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<mods:title id="400F0C186D3715F859832FEE0C5C1FE2">Observations on non-didemnid ascidians from Australian waters (1)</mods:title>
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<paragraph id="890B36D7FFDDCD6C1FBFFF2FE1D2FF0C" blockId="46.[125,1172,136,288]" box="[444,928,136,160]" pageId="46" pageNumber="214">
<taxonomicName id="4EB44D54FFDDCD6C1FBFFF2FE1D2FF0C" ID-CoL="WGSV" authority="(Heller, 1878)" baseAuthorityName="Heller" baseAuthorityYear="1878" box="[444,928,136,160]" class="Ascidiacea" family="Styelidae" genus="Cnemidocarpa" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="areolata">
<emphasis id="BBC0EAC5FFDDCD6C1FBFFF2FE082FF0C" bold="true" box="[444,752,136,160]" italics="true" pageId="46" pageNumber="214">Cnemidocarpa areolata</emphasis>
(
<bibRefCitation id="ED254B26FFDDCD6C1D02FF2FE1EAFF0C" author="Heller C" box="[769,920,136,160]" pageId="46" pageNumber="214" pagination="2 - 28" refId="ref27776" refString="Heller C. 1878. Beitrage zur nahern Kenntnis der Tunicaten. Sitzungsberichte der Mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften Wien 77: 2 - 28." type="journal article" year="1878">Heller, 1878</bibRefCitation>
)
</taxonomicName>
</paragraph>
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<subSubSection id="C1AE655CFFDDCD6C1E7EFF0FE01EFE8C" pageId="46" pageNumber="214" type="reference_group">
<paragraph id="890B36D7FFDDCD6C1E7EFF0FE38CFF6C" blockId="46.[125,1172,136,288]" box="[125,510,168,192]" pageId="46" pageNumber="214">
<treatmentCitationGroup id="A9A411F9FFDDCD6C1E7EFF0FE38CFF6C" box="[125,510,168,192]" pageId="46" pageNumber="214">
<treatmentCitation id="081510C6FFDDCD6C1E7EFF0FE388FF6C" author="Heller C" box="[125,506,168,192]" page="26" pageId="46" pageNumber="214" year="1878">
<taxonomicName id="4EB44D54FFDDCD6C1E7EFF0FE388FF6C" ID-CoL="5386B" authority="Heller 1878, p 26" authorityName="Heller" authorityPageNumber="26" authorityYear="1878" box="[125,506,168,192]" class="Ascidiacea" family="Styelidae" genus="Styela" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="areolata">
<emphasis id="BBC0EAC5FFDDCD6C1E7EFF0FE352FF6C" box="[125,288,168,192]" italics="true" pageId="46" pageNumber="214">Styela areolata</emphasis>
<bibRefCitation id="ED254B26FFDDCD6C1F2AFF0FE3CAFF6C" author="Heller C" box="[297,440,168,192]" pageId="46" pageNumber="214" pagination="2 - 28" refId="ref27776" refString="Heller C. 1878. Beitrage zur nahern Kenntnis der Tunicaten. Sitzungsberichte der Mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften Wien 77: 2 - 28." type="journal article" year="1878">Heller 1878</bibRefCitation>
, p 26
</taxonomicName>
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.
