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<mods:title id="4A9195DD3338D04D9BCEF3A1FF21B062">A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea</mods:title>
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<mods:namePart id="7CC0A8D67709D22B817D7773535823DB">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="6DDF729E398891F11F5A2F93B3337515">Kompier, Tom</mods:namePart>
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<mods:namePart id="D9032E212A517D43D4E4C6BB5AAA298B">Karube, Haruki</mods:namePart>
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<mods:namePart id="032910293A80A0E36283BFB1EE66554D">Hayashi, Fumio</mods:namePart>
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<heading id="D33181A9FFFEEC03B594FBF1FD8FD7E6" bold="true" box="[151,560,1142,1168]" fontSize="11" level="1" pageId="27" pageNumber="178" reason="1">
<taxonomicName id="4FC64D46FFFEEC03B594FBF1FD8FD7E6" ID-CoL="3CLZP" authority="Kimmins, 1936" authorityName=": Kimmins" authorityYear="1936" box="[151,560,1142,1168]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="BAB2EAD7FFFEEC03B594FBF1FD8FD7E6" bold="true" box="[151,560,1142,1168]" pageId="27" pageNumber="178">
<emphasis id="BAB2EAD7FFFEEC03B594FBF1FED4D7E6" bold="true" box="[151,363,1142,1168]" italics="true" pageId="27" pageNumber="178">Euphaea superba</emphasis>
<bibRefCitation id="EC574B34FFFEEC03B470FBF1FD8FD7E6" author="Kimmins" box="[371,560,1142,1168]" pageId="27" pageNumber="178" refString="Kimmins, D. E. (1936) A new species of Euphaea (Odonata). The Annals and Magazine of Natural History, Series 10, 17, 147 - 149." type="journal article" year="1936">Kimmins, 1936</bibRefCitation>
</emphasis>
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<paragraph id="887936C5FFFEEC03B594FB3AFD62D659" blockId="27.[151,1436,1213,1327]" pageId="27" pageNumber="178">
<taxonomicName id="4FC64D46FFFEEC03B594FB3AFDBAD7A5" ID-CoL="3CLZP" authority=": Kimmins (1936)" authorityName=": Kimmins" authorityYear="1936" box="[151,517,1213,1235]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="BAB2EAD7FFFEEC03B594FB3AFEF5D7A5" box="[151,330,1213,1235]" italics="true" pageId="27" pageNumber="178">Euphaea superba</emphasis>
:
<bibRefCitation id="EC574B34FFFEEC03B45AFB3AFDBAD7A5" author="Kimmins" box="[345,517,1213,1235]" pageId="27" pageNumber="178" refString="Kimmins, D. E. (1936) A new species of Euphaea (Odonata). The Annals and Magazine of Natural History, Series 10, 17, 147 - 149." type="journal article" year="1936">Kimmins (1936)</bibRefCitation>
</taxonomicName>
, pp. 147149,
<figureCitation id="10FD2A40FFFEEC03B79DFB3AFD47D7A5" box="[670,760,1213,1235]" captionStart-0="FIGURE1" captionStart-1="FIGURE 2" captionStart-2="FIGURE3" captionStartId-0="3.[151,250,1517,1539]" captionStartId-1="4.[151,250,1803,1825]" captionStartId-2="6.[151,250,990,1012]" captionTargetBox-0="[233,1358,627,1465]" captionTargetBox-1="[295,1288,877,1761]" captionTargetBox-2="[186,1420,213,944]" captionTargetId-0="figure@3.[217,1369,615,1495]" captionTargetId-1="figure@4.[281,1305,857,1781]" captionTargetId-2="figure@6.[153,1433,195,968]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURE1. Anisopleurabipugio. [1ac] ♂,A Luoi and [1dh]♀, Hon Ba.(1a)head and thorax;(1b)appendages,dorsal view; (1c) appendages, lateral view; (1d) body; (1e) head, frontal view; (1f) prothorax,lateral view; (1g) prothorax, dorsal view; (1h) basal half of HW." captionText-1="FIGURE 2. Anisopleura qingyuanensis. [2ad] ♂ and [2ef] ♀, Tu Le. (2a) head and thorax; (2b) head, oblique-dorsal view; (2c) appendages, dorsal view; (2d) appendages, lateral view; (2e) head and thorax; (2f) head, oblique-dorsal view." captionText-2="FIGURE3. Anisopleurayunnanensis. [3ac] ♂,Yunnan and [3d]♀,Sapa.(3a) head and thorax; (3b) appendages, dorsalview; (3c) appendages, lateral view; (3d) head and thorax." httpUri-0="https://zenodo.org/record/3096039/files/figure.png" httpUri-1="https://zenodo.org/record/3096041/files/figure.png" httpUri-2="https://zenodo.org/record/3096045/files/figure.png" pageId="27" pageNumber="178">Figs 13</figureCitation>
[Holotype male (at the British Museum of Natural History) from Tonkin (purchased from H. Fruhstorfer)]; van
<bibRefCitation id="EC574B34FFFEEC03B7AEFB5BFC19D784" author="Tol" box="[685,934,1244,1266]" pageId="27" pageNumber="178" refString="van Tol, J. &amp; Rozendaal, F. G. (1995) Records of Calopterygoidea from Vietnam, with descriptions of two new species (Zygoptera: Amphipterygidae, Calopterygidae, Chlorocyphidae, Euphaeidae). Odonatologica, 24, 89 - 107." type="journal article" year="1995">Tol &amp; Rozendaal (1995)</bibRefCitation>
, p. 100 [Male specimen (at the National Museum of Natural History, Leiden) from W. Tonkin, H. Fruhstorfer, ex coll. Förster];
<bibRefCitation id="EC574B34FFFEEC03B102FB7CFB76D666" author="Do" box="[1025,1225,1274,1296]" pageId="27" pageNumber="178" refString="Do, M. C. &amp; Dang, T. T. H. (2007) Checklist of dragonfly from Vietnam. Vietnam National University Publisher, Hanoi, 182 pp." type="book" year="2007">Do &amp; Dang (2007)</bibRefCitation>
, p. 130 [Included in species list as species without detailed locality data].
