600 lines
73 KiB
XML
600 lines
73 KiB
XML
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<mods:title id="52796F83DE6E40914F165AB49B96F5AE">A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea</mods:title>
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<mods:namePart id="CBA88389D6AA4FF83FEAF2EB70D87894">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="86BB55308D564D1ED3F7443FDF7337A0">Kompier, Tom</mods:namePart>
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<mods:namePart id="0875AB33DDDD284B193F104B76B00AE5">Karube, Haruki</mods:namePart>
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<mods:namePart id="482AD27BDCDD2D531A4F80A03DF47ECF">Hayashi, Fumio</mods:namePart>
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<mods:title id="132E2E41C2A27581CE4557088CDE3335">Zootaxa</mods:title>
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<mods:date id="DD939A94073CCA5C3343A3B84B1C410C">2018</mods:date>
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<heading id="D33181A9FFF0EC0DB594FA79FD45D56E" bold="true" box="[151,762,1534,1560]" fontSize="11" level="1" pageId="21" pageNumber="172" reason="1">
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<emphasis id="BAB2EAD7FFF0EC0DB594FA79FD45D56E" bold="true" box="[151,762,1534,1560]" pageId="21" pageNumber="172">
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<taxonomicName id="4FC64D46FFF0EC0DB594FA79FD30D56E" ID-CoL="9X2NT" authority="Kompier & Hayashi" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[151,655,1534,1560]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="sanguinea" status="sp. nov.">
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<emphasis id="BAB2EAD7FFF0EC0DB594FA79FE37D56E" bold="true" box="[151,392,1534,1560]" italics="true" pageId="21" pageNumber="172">Euphaea sanguinea</emphasis>
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Kompier & Hayashi
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</taxonomicName>
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,
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<taxonomicNameLabel id="A18157ACFFF0EC0DB79EFA79FD45D56E" box="[669,762,1534,1560]" pageId="21" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="887936C5FFF0EC0DB594F9A6FEA7D54C" blockId="21.[151,762,1534,1594]" box="[151,280,1569,1594]" pageId="21" pageNumber="172">
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(
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<figureCitation id="10FD2A40FFF0EC0DB59CF9A6FEAFD54C" box="[159,272,1569,1594]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="21" pageNumber="172">Fig. 17f–j</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C0DC654EFFF0EC0DB594F9EEFD43D5E8" pageId="21" pageNumber="172" type="reference_group">
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<paragraph id="887936C5FFF0EC0DB594F9EEFD43D5E8" blockId="21.[151,1331,1641,1694]" pageId="21" pageNumber="172">
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<taxonomicName id="4FC64D46FFF0EC0DB594F9EEFEEAD509" ID-CoL="9WTZV" authorityName=": Selys" authorityYear="1859" box="[151,341,1641,1663]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="ochracea">
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<emphasis id="BAB2EAD7FFF0EC0DB594F9EEFEEAD509" box="[151,341,1641,1663]" italics="true" pageId="21" pageNumber="172">Euphaea ochracea</emphasis>
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</taxonomicName>
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[nec.
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<bibRefCitation id="EC574B34FFF0EC0DB497F9EEFDB6D509" author="Selys" box="[404,521,1641,1663]" pageId="21" pageNumber="172" refString="Selys Longchamps, E. de (1859) Additions au synopsis des Calopterygines. Bulletins de l'Academie royale de Belgique, Serie 2, 7, 437 - 451." type="journal article" year="1859">Selys 1859</bibRefCitation>
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]:
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<bibRefCitation id="EC574B34FFF0EC0DB71AF9EDFD64D509" author="Do" box="[537,731,1641,1663]" pageId="21" pageNumber="172" refString="Do, M. C. & Dang, T. T. H. (2007) Checklist of dragonfly from Vietnam. Vietnam National University Publisher, Hanoi, 182 pp." type="book" year="2007">Do & Dang (2007)</bibRefCitation>
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, p. 36 [Records from
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<collectingRegion id="4A02F827FFF0EC0DB6BBF9EDFB9AD509" box="[952,1061,1642,1663]" country="Vietnam" name="Lam Dong" pageId="21" pageNumber="172">Lam Dong</collectingRegion>
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and
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<collectingRegion id="4A02F827FFF0EC0DB155F9EDFB06D509" box="[1110,1209,1641,1663]" country="Vietnam" name="Dong Nai" pageId="21" pageNumber="172">Dong Nai</collectingRegion>
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Provinces].