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</paragraph>
<paragraph id="890B36D7FFDDCD6C1E7EFF6FE2ADFEAC" blockId="46.[125,1172,136,288]" pageId="46" pageNumber="214">
<treatmentCitationGroup id="A9A411F9FFDDCD6C1E7EFF6FE2ADFEAC" pageId="46" pageNumber="214">
<treatmentCitation id="081510C6FFDDCD6C1E7EFF6FE016FF4C" author="Kott P" box="[125,612,200,224]" page="122" pageId="46" pageNumber="214" year="1985">
<taxonomicName id="4EB44D54FFDDCD6C1E7EFF6FE016FF4C" ID-CoL="WGSV" authority=": Kott 1985, p 122" authorityName="Kott" authorityPageNumber="122" authorityYear="1985" box="[125,612,200,224]" class="Ascidiacea" family="Styelidae" genus="Cnemidocarpa" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="areolata">
<emphasis id="BBC0EAC5FFDDCD6C1E7EFF6FE30CFF4C" box="[125,382,200,224]" italics="true" pageId="46" pageNumber="214">Cnemidocarpa areolata</emphasis>
:
<bibRefCitation id="ED254B26FFDDCD6C1F90FF6FE061FF4C" author="Kott P" box="[403,531,200,224]" pageId="46" pageNumber="214" pagination="1 - 440" refId="ref28254" refString="Kott P. 1985. The Australian Ascidiacea Pt 1, Phlebobranchia and Stolidobranchia. Memoirs of the Queensland Museum 23: 1 - 440." type="journal article" year="1985">Kott 1985</bibRefCitation>
, p 122
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and synonymy;
<treatmentCitation id="081510C6FFDDCD6C1D30FF6FE2A9FEAC" author="Monniot C &amp; Monniot F" page="577" pageId="46" pageNumber="214" year="1984">
<bibRefCitation id="ED254B26FFDDCD6C1D30FF6FE6E2FF4C" author="Monniot C &amp; Monniot F" box="[819,1168,200,224]" pageId="46" pageNumber="214" pagination="567 - 582" refId="ref29238" refString="Monniot C, Monniot F. 1984. Ascidies littorales de Guadeloupe VII. Especes nouvelles et complementaires a l'inventaire. Bulletin du Museum National d'Histoire Naturelle Serie 4 6 A: 567 - 582." type="journal article" year="1984">Monniot and Monniot 1984</bibRefCitation>
, p 577
</treatmentCitation>
.
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</paragraph>
<paragraph id="890B36D7FFDDCD6C1E7EFEAFE01EFE8C" blockId="46.[125,1172,136,288]" box="[125,620,264,288]" pageId="46" pageNumber="214">
<treatmentCitationGroup id="A9A411F9FFDDCD6C1E7EFEAFE01EFE8C" box="[125,620,264,288]" pageId="46" pageNumber="214">
<treatmentCitation id="081510C6FFDDCD6C1E7EFEAFE01AFE8C" author="Nishikawa T" box="[125,616,264,288]" page="96" pageId="46" pageNumber="214" year="1991">
<taxonomicName id="4EB44D54FFDDCD6C1E7EFEAFE01AFE8C" ID-CoL="5ZGMK" authority=": Nishikawa 1991, p 96" authorityName="Nishikawa" authorityPageNumber="96" authorityYear="1991" box="[125,616,264,288]" class="Ascidiacea" family="Styelidae" genus="Cnemidocarpa" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="irene">
<emphasis id="BBC0EAC5FFDDCD6C1E7EFEAFE324FE8C" box="[125,342,264,288]" italics="true" pageId="46" pageNumber="214">Cnemidocarpa irene</emphasis>
:
<bibRefCitation id="ED254B26FFDDCD6C1F65FEAFE054FE8C" author="Nishikawa T" box="[358,550,264,288]" pageId="46" pageNumber="214" pagination="25 - 170" refId="ref29462" refString="Nishikawa T. 1991. The ascidians of the Japan Sea. II. Publications of the Seto Marine Biology Laboratory 35: 25 - 170." type="journal article" year="1991">Nishikawa 1991</bibRefCitation>
, p 96
</taxonomicName>
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.
