB1CFBCE" box="[1140,1240,1045,1071]" name="Ecuador" pageId="7" pageNumber="18">Ecuador</collectingCountry>
. They are recognized in this small collective way as a token of appreciation for the engagement of individuals and communities in this research, and for sharing their biodiversity and knowledge. The patronym is a Latin feminine plural genitive of the nation name.
</paragraph>
<paragraph id="8935E779FFEA2B31FF1645CDFEC8FB16" blockId="7.[151,1437,1245,1951]" box="[151,268,1245,1271]" pageId="7" pageNumber="18">Diagnosis</paragraph>
<paragraph id="8935E779FFEA2B31FF474415FEC0F87E" blockId="7.[151,1437,1245,1951]" pageId="7" pageNumber="18">
(1) Skin of dorsum smooth to finely shagreened with W-shaped occipital-scapular ridges, that of venter areolate; no obvious discoidal folds; dorsolateral fold absent; (2) tympanum and tympanic membrane distinct, tympanic annulus barely or slightly discernible, tympanum width approximately one-third eye width; supratympanic ridge dark; (3) snout subacuminate (weakly pointed) in dorsal view, rounded in lateral profile, canthus rostralis from anterior corner of eye to nostril slightly convex in lateral profile, concave in dorsal view, sharply angled in cross-section; (4) upper eyelid with or without two low tubercles; cranial crests absent; (5) odontophores moderately raised, oval, oblique; (6) males with moderately sized vocal slits, nuptial pads absent; (7) first finger shorter than second; all fingers bearing pads with circumferential grooves; discs slightly truncate, almost rounded, those on IV and III largest; (8) fingers slightly fringed; (9) ulnar tubercles very low or absent; (10) heel bearing small subconical tubercles; tarsal tubercles absent, tarsal fold weak; (11) inner metatarsal tubercle raised, elliptical, lateral margin more convex than medial margin, 4–5 times longer than conical outer metatarsal tubercle; few supernumerary plantar tubercles; (12) toes weakly fringed, lacking webbing; toe V much longer than III; discs approximately of equal size and shape as finger discs; (13) dorsum dark to light brown; W-shaped scapular ridge black in some, black dots at the lower points of the W in all; canthal stripe black; postsacral bar slightly or completely black; posterior surface of thighs brown; venter cream with some dark flecks; (14) adults small, males 17.5–21.9 mm (
<emphasis id="BBFE3B6BFFEA2B31FC60464DFC30F897" box="[993,1012,1885,1910]" italics="true" pageId="7" pageNumber="18">N</emphasis>
= 30), females 24.4–27.0 mm (
<emphasis id="BBFE3B6BFFEA2B31FAEE464DFA46F897" box="[1391,1410,1885,1910]" italics="true" pageId="7" pageNumber="18">N</emphasis>
= 13) SVL.
</paragraph>
<caption id="DDF5B7F1FFE52B3EFF164772FE06F810" httpUri="https://zenodo.org/record/182423/files/figure.png" pageId="8" pageNumber="19" targetBox="[151,1429,212,1567]" targetPageId="8">
<paragraph id="8935E779FFE52B3EFF164772FE06F810" blockId="8.[151,1436,1634,2033]" pageId="8" pageNumber="19">
<emphasis id="BBFE3B6BFFE52B3EFF164772FEDDF99B" bold="true" box="[151,281,1634,1658]" pageId="8" pageNumber="19">FIGURE 5.</emphasis>
Map of localities for reviewed specimens of 
<taxonomicName id="4E8A9CFAFFE52B3EFD7A4773FCB5F99B" box="[763,881,1635,1658]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="19" phylum="Chordata" rank="species" species="altamnis">
<emphasis id="BBFE3B6BFFE52B3EFD7A4773FCB5F99B" box="[763,881,1635,1658]" italics="true" pageId="8" pageNumber="19">P. altamnis</emphasis>
</taxonomicName>
(white dot), 
<taxonomicName id="4E8A9CFAFFE52B3EFC7A4773FB4FF99B" box="[1019,1163,1635,1658]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="19" phylum="Chordata" rank="species" species="kichwarum">
<emphasis id="BBFE3B6BFFE52B3EFC7A4773FB4FF99B" box="[1019,1163,1635,1658]" italics="true" pageId="8" pageNumber="19">P. kichwarum</emphasis>
</taxonomicName>
(grey dot), and 
<taxonomicName id="4E8A9CFAFFE52B3EFAB74773FA58F99B" box="[1334,1436,1635,1658]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="19" phylum="Chordata" rank="species" species="achuar">
<emphasis id="BBFE3B6BFFE52B3EFAB74773FA58F99B" box="[1334,1436,1635,1658]" italics="true" pageId="8" pageNumber="19">P. achuar</emphasis>
</taxonomicName>
(black dot). The general area of the Cuyabeno Reserve and Yasuní National Park are noted, as mentioned in the text for some collections. For simplicity, some nearby localities share a dot. (1) Chonta Yacu, Lumbaqui; (2) Centro Shuar Yawi; (3) Serena; (4) Talag or Cando; (5) Jatun Sacha Biological Station and Arajuno Jungle Lodge; (6) Puca Chicta, Misahuallí; (7) Auca 14 Rd near Dayuma; (8) Parque Nacional Yasuní: Estación Científica, Onkonegare, and pozo Capiron; (9) Llanganates Mountains, Canton Arosamena Tola; (10) La Selva Lodge, Pañacocha; (11) Y of road Tena to Puyo, Canton Sta Clara; (12) Fundación Hola Vida, El Porvenir; (13) Kapawi Lodge; (14) Lorocachi; (15) Petrolera Garza 1, Montalvo; (16) Bloque Shiripuno, Parque Nacional Yasuní; (17) Rio Huataracu, via Hollín-Loreto; (18) Indillama; (19) Yuturi; (20) Bataburo Lodge, Rio Tiguño; (21) Santa Rita; (22) Parque Nacional Yasuní, Carretera Pompeya Sur-Iro Km 21, 38 or 41; (23) Comuna El Descanso; (24) Parque Nacional Yasuní Road to MAXUS km 107 and AMO 1; (25) Parque Nacional Yasuní Bloque 31.