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<taxonomicName id="4FC64D46FFF0EC0DB594F90FFEEAD5E8" ID-CoL="9WTZV" authorityName=": Selys" authorityYear="1859" box="[151,341,1672,1694]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="ochracea">
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||
<emphasis id="BAB2EAD7FFF0EC0DB594F90FFEEAD5E8" box="[151,341,1672,1694]" italics="true" pageId="21" pageNumber="172">Euphaea ochracea</emphasis>
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</taxonomicName>
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||
?ssp.:
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||
<bibRefCitation id="EC574B34FFF0EC0DB498F90FFD81D5E8" author="Kosterin" box="[411,574,1672,1694]" pageId="21" pageNumber="172" refString="Kosterin, O. E. (2016) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2016. International Dragonfly Fund Report. Journal of the International Dragonfly Fund, 98, 1 - 85." type="journal article" year="2016">Kosterin (2016)</bibRefCitation>
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||
, p. 24–26, Fig. 25.
|
||
</paragraph>
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</subSubSection>
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<subSubSection id="C0DC654EFFF0EC0DB594F94DFBD2D471" pageId="21" pageNumber="172" type="materials_examined">
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<paragraph id="887936C5FFF0EC0DB594F94DFBD2D471" blockId="21.[151,1437,1738,2015]" pageId="21" pageNumber="172">
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<materialsCitation id="38AE3C98FFF0EC0DB594F94DFBD5D471" ID-GBIF-Occurrence="1805098078" collectingDate="2016-06-17" collectionCode="TMUZ" collectorName="T. Kompier & Deposited" country="Japan" county="Systematic Zoology Institute in Tokyo Metropolitan University" location="Bao Loc" municipality="Zoological Collection of Systematic Zoology Institute" pageId="21" pageNumber="172" specimenCount="1" specimenCount-male="1" stateProvince="Tokyo" typeStatus="holotype">
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<emphasis id="BAB2EAD7FFF0EC0DB594F94DFEB4D595" bold="true" box="[151,267,1738,1763]" pageId="21" pageNumber="172">
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<typeStatus id="577D8867FFF0EC0DB594F94DFEB9D595" box="[151,262,1738,1763]" pageId="21" pageNumber="172" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCount id="9EC0FD4CFFF0EC0DB415F94CFE92D594" box="[278,301,1739,1762]" pageId="21" pageNumber="172" type="male">♂</specimenCount>
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,
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<location id="8D19601EFFF0EC0DB43DF94CFE1AD594" LSID="urn:lsid:plazi:treatment:006F87D3FFF0EC0FB594FA79FEEED063:8D19601EFFF0EC0DB43DF94CFE1AD594" box="[318,421,1739,1762]" country="Japan" county="Systematic Zoology Institute in Tokyo Metropolitan University" municipality="Zoological Collection of Systematic Zoology Institute" name="Bao Loc" pageId="21" pageNumber="172" stateProvince="Tokyo">Bao Loc</location>
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,
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<location id="8D19601EFFF0EC0DB4B6F94CFDC4D594" LSID="urn:lsid:plazi:treatment:006F87D3FFF0EC0FB594FA79FEEED063:8D19601EFFF0EC0DB4B6F94CFDC4D594" box="[437,635,1739,1763]" country="Japan" county="Systematic Zoology Institute in Tokyo Metropolitan University" municipality="Zoological Collection of Systematic Zoology Institute" name="Lam Dong Prov." pageId="21" pageNumber="172" stateProvince="Tokyo">Lam Dong Prov.</location>
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,
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<collectingDate id="EC3CE9EDFFF0EC0DB789F94DFCB3D595" box="[650,780,1738,1763]" pageId="21" pageNumber="172" value="2016-06-17">
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<date id="FC781005FFF0EC0DB789F94DFCB3D595" box="[650,780,1738,1763]" pageId="21" pageNumber="172" value="2016-06-17">17.VI.2016</date>
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</collectingDate>
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,
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<collectorName id="25335313FFF0EC0DB61EF94CFC1CD595" box="[797,931,1738,1763]" pageId="21" pageNumber="172">T. Kompier</collectorName>
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leg.