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</paragraph>
</subSubSection>
<subSubSection id="C1AE655CFFDDCD6C1E7EFEF7E172FDFB" pageId="46" pageNumber="214" type="distribution">
<paragraph id="890B36D7FFDDCD6C1E7EFEF7E28FFEC4" blockId="46.[125,253,336,360]" box="[125,253,336,360]" pageId="46" pageNumber="214">
<emphasis id="BBC0EAC5FFDDCD6C1E7EFEF7E28FFEC4" box="[125,253,336,360]" italics="true" pageId="46" pageNumber="214">Distribution</emphasis>
</paragraph>
<paragraph id="890B36D7FFDDCD6C1E7EFE27E172FDFB" blockId="46.[125,1173,384,599]" pageId="46" pageNumber="214">
Previously recorded (see
<bibRefCitation id="ED254B26FFDDCD6C1FB6FE27E165FE34" author="Monniot C &amp; Monniot F" box="[437,791,384,408]" pageId="46" pageNumber="214" pagination="567 - 582" refId="ref29238" refString="Monniot C, Monniot F. 1984. Ascidies littorales de Guadeloupe VII. Especes nouvelles et complementaires a l'inventaire. Bulletin du Museum National d'Histoire Naturelle Serie 4 6 A: 567 - 582." type="journal article" year="1984">Monniot and Monniot 1984</bibRefCitation>
;
<bibRefCitation id="ED254B26FFDDCD6C1D29FE27E1DFFE34" author="Kott P" box="[810,941,384,408]" pageId="46" pageNumber="214" pagination="1 - 440" refId="ref28254" refString="Kott P. 1985. The Australian Ascidiacea Pt 1, Phlebobranchia and Stolidobranchia. Memoirs of the Queensland Museum 23: 1 - 440." type="journal article" year="1985">Kott 1985</bibRefCitation>
;
<bibRefCitation id="ED254B26FFDDCD6C1DC3FE27E6F5FE34" author="Nishikawa T" box="[960,1159,384,408]" pageId="46" pageNumber="214" pagination="25 - 170" refId="ref29462" refString="Nishikawa T. 1991. The ascidians of the Japan Sea. II. Publications of the Seto Marine Biology Laboratory 35: 25 - 170." type="journal article" year="1991">Nishikawa 1991</bibRefCitation>
): Western Australia (Dampier Archipelago, Cape Jaubert, NW Cape, Port Hedland, Carnarvon, Shark Bay, Cockburn Sound, Albany); South Australia (Spencer Gulf); Victoria (Bass Strait); New South Wales (Port Hacking, Long Reef); Queensland (Moreton Bay, Capricorn Group, Low Isles); Northern Territory (Darwin);
<collectingCountry id="F1A37647FFDDCD6C1D9BFDA7E1ABFDB4" box="[920,985,512,536]" name="Palau" pageId="46" pageNumber="214">Palau</collectingCountry>
Is,
<collectingCountry id="F1A37647FFDDCD6C1A08FDA7E6FDFDB4" box="[1035,1167,512,536]" name="Philippines" pageId="46" pageNumber="214">Philippines</collectingCountry>
, Noumea,
<collectingCountry id="F1A37647FFDDCD6C1EF7FD87E3DDFD94" box="[244,431,544,568]" name="New Caledonia" pageId="46" pageNumber="214">New Caledonia</collectingCountry>
, Marianas Is,
<collectingCountry id="F1A37647FFDDCD6C1C58FD87E0EDFD94" box="[603,671,544,568]" name="Japan" pageId="46" pageNumber="214">Japan</collectingCountry>
,
<collectingCountry id="F1A37647FFDDCD6C1CACFD87E0ABFD94" box="[687,729,544,568]" name="Fiji" pageId="46" pageNumber="214">Fiji</collectingCountry>
, South
<collectingCountry id="F1A37647FFDDCD6C1D3BFD87E10DFD94" box="[824,895,544,568]" name="China" pageId="46" pageNumber="214">China</collectingCountry>
Sea,
<collectingCountry id="F1A37647FFDDCD6C1DC6FD87E67BFD94" box="[965,1033,544,568]" name="Japan" pageId="46" pageNumber="214">Japan</collectingCountry>