</paragraph>
</caption>
<paragraph id="8935E779FFE42B3FFF164188FEE4FF53" blockId="9.[151,1437,152,1658]" box="[151,288,152,178]" pageId="9" pageNumber="20">Description</paragraph>
<paragraph id="8935E779FFE42B3FFF4741D0FE7CFC33" blockId="9.[151,1437,152,1658]" pageId="9" pageNumber="20">
Head slightly wider than body, approximately as long as wide; head width of females 36–39% SVL (x = 37%, 
<emphasis id="BBFE3B6BFFE42B3FFF5A41F8FF2AFEE0" box="[219,238,232,257]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 5), of males 33–39% SVL (x = 36%, 
<emphasis id="BBFE3B6BFFE42B3FFD3E41F8FD16FEE0" box="[703,722,232,257]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 9); snout subacuminate (weakly pointed) in dorsal view, in lateral view rounded; upper jaw overhanging lower jaw at anterior tip; snout of moderate length, eye-nostril distance of females 90–98% of eye width (x = 94%, 
<emphasis id="BBFE3B6BFFE42B3FFD7C4028FCD4FEB0" box="[765,784,312,337]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 5), of males 78–104% (x = 93%, 
<emphasis id="BBFE3B6BFFE42B3FFB354028FB03FEB0" box="[1204,1223,312,337]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 9); canthus rostralis from anterior corner of eye to nostril slightly convex in lateral profile, canthus rostralis concave in dorsal view, sharply angled in cross-section; loreal region slightly concave, lips not flared; nostrils slightly protuberant, directed laterally with slight dorsal inclination; interorbital space flat; cranial crests absent, interorbital distance in females 49–58% of head width (x = 53%, 
<emphasis id="BBFE3B6BFFE42B3FFCA140C8FCF7FE10" box="[800,819,472,497]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 5), in males 52–61% (x = 57%, 
<emphasis id="BBFE3B6BFFE42B3FFB5B40C8FB29FE10" box="[1242,1261,472,497]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 9); two low tubercles on mid eyelids in some specimens; temporal region vertical, sloping slightly laterally towards jaw; supratympanic ridge, darkened, not obscuring tympanum, postrictal tubercle present; tympanum visible, tympanic membrane distinct, tympanic annulus barely or slightly discernable, in females tympanum width 27– 44% of eye width (x = 34%, 
<emphasis id="BBFE3B6BFFE42B3FFE704368FDC0FD70" box="[497,516,632,657]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 5), in males tympanum width 29–43% of eye width (x = 33%, 
<emphasis id="BBFE3B6BFFE42B3FFA904368FAE0FD70" box="[1297,1316,632,657]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 9), size and shape not sexually dimorphic; distance from eye to tympanum less than diameter of tympanum; choanae medium-sized, round, moderately elliptical, well within borders of jaws; odontophores moderately raised and oblique (less raised in females), slightly larger than and posterior to choanae, separated medially by 2–3 times width of odontophore, females with more teeth than males (median 5 left/5 right in females, median 2 left/2 right in males), teeth of males somewhat covered by mucosal tissue in some specimens; tongue longer than wide, with slight posterior notch, attached at front of mouth, posteriorly and laterally not adherent to floor of mouth; vocal slits lateral, moderate sized; in some male specimens slightly distended subgular vocal sac visible. Nuptial pads absent.
</paragraph>
<paragraph id="8935E779FFE42B3FFF4742F0FCA6FA83" blockId="9.[151,1437,152,1658]" pageId="9" pageNumber="20">Skin of dorsum smooth to finely shagreened; skin of venter areolate, skin of flanks shagreened; no apparent discoidal folds (that can be distinguished from preservation artefact); pair of W-shaped prominent ridges from posterior eyelid to scapular region, two posterior scapular tubercles located at base of W; postsacral bar with low tubercles; small (generally low) tubercles on elbow, smooth forearm, ulnar tubercles low or absent; palmar tubercle bifid, crease at base of U in most specimens, larger than elongate thenar tubercle; elongate palmar tubercles smaller than subarticular tubercles; subarticular tubercles elliptical, raised and flat in lateral view, supernumerary tubercles on finger I and II; distal subarticular tubercles flattened and rounded; proximal subarticular tubercles subconical; fingers with weak lateral fringes; all fingers bearing discs on pads, all pads wider than long, slightly truncate, equally distinct on fingers IV-II, pads in decreasing order of size IV=III-II-I (barely wider than digit); decreasing finger length III-IV-II-I.</paragraph>
<paragraph id="8935E779FFE42B3FFF474460FF2DF99B" blockId="9.[151,1437,152,1658]" pageId="9" pageNumber="20">
Heel bearing one or two small subconical tubercles; inner and outer edge of tarsus lacking tubercles; tarsal fold weak; inner metatarsal tubercle raised, elliptical and lateral margin more convex than medial margin, 4–5 times longer than smaller conical outer metatarsal tubercle; subarticular tubercles subconical, most distal subarticular tubercles flatter and smaller; toes weakly fringed, lacking webbing; toe discs wider than long, slightly truncate, approximately equal size to finger discs, in order of descending size, IV-V=III-II-I; hindlimbs of moderate length, shank of female 53–58% SVL (x = 55%, 
<emphasis id="BBFE3B6BFFE42B3FFC374728FC0DF9B0" box="[950,969,1592,1617]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 5), of males 54–60% SVL (x = 58%, 
<emphasis id="BBFE3B6BFFE42B3FFF164770FF6EF998" box="[151,170,1632,1657]" italics="true" pageId="9" pageNumber="20">N</emphasis>
= 9).
</paragraph>
<paragraph id="8935E779FFE42B3FFF1647A0FE5BF92B" blockId="9.[151,1436,1712,2018]" box="[151,415,1712,1738]" pageId="9" pageNumber="20">Colour in preservation</paragraph>
<paragraph id="8935E779FFE42B3CFF4747C8FBB8FE9B" blockId="9.[151,1436,1712,2018]" lastBlockId="10.[151,1437,152,378]" lastPageId="10" lastPageNumber="21" pageId="9" pageNumber="20">Central dorsum dark to light brown, paler on flanks, diagonal brown stripes on flanks, thin white mid-dorsal line extending from tip of snout to vent in some (e.g. QCAZ 18030, 18110, 18126, 25554, 25558); sacral chevron present in some, dark anal triangle present; scapular W-shaped ridge, outer lines of W black, inner lines of W black in some, and/or with white along ridge (in individuals with thin white snout-to-vent line), base of W with two black dots on each side separate or longitudinally joined; background colour of head and scapular region brown or grey, approximately same colour as dorsum; canthal stripe black; prominent dark labial bar at anterior margin of eye; less prominent labial bar at posterior margin of eye in some; supratympanic ridge with thin black line; eyelids grey; dark band at mid-forearm, upper arm lacking markings, elbow with dark spot in some; brown bars on thigh, shank, and tarsus; anterior and posterior surfaces of thighs uniform brown; postsacral dorsal bar partially or fully black; discs of fingers and toes paler than remainder of digits; venter cream (under microscope small deposits of guanine pigment visible on ventral areolar warts) with small dark spotting, more concentrated around jaw and flank; ventral surface of thighs brown, mid-ventral region of thigh and shank with absence of melanophores producing a lighter line.</paragraph>
<paragraph id="8935E779FFE72B3CFF1640A0FEF1FE2B" blockId="10.[151,1437,432,858]" box="[151,309,432,458]" pageId="10" pageNumber="21">Colour in life</paragraph>
<paragraph id="8935E779FFE72B3CFF4740C8FB66FCBB" blockId="10.[151,1437,432,858]" pageId="10" pageNumber="21">Dorsum light to medium brown, lighter laterally, occasionally with posterio-lateral stripes and/or reticulate pattern, with or without darker mid-dorsal chevron(s); some specimens with reddish tinge lateral to W and around upper arms (e.g. QCAZ 18085, 18165), some specimens with random grey dorsal flecks (e.g. QCAZ 18033, 18039, 18069, 18077, 18085, 18132, 18137, 18165), some specimens with thin white snout-vent line (e.g. QCAZ 18030, 18110, 18126, 25554, 25558); points at base of scapular W black, W-shaped ridge most commonly black (except, e.g. QCAZ 18071, 18092); some specimens with lighter brown patch extending from nose to top of head (between eyes) (e.g. QCAZ 18030, 18077, 18085, 18139) or to W (e.g. QCAZ 18117), or extending from top of head (between eyes) to W (e.g. QCAZ 18069); venter cream, lightly specked especially around throat, throat tinged with yellow in some males (e.g. QCAZ 25552, 25555, 25558, 25564). Iris is bronze with a darker lateral stripe, lighter bronze above and darker below median.</paragraph>
<paragraph id="8935E779FFE72B3CFF164280FE1DFC4B" blockId="10.[151,1437,912,1178]" box="[151,473,912,938]" pageId="10" pageNumber="21">
Description of the 
<typeStatus id="563159DBFFE72B3CFEF34280FE1DFC4B" box="[370,473,912,938]" pageId="10" pageNumber="21" type="holotype">holotype</typeStatus>
</paragraph>
<paragraph id="8935E779FFE72B3CFF4742A8FE9EFB7B" blockId="10.[151,1437,912,1178]" pageId="10" pageNumber="21">
Measurements (in mm): SVL 20.2, tibia length 12.3, foot length 15.7, head width 7.6, head length 7.9, IOD 3.9, 
<collectingCountry id="F19DA7E9FFE72B3CFE8442F0FE85FC1B" box="[261,321,992,1018]" name="India" pageId="10" pageNumber="21">END</collectingCountry>
2.9, eye width 2.6, tympanum width 0.9. Colour in preservative: medium brown with reticulate patterning, scapular W marking white with very thin black outline, base points of W black, thin white snoutvent line, finger and toe pads lighter than digits. Right foot of the 
<typeStatus id="563159DBFFE72B3CFC384520FBDBFBAB" box="[953,1055,1072,1098]" pageId="10" pageNumber="21" type="holotype">holotype</typeStatus>
with unusually small toe discs. Colour in life: “mottled medium dark brown dorsum, black and light (paler) outlines of scapular W, thin white snout-vent line”.