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<collectorName id="25335313FFF0EC0DB6E1F94DFBEBD595" box="[994,1108,1738,1763]" pageId="21" pageNumber="172">Deposited</collectorName>
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at
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<collectingMunicipality id="681DACBFFFF0EC0DB183F94DFE65D471" pageId="21" pageNumber="172">
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Zoological Collection of
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<collectingCounty id="61184E49FFF0EC0DB594F969FE65D471" box="[151,474,1774,1799]" pageId="21" pageNumber="172">Systematic Zoology Institute</collectingCounty>
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</collectingMunicipality>
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in Tokyo Metropolitan University,
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<collectingRegion id="4A02F827FFF0EC0DB66BF969FC0DD471" box="[872,946,1774,1799]" country="Japan" name="Tokyo" pageId="21" pageNumber="172">Tokyo</collectingRegion>
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,
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||
<collectingCountry id="F0D17655FFF0EC0DB6BEF968FC42D470" box="[957,1021,1775,1798]" name="Japan" pageId="21" pageNumber="172">Japan</collectingCountry>
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(
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<collectionCode id="EED7AE00FFF0EC0DB10FF968FBDDD471" box="[1036,1122,1775,1799]" pageId="21" pageNumber="172">TMUZ</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<materialsCitation id="38AE3C98FFF0EC0DB5C4F895FA36D439" ID-GBIF-Occurrence="1805098090" collectingDate="1996-06-15" collectingDateMax="2002-06-01" collectingDateMin="1996-06-15" collectionCode="TMUZ" collectorName="H. Karube" county="Lam Dong Prov." location="Bao Loc" municipality="Lam Dong Prov." pageId="21" pageNumber="172" specimenCount="4" specimenCount-male="4" typeStatus="paratype">
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<emphasis id="BAB2EAD7FFF0EC0DB5C4F895FE34D45D" bold="true" box="[199,395,1810,1835]" pageId="21" pageNumber="172">
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Male
|
||
<typeStatus id="577D8867FFF0EC0DB40DF895FE39D45D" box="[270,390,1810,1835]" pageId="21" pageNumber="172" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9EC0FD4CFFF0EC0DB496F894FE07D45D" box="[405,440,1811,1835]" pageId="21" pageNumber="172" type="male">1♂</specimenCount>
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the same location, date and collector as
|
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<typeStatus id="577D8867FFF0EC0DB69BF895FC45D45D" box="[920,1018,1810,1835]" pageId="21" pageNumber="172" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="EED7AE00FFF0EC0DB10FF894FBDDD45D" box="[1036,1122,1811,1835]" pageId="21" pageNumber="172">TMUZ</collectionCode>
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);
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<specimenCount id="9EC0FD4CFFF0EC0DB17BF894FB22D45D" box="[1144,1181,1811,1835]" pageId="21" pageNumber="172" type="male">2♂</specimenCount>
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,
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<location id="8D19601EFFF0EC0DB1AFF894FAAED45C" LSID="urn:lsid:plazi:treatment:006F87D3FFF0EC0FB594FA79FEEED063:8D19601EFFF0EC0DB1AFF894FAAED45C" box="[1196,1297,1811,1834]" county="Lam Dong Prov." municipality="Lam Dong Prov." name="Bao Loc" pageId="21" pageNumber="172">Bao Loc</location>
|
||
,
|
||
<collectingMunicipality id="681DACBFFFF0EC0DB023F894FF6BD438" pageId="21" pageNumber="172">Lam Dong Prov.</collectingMunicipality>
|
||
,
|
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<collectingDate id="EC3CE9EDFFF0EC0DB5DCF8B1FE34D439" box="[223,395,1846,1871]" pageId="21" pageNumber="172" value="1996-06-15" valueMax="1996-06-17" valueMin="1996-06-15">
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<date id="FC781005FFF0EC0DB5DCF8B1FE34D439" box="[223,395,1846,1871]" pageId="21" pageNumber="172" value="1996-06-15" valueMax="1996-06-17" valueMin="1996-06-15">
|
||
15–17.
|
||
<bibRefCitation id="EC574B34FFF0EC0DB42AF8B1FE34D439" author="VI" box="[297,395,1846,1871]" pageId="21" pageNumber="172" refString="Davies, D. A. L. & Yang, B. (1996) New species of Bayadera Selys and Schmidtiphaea Asahina from China (Odonata, Euphaeidae). Tijdschrift voor Entomologie, 139, 145 - 155." type="journal article" year="1996">VI.1996</bibRefCitation>
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||
</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="25335313FFF0EC0DB495F8B0FDB4D439" box="[406,523,1846,1871]" pageId="21" pageNumber="172">H. Karube</collectorName>
|
||
leg. (KMP);
|
||
<specimenCount id="9EC0FD4CFFF0EC0DB7A3F8B0FD7AD439" box="[672,709,1847,1871]" pageId="21" pageNumber="172" type="male">1♂</specimenCount>
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||
, Bao Loc,
|
||
<collectingCounty id="61184E49FFF0EC0DB63CF8B0FC43D438" box="[831,1020,1847,1871]" pageId="21" pageNumber="172">Lam Dong Prov.</collectingCounty>
|
||
,
|
||
<collectingDate id="EC3CE9EDFFF0EC0DB104F8B0FBC5D439" box="[1031,1146,1847,1871]" pageId="21" pageNumber="172" value="2002-06-01">
|
||
<date id="FC781005FFF0EC0DB104F8B0FBC5D439" box="[1031,1146,1847,1871]" pageId="21" pageNumber="172" value="2002-06-01">1.VI.2002</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="25335313FFF0EC0DB185F8B0FB44D439" box="[1158,1275,1846,1871]" pageId="21" pageNumber="172">H. Karube</collectorName>
|
||
leg. (KMP);
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0DC654EFFF0EC0DB5C4F8DDFF46D4E1" pageId="21" pageNumber="172" type="materials_examined">
|
||
<paragraph id="887936C5FFF0EC0DB5C4F8DDFF46D4E1" blockId="21.[151,1437,1738,2015]" pageId="21" pageNumber="172">
|
||
<materialsCitation id="38AE3C98FFF0EC0DB5C4F8DDFF4AD4E1" ID-GBIF-Occurrence="1805098096" collectingDate="2017-10-08" collectionCode="PQTC" collectorName="Hoang Quang Duy" location="Dak Lak Prov." municipality="Other" pageId="21" pageNumber="172" specimenCount="1" specimenCount-male="1" stateProvince="Ka">
|
||
<emphasis id="BAB2EAD7FFF0EC0DB5C4F8DDFDBDD405" bold="true" box="[199,514,1882,1907]" pageId="21" pageNumber="172">
|
||
<collectingMunicipality id="681DACBFFFF0EC0DB5C4F8DDFEB0D405" box="[199,271,1882,1907]" pageId="21" pageNumber="172">Other</collectingMunicipality>
|
||
materials examined
|
||
</emphasis>
|
||
.