,
<collectingCountry id="F1A37647FFDDCD6C1A1BFD87E6E3FD94" box="[1048,1169,544,568]" name="Sri Lanka" pageId="46" pageNumber="214">Sri Lanka</collectingCountry>
. New record: Queensland (18.845
<emphasis id="BBC0EAC5FFDDCD6C1C05FD98E062FDFB" bold="true" box="[518,528,575,599]" italics="true" pageId="46" pageNumber="214">°</emphasis>
S, 146.795
<emphasis id="BBC0EAC5FFDDCD6C1C92FD98E0E9FDFB" bold="true" box="[657,667,575,599]" italics="true" pageId="46" pageNumber="214">°</emphasis>
E,
<quantity id="4E4C9B32FFDDCD6C1CBDFD98E08BFDFB" box="[702,761,575,599]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="46" pageNumber="214" unit="m" value="26.0">26 m</quantity>
)
</paragraph>
</subSubSection>
<subSubSection id="C1AE655CFFDDCD6C1E7EFD20E6E7FB62" pageId="46" pageNumber="214" type="discussion">
<paragraph id="890B36D7FFDDCD6C1E7EFD20E2AEFD33" blockId="46.[125,220,647,671]" box="[125,220,647,671]" pageId="46" pageNumber="214">
<emphasis id="BBC0EAC5FFDDCD6C1E7EFD20E2AEFD33" box="[125,220,647,671]" italics="true" pageId="46" pageNumber="214">Remarks</emphasis>
</paragraph>
<paragraph id="890B36D7FFDDCD6C1E7EFD10E016FBE2" blockId="46.[125,1173,695,1230]" pageId="46" pageNumber="214">
<bibRefCitation id="ED254B26FFDDCD6C1E7EFD10E396FD63" author="Monniot C &amp; Monniot F" box="[125,484,695,719]" pageId="46" pageNumber="214" pagination="567 - 582" refId="ref29238" refString="Monniot C, Monniot F. 1984. Ascidies littorales de Guadeloupe VII. Especes nouvelles et complementaires a l'inventaire. Bulletin du Museum National d'Histoire Naturelle Serie 4 6 A: 567 - 582." type="journal article" year="1984">Monniot and Monniot (1984)</bibRefCitation>
examined material from Hellers
<typeStatus id="560F8875FFDDCD6C1D7BFD10E1DBFD63" box="[888,937,695,719]" pageId="46" pageNumber="214">type</typeStatus>
locality,
<collectingCountry id="F1A37647FFDDCD6C1A1AFD10E6E2FD63" box="[1049,1168,695,719]" name="Sri Lanka" pageId="46" pageNumber="214">Sri Lanka</collectingCountry>
, and found it agreed with the
<typeStatus id="560F8875FFDDCD6C1FE1FD70E061FD43" box="[482,531,727,751]" pageId="46" pageNumber="214">type</typeStatus>
specimen of
<taxonomicName id="4EB44D54FFDDCD6C1CB4FD70E18DFD43" authority="Heller, 1878" authorityName="Heller" authorityYear="1878" box="[695,1023,727,751]" class="Ascidiacea" family="Styelidae" genus="Styela" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="areolata">
<emphasis id="BBC0EAC5FFDDCD6C1CB4FD70E12EFD43" box="[695,860,727,751]" italics="true" pageId="46" pageNumber="214">Styela areolata</emphasis>
<bibRefCitation id="ED254B26FFDDCD6C1D64FD70E18DFD43" author="Heller C" box="[871,1023,727,751]" pageId="46" pageNumber="214" pagination="2 - 28" refId="ref27776" refString="Heller C. 1878. Beitrage zur nahern Kenntnis der Tunicaten. Sitzungsberichte der Mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften Wien 77: 2 - 28." type="journal article" year="1878">Heller, 1878</bibRefCitation>
</taxonomicName>
, in having a deeply curved gut loop and about 12 openings of the short vas deferens on the surface of the ovary.