</paragraph>
<paragraph id="8935E779FFE72B3CFF1645C0FEB9FB0B" blockId="10.[151,1436,1232,1418]" box="[151,381,1232,1258]" pageId="10" pageNumber="21">Molecular diversity</paragraph>
<paragraph id="8935E779FFE72B3CFF4745E8FCE4FA6B" blockId="10.[151,1436,1232,1418]" pageId="10" pageNumber="21">
MtDNA cytochrome 
<emphasis id="BBFE3B6BFFE72B3CFE4545E8FE17FAF0" box="[452,467,1272,1297]" italics="true" pageId="10" pageNumber="21">b</emphasis>
sequences examined for 44 
<typeStatus id="563159DBFFE72B3CFCA945E8FC49FAF3" box="[808,909,1272,1298]" pageId="10" pageNumber="21" type="paratype">paratype</typeStatus>
specimens and the 
<typeStatus id="563159DBFFE72B3CFBF845E8FB1BFAF3" box="[1145,1247,1272,1298]" pageId="10" pageNumber="21" type="holotype">holotype</typeStatus>
resulted in five haplotypes with a maximum uncorrected 
<emphasis id="BBFE3B6BFFE72B3CFD024430FD56FAD8" box="[643,658,1312,1337]" italics="true" pageId="10" pageNumber="21">p</emphasis>
-distance between them of 0.63%. MtDNA 16S sequences for six 
<typeStatus id="563159DBFFE72B3CFF164458FEC3FA83" box="[151,263,1352,1378]" pageId="10" pageNumber="21" type="paratype">paratypes</typeStatus>
resulted in three haplotypes with a maximum uncorrected 
<emphasis id="BBFE3B6BFFE72B3CFC3A4458FC0EFA80" box="[955,970,1352,1377]" italics="true" pageId="10" pageNumber="21">p</emphasis>
-distance of 0.39%. See Appendix 1 for cytochrome 
<emphasis id="BBFE3B6BFFE72B3CFEA94460FEF3FA68" box="[296,311,1392,1417]" italics="true" pageId="10" pageNumber="21">b</emphasis>
and/or 16S GenBank accession numbers.
</paragraph>
<paragraph id="8935E779FFE72B3CFF1644D0FE83FA3B" blockId="10.[151,1436,1472,1658]" box="[151,327,1472,1498]" pageId="10" pageNumber="21">Natural history</paragraph>
<paragraph id="8935E779FFE72B3CFF4744F8FA92F99B" blockId="10.[151,1436,1472,1658]" pageId="10" pageNumber="21">
Individuals were found in primary and secondary forest. By night they were found perched on the leaves or stems of low vegetation from 0.2 to 1.5 m elevation. By day they were found in the leaflitter. The following specimens were collected in amplexus: QCAZ 25287 (
<typeStatus id="563159DBFFE72B3CFCA44728FC48F9B3" box="[805,908,1592,1618]" pageId="10" pageNumber="21" type="paratype">paratype</typeStatus>
female) with 25288 (
<typeStatus id="563159DBFFE72B3CFB0E4728FB32F9B3" box="[1167,1270,1592,1618]" pageId="10" pageNumber="21" type="paratype">paratype</typeStatus>
male), 25751 (
<typeStatus id="563159DBFFE72B3CFF1F4770FEC1F99B" box="[158,261,1632,1658]" pageId="10" pageNumber="21" type="paratype">paratype</typeStatus>
female) with 25752 (referred male), and 25782 (
<typeStatus id="563159DBFFE72B3CFCC34770FC6DF99B" box="[834,937,1632,1658]" pageId="10" pageNumber="21" type="paratype">paratype</typeStatus>
female) with 25783 (referred male).
</paragraph>
<paragraph id="8935E779FFE72B3CFF1647A0FEE1F92B" blockId="10.[151,1437,1712,1938]" box="[151,293,1712,1738]" pageId="10" pageNumber="21">Distribution</paragraph>
<paragraph id="8935E779FFE72B3CFF4747C8FB89F873" blockId="10.[151,1437,1712,1938]" pageId="10" pageNumber="21">
<taxonomicName id="4E8A9CFAFFE72B3CFF4747C8FE20F910" box="[198,484,1752,1777]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="21" phylum="Chordata" rank="species" species="kichwarum">
<emphasis id="BBFE3B6BFFE72B3CFF4747C8FE20F910" box="[198,484,1752,1777]" italics="true" pageId="10" pageNumber="21">Pristimantis kichwarum</emphasis>
</taxonomicName>
has been found throughout the humid tropical forests of eastern lowlands of 
<collectingCountry id="F19DA7E9FFE72B3CFF164610FF3DF8FB" box="[151,249,1792,1818]" name="Ecuador" pageId="10" pageNumber="21">Ecuador</collectingCountry>
in the Napo, Orellana, and Sucumbíos provinces, at elevations of approximately 500 masl and below (see 
<figureCitation id="11B1FBFCFFE72B3CFF534638FED9F8A3" box="[210,285,1832,1858]" captionStart="FIGURE 5" captionStartId="8.[151,255,1634,1658]" captionTargetBox="[151,1429,212,1567]" captionTargetId="figure@8.[151,1436,194,1610]" captionTargetPageId="8" captionText="FIGURE 5. Map of localities for reviewed specimens of P. altamnis (white dot), P. kichwarum (grey dot), and P. achuar (black dot). The general area of the Cuyabeno Reserve and Yasuní National Park are noted, as mentioned in the text for some collections. For simplicity, some nearby localities share a dot. (1) Chonta Yacu, Lumbaqui; (2) Centro Shuar Yawi; (3) Serena; (4) Talag or Cando; (5) Jatun Sacha Biological Station and Arajuno Jungle Lodge; (6) Puca Chicta, Misahuallí; (7) Auca 14 Rd near Dayuma; (8) Parque Nacional Yasuní: Estación Científica, Onkonegare, and pozo Capiron; (9) Llanganates Mountains, Canton Arosamena Tola; (10) La Selva Lodge, Pañacocha; (11) Y of road Tena to Puyo, Canton Sta Clara; (12) Fundación Hola Vida, El Porvenir; (13) Kapawi Lodge; (14) Lorocachi; (15) Petrolera Garza 1, Montalvo; (16) Bloque Shiripuno, Parque Nacional Yasuní; (17) Rio Huataracu, via Hollín-Loreto; (18) Indillama; (19) Yuturi; (20) Bataburo Lodge, Rio Tiguño; (21) Santa Rita; (22) Parque Nacional Yasuní, Carretera Pompeya Sur-Iro Km 21, 38 or 41; (23) Comuna El Descanso; (24) Parque Nacional Yasuní Road to MAXUS km 107 and AMO 1; (25) Parque Nacional Yasuní Bloque 31." httpUri="https://zenodo.org/record/182423/files/figure.png" pageId="10" pageNumber="21">Fig. 5</figureCitation>
). The extent of its distribution to the east and north is not known. The species is apparently replaced by 
<taxonomicName id="4E8A9CFAFFE72B3CFEA94640FE5DF888" box="[296,409,1872,1897]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="21" phylum="Chordata" rank="species" species="achuar">
<emphasis id="BBFE3B6BFFE72B3CFEA94640FE5DF888" box="[296,409,1872,1897]" italics="true" pageId="10" pageNumber="21">P. achuar</emphasis>