|
||
<specimenCount id="9EC0FD4CFFF0EC0DB712F8DCFD89D405" box="[529,566,1883,1907]" pageId="21" pageNumber="172" type="male">1♂</specimenCount>
|
||
, Nam
|
||
<collectingRegion id="4A02F827FFF0EC0DB781F8DCFD1CD404" box="[642,675,1883,1906]" country="Bhutan" name="Ha" pageId="21" pageNumber="172">Ka</collectingRegion>
|
||
Nature Reserve,
|
||
<location id="8D19601EFFF0EC0DB66AF8DDFBAED404" LSID="urn:lsid:plazi:treatment:006F87D3FFF0EC0FB594FA79FEEED063:8D19601EFFF0EC0DB66AF8DDFBAED404" box="[873,1041,1882,1907]" municipality="Other" name="Dak Lak Prov." pageId="21" pageNumber="172" stateProvince="Ka">Dak Lak Prov.</location>
|
||
,
|
||
<collectingDate id="EC3CE9EDFFF0EC0DB11EF8DCFB37D405" box="[1053,1160,1883,1907]" pageId="21" pageNumber="172" value="2017-10-08">
|
||
<date id="FC781005FFF0EC0DB11EF8DCFB37D405" box="[1053,1160,1883,1907]" pageId="21" pageNumber="172" value="2017-10-08">8.X.2017</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="25335313FFF0EC0DB196F8DCFAD4D405" box="[1173,1387,1883,1907]" pageId="21" pageNumber="172">Hoang Quang Duy</collectorName>
|
||
leg. (
|
||
<collectionCode id="EED7AE00FFF0EC0DB59CF8F8FF52D4E1" box="[159,237,1919,1943]" pageId="21" pageNumber="172">PQTC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0DC654EFFF0EC0DB5C4F825FB0CD4CD" box="[199,1203,1954,1979]" pageId="21" pageNumber="172" type="description">
|
||
<paragraph id="887936C5FFF0EC0DB5C4F825FB0CD4CD" blockId="21.[151,1437,1738,2015]" box="[199,1203,1954,1979]" pageId="21" pageNumber="172">
|
||
<materialsCitation id="38AE3C98FFF0EC0DB5C4F825FB0CD4CD" ID-GBIF-Occurrence="1805098103" box="[199,1203,1954,1979]" collectingDate="2016-06-17" collectionCode="TMUZ" collectorName="T. Kompier" location="Bao Loc" pageId="21" pageNumber="172" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong" typeStatus="paratype">
|
||
<emphasis id="BAB2EAD7FFF0EC0DB5C4F825FE27D4CD" bold="true" box="[199,408,1954,1979]" pageId="21" pageNumber="172">
|
||
Female
|
||
<typeStatus id="577D8867FFF0EC0DB426F825FE2CD4CD" box="[293,403,1954,1979]" pageId="21" pageNumber="172" type="paratype">paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9EC0FD4CFFF0EC0DB49DF824FE7ED4CD" box="[414,449,1955,1979]" pageId="21" pageNumber="172" type="female">1♀</specimenCount>
|
||
,
|
||
<location id="8D19601EFFF0EC0DB4CCF824FD8ED4CC" LSID="urn:lsid:plazi:treatment:006F87D3FFF0EC0FB594FA79FEEED063:8D19601EFFF0EC0DB4CCF824FD8ED4CC" box="[463,561,1955,1978]" name="Bao Loc" pageId="21" pageNumber="172" stateProvince="Lam Dong">Bao Loc</location>
|
||
,
|
||
<collectingRegion id="4A02F827FFF0EC0DB73EF824FD0AD4CD" box="[573,693,1955,1979]" country="Vietnam" name="Lam Dong" pageId="21" pageNumber="172">Lam Dong</collectingRegion>
|
||
Prov.,
|
||
<collectingDate id="EC3CE9EDFFF0EC0DB607F825FC39D4CD" box="[772,902,1954,1979]" pageId="21" pageNumber="172" value="2016-06-17">
|
||
<date id="FC781005FFF0EC0DB607F825FC39D4CD" box="[772,902,1954,1979]" pageId="21" pageNumber="172" value="2016-06-17">17.VI.2016</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="25335313FFF0EC0DB691F824FBACD4CD" box="[914,1043,1954,1979]" pageId="21" pageNumber="172">T. Kompier</collectorName>
|
||
leg. (
|
||
<collectionCode id="EED7AE00FFF0EC0DB152F824FB18D4CD" box="[1105,1191,1955,1979]" pageId="21" pageNumber="172">TMUZ</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0DC654EFFF0EC0DB5C4F841FA2FD4A9" box="[199,1424,1990,2015]" pageId="21" pageNumber="172" type="etymology">
|
||
<paragraph id="887936C5FFF0EC0DB5C4F841FA2FD4A9" blockId="21.[151,1437,1738,2015]" box="[199,1424,1990,2015]" pageId="21" pageNumber="172">
|
||
<emphasis id="BAB2EAD7FFF0EC0DB5C4F841FEF1D4A9" bold="true" box="[199,334,1990,2015]" pageId="21" pageNumber="172">Etymology.</emphasis>
|
||
The specific name “
|
||