<bibRefCitation id="ED254B26FFDDCD6C1EFAFCB0E3B8FC83" author="Nishikawa T" box="[249,458,791,815]" pageId="46" pageNumber="214" pagination="25 - 170" refId="ref29462" refString="Nishikawa T. 1991. The ascidians of the Japan Sea. II. Publications of the Seto Marine Biology Laboratory 35: 25 - 170." type="journal article" year="1991">Nishikawa (1991)</bibRefCitation>
found three to four similar openings in some from
<collectingCountry id="F1A37647FFDDCD6C1A20FCB0E616FC83" box="[1059,1124,791,815]" name="Japan" pageId="46" pageNumber="214">Japan</collectingCountry>
but never as many as the
<collectingCountry id="F1A37647FFDDCD6C1F75FC90E394FCE3" box="[374,486,823,847]" name="Sri Lanka" pageId="46" pageNumber="214">Sri Lanka</collectingCountry>
specimens. As a consequence, these authors proposed that
<taxonomicName id="4EB44D54FFDDCD6C1E7EFCF0E3B3FCC3" authority="(Hartmeyer, 1906)" baseAuthorityName="Hartmeyer" baseAuthorityYear="1906" box="[125,449,855,879]" class="Ascidiacea" family="Styelidae" genus="Cnemidocarpa" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="irene">
<emphasis id="BBC0EAC5FFDDCD6C1E7EFCF0E2A5FCC3" box="[125,215,855,879]" italics="true" pageId="46" pageNumber="214">C. irene</emphasis>
(
<bibRefCitation id="ED254B26FFDDCD6C1EE9FCF0E3C8FCC3" author="Hartmeyer R" box="[234,442,855,879]" pageId="46" pageNumber="214" pagination="1 - 30" refId="ref27638" refString="Hartmeyer R. 1906. Ein Beitrag zur Kenntnis der japanischen Ascidienfauna. Zoologischer Anzeiger 31: 1 - 30." type="journal article" year="1906">Hartmeyer, 1906</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F1A37647FFDDCD6C1C13FCF0E023FCC3" box="[528,593,855,879]" name="Japan" pageId="46" pageNumber="214">Japan</collectingCountry>
(with
<taxonomicName id="4EB44D54FFDDCD6C1CA6FCF0E18BFCC3" authority="Hartmeyer, 1919" authorityName="Hartmeyer" authorityYear="1919" box="[677,1017,855,879]" class="Ascidiacea" family="Styelidae" genus="Cnemidocarpa" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="valborgi">
<emphasis id="BBC0EAC5FFDDCD6C1CA6FCF0E150FCC3" box="[677,802,855,879]" italics="true" pageId="46" pageNumber="214">C. valborgi</emphasis>
<bibRefCitation id="ED254B26FFDDCD6C1D2DFCF0E18BFCC3" author="Hartmeyer R" box="[814,1017,855,879]" pageId="46" pageNumber="214" pagination="1 - 150" refId="ref27688" refString="Hartmeyer R. 1919. Ascidien. In: Results of Dr E. Mjoberg's Swedish scientific expedtions to Australia 1910 - 13. Kungliga Svenska Vetenskapsakademiens Handlingar 60: 1 - 150." type="journal article" year="1919">Hartmeyer, 1919</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4EB44D54FFDDCD6C1A38FCF0E33AFC23" authority="Hartmeyer, 1927" authorityName="Hartmeyer" authorityYear="1927" class="Ascidiacea" family="Styelidae" genus="Cnemidocarpa" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="irma">
<emphasis id="BBC0EAC5FFDDCD6C1A38FCF0E6E6FCC3" box="[1083,1172,855,879]" italics="true" pageId="46" pageNumber="214">C. irma</emphasis>
<bibRefCitation id="ED254B26FFDDCD6C1E7EFCD0E33AFC23" author="Hartmeyer R" box="[125,328,887,911]" pageId="46" pageNumber="214" pagination="157 - 194" refId="ref27751" refString="Hartmeyer R. 1927. Zur Kenntnis phlebobranchiater und dictyobranchiater Ascidien. Mitteilungen aus dem Zoologischen Museum in Berlin 13: 157 - 194." type="journal article" year="1927">Hartmeyer, 1927</bibRefCitation>
</taxonomicName>
from northwestern
<collectingCountry id="F1A37647FFDDCD6C1C34FCD0E0EDFC23" box="[567,671,887,911]" name="Australia" pageId="46" pageNumber="214">Australia</collectingCountry>
as junior synonyms) was distinct from
<taxonomicName id="4EB44D54FFDDCD6C1A7AFCD0E300FC02" authority="Heller, 1878" authorityName="Heller" authorityYear="1878" class="Ascidiacea" family="Styelidae" genus="Styela" kingdom="Animalia" order="Stolidobranchia" pageId="46" pageNumber="214" phylum="Chordata" rank="species" species="areolata">