</taxonomicName>
to the south in the Pastaza basin. In the upper regions of the Napo River (Jatun Sacha Biological Station), 
<taxonomicName id="4E8A9CFAFFE72B3CFE024668FDE6F870" box="[387,546,1912,1937]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="21" phylum="Chordata" rank="species" species="kichwarum">
<emphasis id="BBFE3B6BFFE72B3CFE024668FDE6F870" box="[387,546,1912,1937]" italics="true" pageId="10" pageNumber="21">P. kichwarum</emphasis>
</taxonomicName>
has been found sympatrically with 
<taxonomicName id="4E8A9CFAFFE72B3CFC454668FB82F870" box="[964,1094,1912,1937]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="21" phylum="Chordata" rank="species" species="altamnis">
<emphasis id="BBFE3B6BFFE72B3CFC454668FB82F870" box="[964,1094,1912,1937]" italics="true" pageId="10" pageNumber="21">P. altamnis</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DDF5B7F1FFE62B3DFF164187FC3DFF30" ID-Table-UUID="DDF5B7F1FFE62B3DFF164187FC3DFF30" httpUri="http://table.plazi.org/id/DDF5B7F1FFE62B3DFF164187FC3DFF30" pageId="11" pageNumber="22" targetBox="[194,1413,255,701]" targetIsTable="true" targetPageId="11">
<paragraph id="8935E779FFE62B3DFF164187FC3DFF30" blockId="11.[151,1436,151,209]" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFF164187FECDFF4E" bold="true" box="[151,265,151,175]" pageId="11" pageNumber="22">TABLE 1.</emphasis>
Morphological characters used to distinguish various upper Amazonian or east Andean 
<taxonomicName id="4E8A9CFAFFE62B3DFB374188FAB3FF4E" box="[1206,1399,152,175]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="BBFE3B6BFFE62B3DFB374188FAB3FF4E" box="[1206,1399,152,175]" italics="true" pageId="11" pageNumber="22">Eleutherodactylus</emphasis>
</taxonomicName>
(+ indicates presence, - indicates absence,? indicates not available or inconclusive).
</paragraph>
</caption>
<paragraph id="8935E779FFE62B3DFE0841EFFA9BFD42" pageId="11" pageNumber="22">
<table id="FB8A15D9FFE6D4C9FF4341EFFA41FD5C" box="[194,1413,255,701]" gridcols="7" gridrows="5" pageId="11" pageNumber="22">
<tr id="37BAE53BFFE6D4C9FF4341EFFA41FECE" box="[194,1413,255,303]" gridrow="0" pageId="11" pageNumber="22" rowspan-0="1">
<th id="746B8C47FFE6D4C9FE0841EFFDE1FECE" box="[393,549,255,303]" gridcol="1" gridrow="0" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFE0841EFFE32FECE" bold="true" pageId="11" pageNumber="22">W on scapular region</emphasis>
</th>
<th id="746B8C47FFE6D4C9FDED41EFFD21FECE" box="[620,741,255,303]" gridcol="2" gridrow="0" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFDF241EFFD24FEF5" bold="true" box="[627,736,255,276]" pageId="11" pageNumber="22">SVL (mm)</emphasis>
</th>
<th id="746B8C47FFE6D4C9FCAE41EFFC46FECE" box="[815,898,255,303]" gridcol="3" gridrow="0" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFCAE41EFFCB1FECE" bold="true" box="[815,898,255,303]" pageId="11" pageNumber="22">Canthal stripe</emphasis>
</th>
<th id="746B8C47FFE6D4C9FC2B41EFFBC3FECE" box="[938,1031,255,303]" gridcol="4" gridrow="0" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFC3941EFFBC3FECE" bold="true" pageId="11" pageNumber="22">Eyelid tubercles</emphasis>
</th>
<th id="746B8C47FFE6D4C9FBB041EFFB69FECE" box="[1073,1197,255,303]" gridcol="5" gridrow="0" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFBB541EFFB6EFEF5" bold="true" box="[1076,1194,255,276]" pageId="11" pageNumber="22">Tympanum</emphasis>
</th>
<th id="746B8C47FFE6D4C9FB5641EFFA41FECE" box="[1239,1413,255,303]" gridcol="6" gridrow="0" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFB6B41EFFA41FECE" bold="true" pageId="11" pageNumber="22">Thigh colour, posterior surface</emphasis>
</th>
</tr>
<tr id="37BAE53BFFE6D4C9FF43405FFA41FE78" box="[194,1413,335,409]" gridrow="1" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF43405FFE8CFE78" box="[194,328,335,409]" gridcol="0" gridrow="1" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF56405FFEF1FE85" box="[215,309,335,356]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="achuar">
<emphasis id="BBFE3B6BFFE62B3DFF56405FFEF1FE85" bold="true" box="[215,309,335,356]" italics="true" pageId="11" pageNumber="22">P. achuar</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE08405FFDE1FE78" box="[393,549,335,409]" gridcol="1" gridrow="1" pageId="11" pageNumber="22">present as thin ridge, usually black</td>
<td id="746B8C47FFE6D4C9FDED405FFD21FE78" box="[620,741,335,409]" gridcol="2" gridrow="1" pageId="11" pageNumber="22">Ψ 19.6-23.5; ɗ 14.3-16.6</td>
<td id="746B8C47FFE6D4C9FCAE405FFC46FE78" box="[815,898,335,409]" gridcol="3" gridrow="1" pageId="11" pageNumber="22">-</td>
<td id="746B8C47FFE6D4C9FC2B405FFBC3FE78" box="[938,1031,335,409]" gridcol="4" gridrow="1" pageId="11" pageNumber="22">+</td>
<td id="746B8C47FFE6D4C9FBB0405FFB69FE78" box="[1073,1197,335,409]" gridcol="5" gridrow="1" pageId="11" pageNumber="22">visible</td>
<td id="746B8C47FFE6D4C9FB56405FFA41FE78" box="[1239,1413,335,409]" gridcol="6" gridrow="1" pageId="11" pageNumber="22">uniform brown</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF4340A9FA41FDE5" box="[194,1413,441,516]" gridrow="2" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF4340A9FE8CFDE5" box="[194,328,441,516]" gridcol="0" gridrow="2" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFF4E40A9FEFDFE2F" bold="true" box="[207,313,441,462]" italics="true" pageId="11" pageNumber="22">P. a l ta m n i s</emphasis>
</th>
<td id="746B8C47FFE6D4C9FE0840A9FDE1FDE5" box="[393,549,441,516]" gridcol="1" gridrow="2" pageId="11" pageNumber="22">present as thin ridge, usually black</td>
<td id="746B8C47FFE6D4C9FDED40A9FD21FDE5" box="[620,741,441,516]" gridcol="2" gridrow="2" pageId="11" pageNumber="22">Ψ 18.8-28.3; ɗ 15.1-22.3</td>