<taxonomicName id="4FC64D46FFF0EC0DB736F840FD17D4A8" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[565,680,1991,2014]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF0EC0DB736F840FD17D4A8" box="[565,680,1991,2014]" italics="true" pageId="21" pageNumber="172">sanguinea</emphasis>
|
||
</taxonomicName>
|
||
” refers to the clear-cut red coloration of abdomen segments S2–6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0DC654EFFF3EC0EB5C4FF10FC76D60E" pageId="22" pageNumber="173" type="description">
|
||
<paragraph id="887936C5FFF3EC0EB5C4FF10FBC1D3A3" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FF10FE50D3C6" bold="true" box="[199,495,151,176]" pageId="22" pageNumber="173">
|
||
Description of
|
||
<typeStatus id="577D8867FFF3EC0EB481FF10FE54D3C6" box="[386,491,151,176]" pageId="22" pageNumber="173" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="BAB2EAD7FFF3EC0EB4FFFF10FD86D3C6" box="[508,569,151,176]" italics="true" pageId="22" pageNumber="173">Head</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB74DFF10FD09D3C6" box="[590,694,151,176]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17h</figureCitation>
|
||
). Labrum, postclypeus, anteclypeus, genae, base of mandibles, antennae, and dorsal surface of head all black. Compound eyes dark chestnut when alive.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FF58FDC8D2FE" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FF58FEA7D38E" box="[199,280,223,248]" italics="true" pageId="22" pageNumber="173">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB42EFF58FE2FD38E" box="[301,400,223,248]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17f</figureCitation>
|
||
). Prothorax black but middle lobe with a pair of orange marks. Synthorax with black mesepisternum with narrow but distinct orange dorsal stripe and antehumeral stripe. Mesepimeron black with distinct orange stripe over its center, slightly hooked downwards at its distal end. Metepisternum and metepimeron red, with black line over centers and covering metapleural suture. Mesinfraepisternum black with some red, metinfraepisternum red with black margin.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FE12FD2CD2DB" blockId="22.[151,1437,151,2013]" box="[199,659,404,429]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FE12FF43D2DA" box="[199,252,405,428]" italics="true" pageId="22" pageNumber="173">Legs</emphasis>
|
||
black. Coxae and trochanters black.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FE3EFCD4D16E" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FE3EFEB5D2A6" box="[199,266,441,464]" italics="true" pageId="22" pageNumber="173">Wings</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB423FE30FE3CD2A6" box="[288,387,439,464]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17f</figureCitation>
|
||
). Strongly colored reddish orange from base to proximal third of outer wing, thereafter gradually paler amber. Veins are similarly reddish turning amber, but largest veins dark. FW has 30 Ax and 42 Px, HW 26 Ax and 34 Px. Pterostigma dark brown, covering 7 cells.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FDA3FE34D116" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FDA3FE8ED14B" box="[199,305,548,573]" italics="true" pageId="22" pageNumber="173">Abdomen</emphasis>
|
||
. Abdominal S2–6 bright red, ringed black at posterior margins, progressively more distinctly so distally; S7–10 black.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FDEBFB63D1F3" blockId="22.[151,1437,151,2013]" box="[199,1244,620,645]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FDEBFEDCD1F3" box="[199,355,620,645]" italics="true" pageId="22" pageNumber="173">Genital ligula</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB471FDEBFE6DD1F2" box="[370,466,620,645]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17c</figureCitation>
|
||
).