<emphasis id="BBC0EAC5FFDDCD6C1A7AFCD0E2A6FC02" italics="true" pageId="46" pageNumber="214">C. areolata</emphasis>
<bibRefCitation id="ED254B26FFDDCD6C1EDDFC31E300FC02" author="Heller C" box="[222,370,918,942]" pageId="46" pageNumber="214" pagination="2 - 28" refId="ref27776" refString="Heller C. 1878. Beitrage zur nahern Kenntnis der Tunicaten. Sitzungsberichte der Mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften Wien 77: 2 - 28." type="journal article" year="1878">Heller, 1878</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F1A37647FFDDCD6C1FBEFC31E047FC02" box="[445,565,918,942]" name="Sri Lanka" pageId="46" pageNumber="214">Sri Lanka</collectingCountry>
. Specimens assigned to this species by
<bibRefCitation id="ED254B26FFDDCD6C1A05FC31E6E6FC02" author="Kott P" box="[1030,1172,918,942]" pageId="46" pageNumber="214" pagination="1 - 440" refId="ref28254" refString="Kott P. 1985. The Australian Ascidiacea Pt 1, Phlebobranchia and Stolidobranchia. Memoirs of the Queensland Museum 23: 1 - 440." type="journal article" year="1985">Kott (1985)</bibRefCitation>
from Australian locations (Heron Is, Cockburn Sound, South Australia) have been reexamined together with specimens from
<collectingCountry id="F1A37647FFDDCD6C1C53FC71E005FC42" box="[592,631,982,1006]" name="Fiji" pageId="46" pageNumber="214">Fiji</collectingCountry>
and the South
<collectingCountry id="F1A37647FFDDCD6C1D24FC71E11CFC42" box="[807,878,982,1006]" name="China" pageId="46" pageNumber="214">China</collectingCountry>
Sea (see
<bibRefCitation id="ED254B26FFDDCD6C1DDAFC71E621FC42" author="Kott P" box="[985,1107,982,1006]" pageId="46" pageNumber="214" pagination="147 - 212" refId="ref28225" refString="Kott P. 1981. The ascidians of the reef flats of Fiji. Proceedings of the Linnean Society of New South Wales 105: 147 - 212." type="journal article" year="1981">Kott 1981</bibRefCitation>
;
<bibRefCitation id="ED254B26FFDDCD6C1A5CFC71E303FBA2" author="Kott P &amp; Goodbody IG" pageId="46" pageNumber="214" pagination="503 - 554" refId="ref28652" refString="Kott P, Goodbody IG. 1982. The ascidians of Hong Kong. In: Morton BS, Tseng CK, editors. Proceedings of the First International Marine Biological Workshop: the flora and fauna of Hong Kong and Southern China, Hong Kong. Volume 1. Hong Kong: Hong Kong University Press. p 503 - 554." type="book chapter" year="1982">Kott and Goodbody 1982</bibRefCitation>
). In all the specimens examined the general morphology is similar, the vas deferens of each gonad opens by a single simple opening on top of the oviducal opening and multiple openings were not detected.
</paragraph>
<paragraph id="890B36D7FFDDCD6C1E9BFBF1E6E7FB62" blockId="46.[125,1173,695,1230]" pageId="46" pageNumber="214">
Nevertheless in view of
<bibRefCitation id="ED254B26FFDDCD6C1FA7FBF1E0F6FBC2" author="Nishikawa T" box="[420,644,1110,1134]" pageId="46" pageNumber="214" pagination="25 - 170" refId="ref29462" refString="Nishikawa T. 1991. The ascidians of the Japan Sea. II. Publications of the Seto Marine Biology Laboratory 35: 25 - 170." type="journal article" year="1991">Nishikawas (1991)</bibRefCitation>
reports of openings of the male duct similar to those in the
<collectingCountry id="F1A37647FFDDCD6C1F0AFBD1E30BFB22" box="[265,377,1142,1166]" name="Sri Lanka" pageId="46" pageNumber="214">Sri Lanka</collectingCountry>
specimen, albeit not so numerous, it appears that the openings of the male duct are sometimes subdivided in this species. At this stage, it appears that such subdivision may be the result of intra-specific variation rather than an indication of speciation.
</paragraph>
</subSubSection>
</treatment>
</document>