<td id="746B8C47FFE6D4C9FCAE40A9FC46FDE5" box="[815,898,441,516]" gridcol="3" gridrow="2" pageId="11" pageNumber="22">-</td>
<td id="746B8C47FFE6D4C9FC2B40A9FBC3FDE5" box="[938,1031,441,516]" gridcol="4" gridrow="2" pageId="11" pageNumber="22">+</td>
<td id="746B8C47FFE6D4C9FBB040A9FB69FDE5" box="[1073,1197,441,516]" gridcol="5" gridrow="2" pageId="11" pageNumber="22">visible</td>
<td id="746B8C47FFE6D4C9FB5640A9FA41FDE5" box="[1239,1413,441,516]" gridcol="6" gridrow="2" pageId="11" pageNumber="22">uniform brown</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF434333FA41FD8F" box="[194,1413,547,622]" gridrow="3" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF434333FE8CFD8F" box="[194,328,547,622]" gridcol="0" gridrow="3" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFF434333FE83FDD9" bold="true" box="[194,327,547,568]" italics="true" pageId="11" pageNumber="22">P. k i c h w a r u m</emphasis>
</th>
<td id="746B8C47FFE6D4C9FE084333FDE1FD8F" box="[393,549,547,622]" gridcol="1" gridrow="3" pageId="11" pageNumber="22">present as thin ridge, usually black</td>
<td id="746B8C47FFE6D4C9FDED4333FD21FD8F" box="[620,741,547,622]" gridcol="2" gridrow="3" pageId="11" pageNumber="22">Ψ 19.3-30.2; ɗ 15.2-22.4</td>
<td id="746B8C47FFE6D4C9FCAE4333FC46FD8F" box="[815,898,547,622]" gridcol="3" gridrow="3" pageId="11" pageNumber="22">+</td>
<td id="746B8C47FFE6D4C9FC2B4333FBC3FD8F" box="[938,1031,547,622]" gridcol="4" gridrow="3" pageId="11" pageNumber="22">+</td>
<td id="746B8C47FFE6D4C9FBB04333FB69FD8F" box="[1073,1197,547,622]" gridcol="5" gridrow="3" pageId="11" pageNumber="22">visible</td>
<td id="746B8C47FFE6D4C9FB564333FA41FD8F" box="[1239,1413,547,622]" gridcol="6" gridrow="3" pageId="11" pageNumber="22">uniform brown</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF43439DFA41FD5C" box="[194,1413,653,701]" gridrow="4" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF43439DFE8CFD5C" box="[194,328,653,701]" gridcol="0" gridrow="4" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF42439EFE8CFD42" box="[195,328,654,675]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="BBFE3B6BFFE62B3DFF42439EFE8CFD42" box="[195,328,654,675]" italics="true" pageId="11" pageNumber="22">P. acuminatus</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE08439DFDE1FD5C" box="[393,549,653,701]" gridcol="1" gridrow="4" pageId="11" pageNumber="22">absent</td>
<td id="746B8C47FFE6D4C9FDED439DFD21FD5C" box="[620,741,653,701]" gridcol="2" gridrow="4" pageId="11" pageNumber="22">Ψ 26-34; ɗ 17-23[9]</td>
<td id="746B8C47FFE6D4C9FCAE439DFC46FD5C" box="[815,898,653,701]" gridcol="3" gridrow="4" pageId="11" pageNumber="22">+[3]</td>
<td id="746B8C47FFE6D4C9FC2B439DFBC3FD5C" box="[938,1031,653,701]" gridcol="4" gridrow="4" pageId="11" pageNumber="22">-</td>
<td id="746B8C47FFE6D4C9FBB0439DFB69FD5C" box="[1073,1197,653,701]" gridcol="5" gridrow="4" pageId="11" pageNumber="22">concealed[3]</td>
<td id="746B8C47FFE6D4C9FB56439DFA41FD5C" box="[1239,1413,653,701]" gridcol="6" gridrow="4" pageId="11" pageNumber="22">pale green</td>
</tr>
</table>
</paragraph>
<paragraph id="8935E779FFE62B3DFF4D43CEFA9FF91E" pageId="11" pageNumber="22">
<table id="FB8A15D9FFE6D4C9FF2043CDFA4BF8FB" box="[161,1423,733,1818]" gridcols="7" gridrows="12" pageId="11" pageNumber="22">
<tr id="37BAE53BFFE6D4C9FF2043CDFA4BFCC9" box="[161,1423,733,808]" gridrow="0" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF2043CDFEAEFCC9" box="[161,362,733,808]" gridcol="0" gridrow="0" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF4D43CEFE84FD12" box="[204,320,734,755]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="BBFE3B6BFFE62B3DFF4D43CEFE84FD12" box="[204,320,734,755]" italics="true" pageId="11" pageNumber="22">P. carvalhoi</emphasis>
</taxonomicName>
</th>
<th id="746B8C47FFE6D4C9FE0043CDFDE9FCC9" box="[385,557,733,808]" gridcol="1" gridrow="0" pageId="11" pageNumber="22">as a marking, present or absent[9]</th>
<th id="746B8C47FFE6D4C9FDDE43CDFD37FCC9" box="[607,755,733,808]" gridcol="2" gridrow="0" pageId="11" pageNumber="22">Ψ 17.4-21.9; ɗ 12.8-18.2[2]</th>
<th id="746B8C47FFE6D4C9FCA943CDFC4EFCC9" box="[808,906,733,808]" gridcol="3" gridrow="0" pageId="11" pageNumber="22">-[9]</th>
<th id="746B8C47FFE6D4C9FC2443CDFBCFFCC9" box="[933,1035,733,808]" gridcol="4" gridrow="0" pageId="11" pageNumber="22">+[9 photo]</th>
<th id="746B8C47FFE6D4C9FBA543CDFB73FCC9" box="[1060,1207,733,808]" gridcol="5" gridrow="0" pageId="11" pageNumber="22">concealed[1] or barely evident [9]</th>
<th id="746B8C47FFE6D4C9FB4C43CDFA4BFCC9" box="[1229,1423,733,808]" gridcol="6" gridrow="0" pageId="11" pageNumber="22">?</th>
</tr>
<tr id="37BAE53BFFE6D4C9FF204258FA4BFC99" box="[161,1423,840,888]" gridrow="1" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF204258FEAEFC99" box="[161,362,840,888]" gridcol="0" gridrow="1" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF324258FE9DFCBC" box="[179,345,840,861]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="croceoinguinis">
<emphasis id="BBFE3B6BFFE62B3DFF324258FE9DFCBC" box="[179,345,840,861]" italics="true" pageId="11" pageNumber="22">P. croceoinguinis</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE004258FDE9FC99" box="[385,557,840,888]" gridcol="1" gridrow="1" pageId="11" pageNumber="22">present as low thick ridge</td>
<td id="746B8C47FFE6D4C9FDDE4258FD37FC99" box="[607,755,840,888]" gridcol="2" gridrow="1" pageId="11" pageNumber="22">Ψ 24.9; ɗ 17.2-21.6[8]</td>
<td id="746B8C47FFE6D4C9FCA94258FC4EFC99" box="[808,906,840,888]" gridcol="3" gridrow="1" pageId="11" pageNumber="22">-</td>
<td id="746B8C47FFE6D4C9FC244258FBCFFC99" box="[933,1035,840,888]" gridcol="4" gridrow="1" pageId="11" pageNumber="22">+</td>