|
||
<taxonomicName id="4FC64D46FFF3EC0EB4E5FDEBFD17D1F3" authorityName="ochracea: Laidlaw" authorityYear="1931" box="[486,680,620,645]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="subSpecies" species="ochracea" subSpecies="ochracea">
|
||
Like
|
||
<emphasis id="BAB2EAD7FFF3EC0EB722FDEAFD17D1F3" box="[545,680,620,645]" italics="true" pageId="22" pageNumber="173">E. ochracea</emphasis>
|
||
</taxonomicName>
|
||
. Vesicle coarsely grooved, similar to
|
||
<emphasis id="BAB2EAD7FFF3EC0EB14EFDEAFB63D1F3" box="[1101,1244,620,645]" italics="true" pageId="22" pageNumber="173">
|
||
<taxonomicName id="4FC64D46FFF3EC0EB14EFDEAFB66D1F3" authorityName=": Selys" authorityYear="1859" box="[1101,1241,620,645]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">E. ochracea</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FD08FE1BD199" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FD08FE35D1DE" box="[199,394,655,680]" italics="true" pageId="22" pageNumber="173">Anal appendages</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB49FFD08FE46D1DE" box="[412,505,655,680]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17j</figureCitation>
|
||
). Black, of usual
|
||
<typeStatus id="577D8867FFF3EC0EB7C6FD17FD47D1DE" box="[709,760,656,680]" pageId="22" pageNumber="173">type</typeStatus>
|
||
, with very short paraprocts in the form of small hooks and double lobed cerci. Cerci with two ventral lobes, the first of which has its proximal edge almost straight and perpendicular to cercus.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FD7AFC1FD062" blockId="22.[151,1437,151,2013]" box="[199,928,764,789]" pageId="22" pageNumber="173">Measurements (in mm). HW 27; abdomen (incl. appendages) 32.</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FC98FC0CD02B" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FC98FDA7D04E" bold="true" box="[199,536,799,824]" pageId="22" pageNumber="173">
|
||
Variation in
|
||
<typeStatus id="577D8867FFF3EC0EB463FC98FE74D04E" box="[352,459,799,824]" pageId="22" pageNumber="173" type="paratype">paratype</typeStatus>
|
||
males
|
||
</emphasis>
|
||
. Four
|
||
<typeStatus id="577D8867FFF3EC0EB765FCA7FD6ED04E" box="[614,721,800,824]" pageId="22" pageNumber="173" type="paratype">paratypes</typeStatus>
|
||
show little morphological variation, excluding size;
|
||
<quantity id="4F3E9B20FFF3EC0EB02FFC98FF7CD02A" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="2.85" metricValueMin="2.65" pageId="22" pageNumber="173" unit="mm" value="27.5" valueMax="28.5" valueMin="26.5">26.5–28.5 mm</quantity>
|
||
in HW length and
|
||
<quantity id="4F3E9B20FFF3EC0EB49BFCC3FD99D02A" box="[408,550,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="3.275" metricValueMax="3.35" metricValueMin="3.2" pageId="22" pageNumber="173" unit="mm" value="32.75" valueMax="33.5" valueMin="32.0">32–33.5 mm</quantity>
|
||
in abdomen including appendages.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FCE0FEA0D0BE" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FCE0FD82D0F6" bold="true" box="[199,573,871,896]" pageId="22" pageNumber="173">
|
||
Description of
|
||
<typeStatus id="577D8867FFF3EC0EB479FCE0FE5AD0F6" box="[378,485,871,896]" pageId="22" pageNumber="173" type="paratype">paratype</typeStatus>
|
||
female
|
||
</emphasis>
|
||
.
|
||
<emphasis id="BAB2EAD7FFF3EC0EB74EFCE0FD35D0F6" box="[589,650,871,896]" italics="true" pageId="22" pageNumber="173">Head</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB798FCE0FD47D0F6" box="[667,760,871,896]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">
|
||
Fig.
|
||
<date id="FC781005FFF3EC0EB7D2FCE0FD47D0F6" box="[721,760,871,896]" pageId="22" pageNumber="173" value="2017-01">17i</date>
|
||
</figureCitation>
|
||
). Black, excluding the yellow markings on labrum, base of mandibles, frons at eye margins, and spots between base of antennae and lateral ocelli. Antennae black. Compound eyes brown.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FC53FB9BD72E" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FC53FEA7D09B" box="[199,280,980,1005]" italics="true" pageId="22" pageNumber="173">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB429FC53FE30D09A" box="[298,399,980,1005]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">
|
||
Fig.