</paragraph>
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<emphasis id="BAB2EAD7FFFEEC03B594FADCFE34D602" bold="true" box="[151,395,1371,1396]" pageId="27" pageNumber="178">Materials examined.</emphasis>
No material available to the authors.
</paragraph>
<paragraph id="887936C5FFFEEC03B5C4FAF8FBB8D519" blockId="27.[151,1436,1371,1648]" pageId="27" pageNumber="178">
<emphasis id="BAB2EAD7FFFEEC03B5C4FAF8FEAED6EE" bold="true" box="[199,273,1407,1432]" pageId="27" pageNumber="178">Notes.</emphasis>
A large sized, brownish winged species, somewhat resembling
<taxonomicName id="4FC64D46FFFEEC03B6DDFAF8FAA5D6EE" authority="Selys, 1853" authorityName="Selys" authorityYear="1853" box="[990,1306,1407,1432]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="BAB2EAD7FFFEEC03B6DDFAF8FB31D6EE" box="[990,1166,1407,1432]" italics="true" pageId="27" pageNumber="178">Euphaea opaca</emphasis>
<bibRefCitation id="EC574B34FFFEEC03B197FAF8FAA5D6EE" author="Selys" box="[1172,1306,1407,1432]" pageId="27" pageNumber="178" refString="Selys Longchamps, E. de (1853) Synopsis des Calopterygines. Bulletins de l'Academie Royale des sciences, des lettres et des beaux-arts de Belgique, 20, 1 - 73." type="journal article" year="1853">Selys, 1853</bibRefCitation>
</taxonomicName>
. According to M. Hämäläinen (
<emphasis id="BAB2EAD7FFFEEC03B479FA22FE07D6CA" box="[378,440,1443,1468]" italics="true" pageId="27" pageNumber="178">in litt</emphasis>
.) there is also one male specimen of
<taxonomicName id="4FC64D46FFFEEC03B66FFA24FB8DD6CA" authorityName=": Kimmins" authorityYear="1936" box="[876,1074,1443,1468]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="BAB2EAD7FFFEEC03B66FFA24FB8DD6CA" box="[876,1074,1443,1468]" italics="true" pageId="27" pageNumber="178">Euphaea superba</emphasis>
</taxonomicName>
from Tonkin (without further details) at NRS Stockholm. The specimen originates from Coll. F. Ris, and includes his identification label
<taxonomicName id="4FC64D46FFFEEC03B5A3FA6CFEBBD572" authority="Phan &amp; Kompier &amp; Karube &amp; Hayashi, 2018" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[160,260,1515,1540]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="178" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="BAB2EAD7FFFEEC03B5A3FA6CFEBBD572" box="[160,260,1515,1540]" italics="true" pageId="27" pageNumber="178">Euphaea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A18157ACFFFEEC03B408FA6BFEC0D572" box="[267,383,1516,1540]" pageId="27" pageNumber="178" rank="species">spec. nov.</taxonomicNameLabel>
It is possible that this specimen was also purchased from Hans Fruhstorfer. Obviously this is a very rare species in northern Vietnam. According to information received from Haomiao Zhang and M. Hämäläinen (
<emphasis id="BAB2EAD7FFFEEC03B42DF9B2FEC6D53A" box="[302,377,1587,1612]" italics="true" pageId="27" pageNumber="178">in litt.)</emphasis>
, in China
<taxonomicName id="4FC64D46FFFEEC03B4ECF9B4FD0CD53A" authorityName=": Kimmins" authorityYear="1936" box="[495,691,1587,1612]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="BAB2EAD7FFFEEC03B4ECF9B4FD0CD53A" box="[495,691,1587,1612]" italics="true" pageId="27" pageNumber="178">Euphaea superba</emphasis>
</taxonomicName>
occurs in Guangxi and Guizhou; the published
<taxonomicName id="4FC64D46FFFEEC03B1CAF9B2FAFDD53A" authorityName=": Kimmins" authorityYear="1936" box="[1225,1346,1587,1612]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="BAB2EAD7FFFEEC03B1CAF9B2FAFDD53A" box="[1225,1346,1587,1612]" italics="true" pageId="27" pageNumber="178">E. superba</emphasis>
</taxonomicName>
records from Guangdong and Hong Kong (cf.
<bibRefCitation id="EC574B34FFFEEC03B746F9D0FC89D506" author="Wilson" box="[581,822,1623,1648]" pageId="27" pageNumber="178" refString="Wilson, K. D. P. &amp; Zaifu, X. (2007) Odonata of Guangdong, Hong Kong and Macau, South China, part 1: Zygoptera. International Journal of Odonatology, 10, 87 - 128, pls. I - VIII." type="journal article" year="2007">Wilson &amp; Zaifu 2007</bibRefCitation>
) refer to
<taxonomicName id="4FC64D46FFFEEC03B69EF9DEFBBFD506" authorityName="Selys" authorityYear="1853" box="[925,1024,1625,1648]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="BAB2EAD7FFFEEC03B69EF9DEFBBFD506" box="[925,1024,1625,1648]" italics="true" pageId="27" pageNumber="178">E. opaca</emphasis>
</taxonomicName>
.
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