<td id="746B8C47FFE6D4C9FBA54258FB73FC99" box="[1060,1207,840,888]" gridcol="5" gridrow="1" pageId="11" pageNumber="22">concealed[6]</td>
<td id="746B8C47FFE6D4C9FB4C4258FA4BFC99" box="[1229,1423,840,888]" gridcol="6" gridrow="1" pageId="11" pageNumber="22">grey</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF204287FA4BFC26" box="[161,1423,919,967]" gridrow="2" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF204287FEAEFC26" box="[161,362,919,967]" gridcol="0" gridrow="2" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFF5E4288FEEEFC4C" box="[223,298,920,941]" italics="true" pageId="11" pageNumber="22">P. f r a te r</emphasis>
</th>
<td id="746B8C47FFE6D4C9FE004287FDE9FC26" box="[385,557,919,967]" gridcol="1" gridrow="2" pageId="11" pageNumber="22">?</td>
<td id="746B8C47FFE6D4C9FDDE4287FD37FC26" box="[607,755,919,967]" gridcol="2" gridrow="2" pageId="11" pageNumber="22">Ψ 23.7-25.9; ɗ 15.6-18.8;[1]</td>
<td id="746B8C47FFE6D4C9FCA94287FC4EFC26" box="[808,906,919,967]" gridcol="3" gridrow="2" pageId="11" pageNumber="22">?</td>
<td id="746B8C47FFE6D4C9FC244287FBCFFC26" box="[933,1035,919,967]" gridcol="4" gridrow="2" pageId="11" pageNumber="22">?</td>
<td id="746B8C47FFE6D4C9FBA54287FB73FC26" box="[1060,1207,919,967]" gridcol="5" gridrow="2" pageId="11" pageNumber="22">visible[8]</td>
<td id="746B8C47FFE6D4C9FB4C4287FA4BFC26" box="[1229,1423,919,967]" gridcol="6" gridrow="2" pageId="11" pageNumber="22">uniform brown[8]</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF2042F7FA4BFBF6" box="[161,1423,999,1047]" gridrow="3" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF2042F7FEAEFBF6" box="[161,362,999,1047]" gridcol="0" gridrow="3" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFF4842F8FEFBFC1C" box="[201,319,1000,1021]" italics="true" pageId="11" pageNumber="22">P. in c o m p t u s</emphasis>
</th>
<td id="746B8C47FFE6D4C9FE0042F7FDE9FBF6" box="[385,557,999,1047]" gridcol="1" gridrow="3" pageId="11" pageNumber="22">present as mark- ing [1]</td>
<td id="746B8C47FFE6D4C9FDDE42F7FD37FBF6" box="[607,755,999,1047]" gridcol="2" gridrow="3" pageId="11" pageNumber="22">Ψ 27.5-45.4; ɗ 21.7-27.9[1]</td>
<td id="746B8C47FFE6D4C9FCA942F7FC4EFBF6" box="[808,906,999,1047]" gridcol="3" gridrow="3" pageId="11" pageNumber="22">-[8]</td>
<td id="746B8C47FFE6D4C9FC2442F7FBCFFBF6" box="[933,1035,999,1047]" gridcol="4" gridrow="3" pageId="11" pageNumber="22">-[1]</td>
<td id="746B8C47FFE6D4C9FBA542F7FB73FBF6" box="[1060,1207,999,1047]" gridcol="5" gridrow="3" pageId="11" pageNumber="22">visible[1]</td>
<td id="746B8C47FFE6D4C9FB4C42F7FA4BFBF6" box="[1229,1423,999,1047]" gridcol="6" gridrow="3" pageId="11" pageNumber="22">brown[1]</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF204527FA4BFB7D" box="[161,1423,1079,1180]" gridrow="4" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF204527FEAEFB7D" box="[161,362,1079,1180]" gridcol="0" gridrow="4" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF454527FE83FBAD" box="[196,327,1079,1100]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="lanthanites">
<emphasis id="BBFE3B6BFFE62B3DFF454527FE83FBAD" box="[196,327,1079,1100]" italics="true" pageId="11" pageNumber="22">P. lanthanites</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE004527FDE9FB7D" box="[385,557,1079,1180]" gridcol="1" gridrow="4" pageId="11" pageNumber="22">absent, but with chevron mark- ings and scapular points</td>
<td id="746B8C47FFE6D4C9FDDE4527FD37FB7D" box="[607,755,1079,1180]" gridcol="2" gridrow="4" pageId="11" pageNumber="22">Ψ 18-23; ɗ 13-17[4]</td>
<td id="746B8C47FFE6D4C9FCA94527FC4EFB7D" box="[808,906,1079,1180]" gridcol="3" gridrow="4" pageId="11" pageNumber="22">usually[3]</td>
<td id="746B8C47FFE6D4C9FC244527FBCFFB7D" box="[933,1035,1079,1180]" gridcol="4" gridrow="4" pageId="11" pageNumber="22">+</td>
<td id="746B8C47FFE6D4C9FBA54527FB73FB7D" box="[1060,1207,1079,1180]" gridcol="5" gridrow="4" pageId="11" pageNumber="22">visible</td>
<td id="746B8C47FFE6D4C9FB4C4527FA4BFB7D" box="[1229,1423,1079,1180]" gridcol="6" gridrow="4" pageId="11" pageNumber="22">brown with small yellow-tan flecks[3]</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF2045ACFA4BFB0D" box="[161,1423,1212,1260]" gridrow="5" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF2045ACFEAEFB0D" box="[161,362,1212,1260]" gridcol="0" gridrow="5" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF5145ACFEFFFB30" box="[208,315,1212,1233]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="librarius">
<emphasis id="BBFE3B6BFFE62B3DFF5145ACFEFFFB30" box="[208,315,1212,1233]" italics="true" pageId="11" pageNumber="22">P. librarius</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE0045ACFDE9FB0D" box="[385,557,1212,1260]" gridcol="1" gridrow="5" pageId="11" pageNumber="22">no, but some pale chevron markings</td>
<td id="746B8C47FFE6D4C9FDDE45ACFD37FB0D" box="[607,755,1212,1260]" gridcol="2" gridrow="5" pageId="11" pageNumber="22">Ψ 16-19; ɗ 13-17[4]</td>
<td id="746B8C47FFE6D4C9FCA945ACFC4EFB0D" box="[808,906,1212,1260]" gridcol="3" gridrow="5" pageId="11" pageNumber="22">-</td>
<td id="746B8C47FFE6D4C9FC2445ACFBCFFB0D" box="[933,1035,1212,1260]" gridcol="4" gridrow="5" pageId="11" pageNumber="22">two low tubercles</td>
<td id="746B8C47FFE6D4C9FBA545ACFB73FB0D" box="[1060,1207,1212,1260]" gridcol="5" gridrow="5" pageId="11" pageNumber="22">partially con- cealed[8]</td>
<td id="746B8C47FFE6D4C9FB4C45ACFA4BFB0D" box="[1229,1423,1212,1260]" gridcol="6" gridrow="5" pageId="11" pageNumber="22">reddish in life</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF20441BFA4BFAB7" box="[161,1423,1291,1366]" gridrow="6" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF20441BFEAEFAB7" box="[161,362,1291,1366]" gridcol="0" gridrow="6" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFF55441CFEF0FAC0" box="[212,308,1292,1313]" italics="true" pageId="11" pageNumber="22">P. ma rtia e</emphasis>