|
||
<quantity id="4F3E9B20FFF3EC0EB461FC53FE30D09A" box="[354,399,980,1004]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="22" pageNumber="173" unit="g" value="17.0">17g</quantity>
|
||
</figureCitation>
|
||
). Anterior lobe of prothorax black with two small yellow dots; middle lobe largely black except for two lateral yellow markings; posterior lobe simple, its distal margin smoothly rounded and broadly yellowish; propleuron black with some yellow marking laterally. Synthorax black with yellow marking like male, but mesinfraepisternum yellow with some black, and metinfraepisternum yellow.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FBE2FB02D70B" blockId="22.[151,1437,151,2013]" box="[199,1213,1124,1149]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FBE2FF43D70A" box="[199,252,1125,1148]" italics="true" pageId="22" pageNumber="173">Legs</emphasis>
|
||
dark grey, with a yellow stripe on each inner femur. Coxae and trochanters dark grey.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FB0EFD0FD7B3" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FB0EFEB5D7D6" box="[199,266,1161,1184]" italics="true" pageId="22" pageNumber="173">Wings</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB41FFB00FE3FD7D6" box="[284,384,1159,1184]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">
|
||
Fig.
|
||
<quantity id="4F3E9B20FFF3EC0EB451FB0FFE3FD7D6" box="[338,384,1160,1184]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="22" pageNumber="173" unit="g" value="17.0">17g</quantity>
|
||
</figureCitation>
|
||
). Weakly yellowish, particularly at basal part. Veins dark. FW 25 Ax and 32 Px, HW 19 Ax and 26 Px. Pterostigma brown, covering 7 cells.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FB48FF64D646" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FB48FE8ED79E" box="[199,305,1231,1256]" italics="true" pageId="22" pageNumber="173">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB43CFB48FE1ED79E" box="[319,417,1231,1256]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">
|
||
Fig.
|
||
<quantity id="4F3E9B20FFF3EC0EB470FB57FE1ED79E" box="[371,417,1232,1256]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="22" pageNumber="173" unit="g" value="17.0">17g</quantity>
|
||
</figureCitation>
|
||
). Black, with lateral yellow stripes in two rows (lower one is shorter than upper one) across S1–6, but only a small anterior mark on S7 and no stripe on S8. S9 a posterolateral oval yellow spot. S10 and cerci black.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FABAFCC7D623" blockId="22.[151,1437,151,2013]" box="[199,888,1340,1365]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FABAFE80D622" box="[199,319,1341,1364]" italics="true" pageId="22" pageNumber="173">Ovipositor</emphasis>
|
||
yellowish, but black on ventral valvula and stylus.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0EB5C4FAE6FC76D60E" blockId="22.[151,1437,151,2013]" box="[199,969,1375,1400]" pageId="22" pageNumber="173">Measurements (in mm). HW 27.5; abdomen (incl. appendages) 28.5.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0DC654EFFF3EC0FB5C4FA03FA80D1BB" lastPageId="23" lastPageNumber="174" pageId="22" pageNumber="173" type="discussion">
|
||
<paragraph id="887936C5FFF3EC0EB5C4FA03FCF8D503" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB5C4FA03FE84D6EB" bold="true" box="[199,315,1412,1437]" pageId="22" pageNumber="173">Remarks.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB4ADFA03FD33D6EA" authority="Phan & Kompier & Karube & Hayashi, 2018" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[430,652,1412,1437]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="sanguinea" status="sp. nov.">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB4ADFA03FD33D6EA" box="[430,652,1412,1437]" italics="true" pageId="22" pageNumber="173">Euphaea sanguinea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A18157ACFFF3EC0EB797FA03FD51D6EB" box="[660,750,1412,1437]" pageId="22" pageNumber="173" rank="species">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB797FA03FD51D6EB" bold="true" box="[660,750,1412,1437]" pageId="22" pageNumber="173">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
resemble those of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB6C6FA02FBF3D6EB" authorityName=": Selys" authorityYear="1859" box="[965,1100,1412,1437]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB6C6FA02FBF3D6EB" box="[965,1100,1412,1437]" italics="true" pageId="22" pageNumber="173">E. ochracea</emphasis>
|
||
</taxonomicName>
|
||
, the geographic variability of which is complicated.
|
||
<taxonomicName id="4FC64D46FFF3EC0EB494FA20FD53D6B6" authority="Selys, 1879" authorityName="Selys" authorityYear="1879" box="[407,748,1447,1472]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="brunnea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB494FA20FDDFD6B6" box="[407,608,1447,1472]" italics="true" pageId="22" pageNumber="173">Euphaea brunnea</emphasis>
|
||
<bibRefCitation id="EC574B34FFF3EC0EB76AFA20FD53D6B6" author="Selys" box="[617,748,1447,1472]" pageId="22" pageNumber="173" refString="Selys Longchamps, E. de (1879) Quatriemes additions au Synopsis des Calopterygines. Bulletin s de l'Academie royale de Belgique, Serie 2, 47, 1 - 63." type="journal article" year="1879">Selys, 1879</bibRefCitation>
|
||
</taxonomicName>
|
||
was described from specimens from Khasi Hills, Meghalay, NE
|
||
<collectingCountry id="F0D17655FFF3EC0EB5CBFA4BFEBAD693" box="[200,261,1484,1509]" name="India" pageId="22" pageNumber="173">India</collectingCountry>
|
||
.