</th>
<td id="746B8C47FFE6D4C9FE00441BFDE9FAB7" box="[385,557,1291,1366]" gridcol="1" gridrow="6" pageId="11" pageNumber="22">no, but other markings present[4]</td>
<td id="746B8C47FFE6D4C9FDDE441BFD37FAB7" box="[607,755,1291,1366]" gridcol="2" gridrow="6" pageId="11" pageNumber="22">ɗ 15.3-18.7[7]</td>
<td id="746B8C47FFE6D4C9FCA9441BFC4EFAB7" box="[808,906,1291,1366]" gridcol="3" gridrow="6" pageId="11" pageNumber="22">-[4]</td>
<td id="746B8C47FFE6D4C9FC24441BFBCFFAB7" box="[933,1035,1291,1366]" gridcol="4" gridrow="6" pageId="11" pageNumber="22">low flat warts[4]</td>
<td id="746B8C47FFE6D4C9FBA5441BFB73FAB7" box="[1060,1207,1291,1366]" gridcol="5" gridrow="6" pageId="11" pageNumber="22">concealed[1]</td>
<td id="746B8C47FFE6D4C9FB4C441BFA4BFAB7" box="[1229,1423,1291,1366]" gridcol="6" gridrow="6" pageId="11" pageNumber="22">brown</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF204465FA4BFA21" box="[161,1423,1397,1472]" gridrow="7" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF204465FEAEFA21" box="[161,362,1397,1472]" gridcol="0" gridrow="7" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF504466FEFEFA6A" box="[209,314,1398,1419]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="paululus">
<emphasis id="BBFE3B6BFFE62B3DFF504466FEFEFA6A" box="[209,314,1398,1419]" italics="true" pageId="11" pageNumber="22">P. paululus</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE004465FDE9FA21" box="[385,557,1397,1472]" gridcol="1" gridrow="7" pageId="11" pageNumber="22">some scapular markings, but not a ridge</td>
<td id="746B8C47FFE6D4C9FDDE4465FD37FA21" box="[607,755,1397,1472]" gridcol="2" gridrow="7" pageId="11" pageNumber="22">Ψ 23; ɗ 19[3]</td>
<td id="746B8C47FFE6D4C9FCA94465FC4EFA21" box="[808,906,1397,1472]" gridcol="3" gridrow="7" pageId="11" pageNumber="22">+[3]</td>
<td id="746B8C47FFE6D4C9FC244465FBCFFA21" box="[933,1035,1397,1472]" gridcol="4" gridrow="7" pageId="11" pageNumber="22">-</td>
<td id="746B8C47FFE6D4C9FBA54465FB73FA21" box="[1060,1207,1397,1472]" gridcol="5" gridrow="7" pageId="11" pageNumber="22">visible[3]</td>
<td id="746B8C47FFE6D4C9FB4C4465FA4BFA21" box="[1229,1423,1397,1472]" gridcol="6" gridrow="7" pageId="11" pageNumber="22">greyish-tan[3]</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF2044F0FA4BF9CB" box="[161,1423,1504,1578]" gridrow="8" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF2044F0FEAEF9CB" box="[161,362,1504,1578]" gridcol="0" gridrow="8" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF6044F0FEEEFA14" box="[225,298,1504,1525]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="pecki">
<emphasis id="BBFE3B6BFFE62B3DFF6044F0FEEEFA14" box="[225,298,1504,1525]" italics="true" pageId="11" pageNumber="22">P. pecki</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE0044F0FDE9F9CB" box="[385,557,1504,1578]" gridcol="1" gridrow="8" pageId="11" pageNumber="22">absent; brown interorbital trian- gle[7]</td>
<td id="746B8C47FFE6D4C9FDDE44F0FD37F9CB" box="[607,755,1504,1578]" gridcol="2" gridrow="8" pageId="11" pageNumber="22">Ψ 24.6-31.3; ɗ 19.6-22.5[8]</td>
<td id="746B8C47FFE6D4C9FCA944F0FC4EF9CB" box="[808,906,1504,1578]" gridcol="3" gridrow="8" pageId="11" pageNumber="22">-[7]</td>
<td id="746B8C47FFE6D4C9FC2444F0FBCFF9CB" box="[933,1035,1504,1578]" gridcol="4" gridrow="8" pageId="11" pageNumber="22">+ nonpun- gent[7]</td>
<td id="746B8C47FFE6D4C9FBA544F0FB73F9CB" box="[1060,1207,1504,1578]" gridcol="5" gridrow="8" pageId="11" pageNumber="22">prominent [7]</td>
<td id="746B8C47FFE6D4C9FB4C44F0FA4BF9CB" box="[1229,1423,1504,1578]" gridcol="6" gridrow="8" pageId="11" pageNumber="22">uniform brown[7]</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF20475AFA4BF99B" box="[161,1423,1610,1658]" gridrow="9" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF20475AFEAEF99B" box="[161,362,1610,1658]" gridcol="0" gridrow="9" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF20475BFEAEF981" box="[161,362,1611,1632]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="pseudoacuminatus">
<emphasis id="BBFE3B6BFFE62B3DFF20475BFEAEF981" box="[161,362,1611,1632]" italics="true" pageId="11" pageNumber="22">P. pseudoacuminatus</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE00475AFDE9F99B" box="[385,557,1610,1658]" gridcol="1" gridrow="9" pageId="11" pageNumber="22">absent</td>
<td id="746B8C47FFE6D4C9FDDE475AFD37F99B" box="[607,755,1610,1658]" gridcol="2" gridrow="9" pageId="11" pageNumber="22">Ψ 21.9-26.8; ɗ 15.7-22.0[6]</td>
<td id="746B8C47FFE6D4C9FCA9475AFC4EF99B" box="[808,906,1610,1658]" gridcol="3" gridrow="9" pageId="11" pageNumber="22">-, + in some[3]</td>
<td id="746B8C47FFE6D4C9FC24475AFBCFF99B" box="[933,1035,1610,1658]" gridcol="4" gridrow="9" pageId="11" pageNumber="22">-[3]</td>
<td id="746B8C47FFE6D4C9FBA5475AFB73F99B" box="[1060,1207,1610,1658]" gridcol="5" gridrow="9" pageId="11" pageNumber="22">visible[3]</td>
<td id="746B8C47FFE6D4C9FB4C475AFA4BF99B" box="[1229,1423,1610,1658]" gridcol="6" gridrow="9" pageId="11" pageNumber="22">uniform pale green to grey-green[3]</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF20478AFA4BF92B" box="[161,1423,1690,1738]" gridrow="10" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF20478AFEAEF92B" box="[161,362,1690,1738]" gridcol="0" gridrow="10" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF36478BFE90F951" box="[183,340,1691,1712]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="quaquaversus">
<emphasis id="BBFE3B6BFFE62B3DFF36478BFE90F951" box="[183,340,1691,1712]" italics="true" pageId="11" pageNumber="22">P. quaquaversus</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE00478AFDE9F92B" box="[385,557,1690,1738]" gridcol="1" gridrow="10" pageId="11" pageNumber="22">absent</td>
<td id="746B8C47FFE6D4C9FDDE478AFD37F92B" box="[607,755,1690,1738]" gridcol="2" gridrow="10" pageId="11" pageNumber="22">Ψ 24.6-31.3; ɗ 19.6–22.5[4]</td>
<td id="746B8C47FFE6D4C9FCA9478AFC4EF92B" box="[808,906,1690,1738]" gridcol="3" gridrow="10" pageId="11" pageNumber="22">-</td>
<td id="746B8C47FFE6D4C9FC24478AFBCFF92B" box="[933,1035,1690,1738]" gridcol="4" gridrow="10" pageId="11" pageNumber="22">+</td>
<td id="746B8C47FFE6D4C9FBA5478AFB73F92B" box="[1060,1207,1690,1738]" gridcol="5" gridrow="10" pageId="11" pageNumber="22">concealed[3]</td>
<td id="746B8C47FFE6D4C9FB4C478AFA4BF92B" box="[1229,1423,1690,1738]" gridcol="6" gridrow="10" pageId="11" pageNumber="22">purplish with white flecks[3]</td>
</tr>
<tr id="37BAE53BFFE6D4C9FF2047FAFA4BF8FB" box="[161,1423,1770,1818]" gridrow="11" pageId="11" pageNumber="22">
<th id="746B8C47FFE6D4C9FF2047FAFEAEF8FB" box="[161,362,1770,1818]" gridcol="0" gridrow="11" pageId="11" pageNumber="22">
<taxonomicName id="4E8A9CFAFFE62B3DFF4D47FAFE84F91E" box="[204,320,1770,1791]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="22" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="BBFE3B6BFFE62B3DFF4D47FAFE84F91E" box="[204,320,1770,1791]" italics="true" pageId="11" pageNumber="22">P. variabilis</emphasis>
</taxonomicName>
</th>
<td id="746B8C47FFE6D4C9FE0047FAFDE9F8FB" box="[385,557,1770,1818]" gridcol="1" gridrow="11" pageId="11" pageNumber="22">absent</td>
<td id="746B8C47FFE6D4C9FDDE47FAFD37F8FB" box="[607,755,1770,1818]" gridcol="2" gridrow="11" pageId="11" pageNumber="22">Ψ 22.0–27.0; ɗ 19[3]</td>
<td id="746B8C47FFE6D4C9FCA947FAFC4EF8FB" box="[808,906,1770,1818]" gridcol="3" gridrow="11" pageId="11" pageNumber="22">+[6]</td>
<td id="746B8C47FFE6D4C9FC2447FAFBCFF8FB" box="[933,1035,1770,1818]" gridcol="4" gridrow="11" pageId="11" pageNumber="22">?</td>
<td id="746B8C47FFE6D4C9FBA547FAFB73F8FB" box="[1060,1207,1770,1818]" gridcol="5" gridrow="11" pageId="11" pageNumber="22">visible</td>
<td id="746B8C47FFE6D4C9FB4C47FAFA4BF8FB" box="[1229,1423,1770,1818]" gridcol="6" gridrow="11" pageId="11" pageNumber="22">brown[6]</td>
</tr>
</table>
</paragraph>
<paragraph id="8935E779FFE62B3AFBF1464AFA8FF869" blockId="11.[1136,1436,1882,1903]" lastBlockId="12.[151,1436,1878,1928]" lastPageId="12" lastPageNumber="23" pageId="11" pageNumber="22">
<emphasis id="BBFE3B6BFFE62B3DFBF1464AFA58F88E" box="[1136,1436,1882,1903]" italics="true" pageId="11" pageNumber="22">...... continued on the next page</emphasis>
[1] 
<bibRefCitation id="ED1B9A88FFE12B3AFF384646FE7EF88A" author="Lynch" box="[185,442,1878,1899]" pageId="12" pageNumber="32" refString="Lynch, J. D. &amp; Duellman, W. E. (1980) The Eleutherodactylus of the Amazonian slopes of the Ecuadorian Andes (Anura: Leptodactylidae). Miscellaneous Publications of the Natural History Museum, University of Kansas, 69, 1 - 86." type="journal article" year="1980">Lynch and Duellman 1980</bibRefCitation>
; [2] Lynch 1980; [3] 
<bibRefCitation id="ED1B9A88FFE12B3AFD044646FCD8F88A" author="Duellman" box="[645,796,1878,1899]" pageId="12" pageNumber="31" refString="Duellman, W. E. (1978) The Biology of an Equatorial Herpetofauna in Amazonian Ecuador. University of Kansas Lawrence, Kansas, USA, 1 - 352." type="book chapter" year="1978">Duellman 1978</bibRefCitation>
; [4] 
<bibRefCitation id="ED1B9A88FFE12B3AFCC94646FC79F88A" author="Lynch" box="[840,957,1878,1899]" pageId="12" pageNumber="32" refString="Lynch, J. D. (1974) New species of frogs (Leptodactylidae: Eleutherodactylus) from the Amazonian lowlands of Ecuador. Occasional Papers of the Museum of Natural History Kansas, 31, 1 - 22." type="journal article" year="1974">Lynch 1974</bibRefCitation>
; [5] IUCN et al.; [6] 
<bibRefCitation id="ED1B9A88FFE12B3AFB044646FB3EF88A" author="Lynch" box="[1157,1274,1878,1899]" pageId="12" pageNumber="32" refString="Lynch, J. D. (1968) Systematic status of some Andean leptodactylid frogs with a description of a new species of Eleutherodactylus. Herpetologica, 24, 289 - 300." type="journal article" year="1968">Lynch 1968</bibRefCitation>
; [7] 
<bibRefCitation id="ED1B9A88FFE12B3AFAA44646FEC8F869" author="Duellman" pageId="12" pageNumber="31" refString="Duellman, W. E. &amp; Lynch, J. D. (1988). Anuran amphibians from the cordillera de Cutucu. Proceedings of the Academy of Natural Sciences of Philadelphia, 140, 125 - 142." type="journal article" year="1988">Duellman &amp; Lynch 1988</bibRefCitation>
; [8] 
<bibRefCitation id="ED1B9A88FFE12B3AFEB64663FE3AF869" author="Flores" box="[311,510,1907,1928]" pageId="12" pageNumber="31" refString="Flores, G. &amp; Vigle, G. O. (1994) A new species of Eleutherodactylus (Anura: Leptodactylidae) from the lowland rainforests of Amazonian Ecuador, with notes on the Eleutherodactylus frater assembly. Journal of Herpetology, 28, 416 - 424." type="journal article" year="1994">
<collectingCountry id="F19DA7E9FFE12B3AFEB64663FEB7F869" box="[311,371,1907,1928]" name="Indonesia" pageId="12" pageNumber="23">Flores</collectingCountry>
&amp; Vigle 1994
</bibRefCitation>
; [9] 
<bibRefCitation id="ED1B9A88FFE12B3AFDAB4663FC80F869" author="Rodriguez" box="[554,836,1907,1928]" pageId="12" pageNumber="32" refString="Rodriguez, L. O. &amp; Duellman, W. D. (1994) Guide to Frogs of the Iquitos Region, Amazonian Peru. University of Kansas Museum of Natural History Special Publication No. 22 Kansas, USA, 1 - 80 pp." type="book chapter" year="1994">Rodriguez &amp; Duellman 1994</bibRefCitation>
; [no citation] KRE/DCC review of QCAZ specimens.
</paragraph>
</subSubSection>
</treatment>
</document>