|
||
<bibRefCitation id="EC574B34FFF3EC0EB417FA4BFD1BD692" author="Hamalainen" box="[276,676,1484,1509]" pageId="22" pageNumber="173" refString="Hamalainen, M. (2016) Calopterygoidea of the world: A synonymic list of extant damselfly species of the superfamily Calopterygoidea (sensu lato) (Odonata: Zygoptera). 1 st Edition. 1 January 2016. Privately published online, Espoo, Finland. Available from: http: // www. caloptera. com / World-Calopterygoidea-List. pdf (accessed 12 July 2017)" type="journal article" year="2016" yearSuffix="p">Hämäläinen (2016, p. 26 note 41)</bibRefCitation>
|
||
discusses the uncertainty of the status of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB198FA4BFAD7D693" authorityName="Selys" authorityYear="1879" box="[1179,1384,1484,1509]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="brunnea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB198FA4BFAD7D693" box="[1179,1384,1484,1509]" italics="true" pageId="22" pageNumber="173">Euphaea brunnea</emphasis>
|
||
</taxonomicName>
|
||
and whether it qualifies as a subspecies of
|
||
<emphasis id="BAB2EAD7FFF3EC0EB746FA76FD6BD57E" box="[581,724,1519,1544]" italics="true" pageId="22" pageNumber="173">
|
||
<taxonomicName id="4FC64D46FFF3EC0EB746FA76FD6ED57E" authorityName=": Selys" authorityYear="1859" box="[581,721,1519,1544]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">E. ochracea</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Superficially, specimens from central and northern
|
||
<collectingCountry id="F0D17655FFF3EC0EB019FA68FAC1D57E" box="[1306,1406,1519,1544]" name="Thailand" pageId="22" pageNumber="173">Thailand</collectingCountry>
|
||
as well as northern
|
||
<collectingCountry id="F0D17655FFF3EC0EB453F993FE10D55B" box="[336,431,1556,1581]" name="Vietnam" pageId="22" pageNumber="173">Vietnam</collectingCountry>
|
||
resemble
|
||
<taxonomicName id="4FC64D46FFF3EC0EB722F993FDC1D55B" authorityName="Selys" authorityYear="1879" box="[545,638,1556,1581]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="brunnea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB722F993FDC1D55B" box="[545,638,1556,1581]" italics="true" pageId="22" pageNumber="173">brunnea</emphasis>
|
||
</taxonomicName>
|
||
better than
|
||
<taxonomicName id="4FC64D46FFF3EC0EB602F993FCD6D55B" authorityName=": Selys" authorityYear="1859" box="[769,873,1556,1581]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB602F993FCD6D55B" box="[769,873,1556,1581]" italics="true" pageId="22" pageNumber="173">ochracea</emphasis>
|
||
</taxonomicName>
|
||
(Hämäläinen
|
||
<emphasis id="BAB2EAD7FFF3EC0EB105F992FBF6D55B" box="[1030,1097,1556,1581]" italics="true" pageId="22" pageNumber="173">in litt.</emphasis>
|
||
). In view of the uncertainty of the validity of the taxon
|
||
<taxonomicName id="4FC64D46FFF3EC0EB4D7F9B0FD8ED526" authorityName="Selys" authorityYear="1879" box="[468,561,1591,1616]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="brunnea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB4D7F9B0FD8ED526" box="[468,561,1591,1616]" italics="true" pageId="22" pageNumber="173">brunnea</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB632F9B0FBB3D526" authorityName=": Selys" authorityYear="1859" box="[817,1036,1591,1616]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB632F9B0FBB3D526" box="[817,1036,1591,1616]" italics="true" pageId="22" pageNumber="173">Euphaea ochracea</emphasis>
|
||
</taxonomicName>
|
||
we refrain from the subspecific identification of the specimens of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB717F9DAFD24D503" authorityName=": Selys" authorityYear="1859" box="[532,667,1628,1653]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB717F9DAFD24D503" box="[532,667,1628,1653]" italics="true" pageId="22" pageNumber="173">E. ochracea</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F0D17655FFF3EC0EB7E3F9DBFCFCD503" box="[736,835,1628,1653]" name="Vietnam" pageId="22" pageNumber="173">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF3EC0FB5C4F9F8FB90D26A" blockId="22.[151,1437,151,2013]" lastBlockId="23.[151,1436,151,789]" lastPageId="23" lastPageNumber="174" pageId="22" pageNumber="173">
|
||
Males of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB431F9F8FDAFD5EE" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[306,528,1663,1688]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB431F9F8FDAFD5EE" box="[306,528,1663,1688]" italics="true" pageId="22" pageNumber="173">Euphaea sanguinea</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the cerci. These differ from those of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB1CBF9F8FA23D5EE" authorityName=": Selys" authorityYear="1859" box="[1224,1436,1663,1688]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB1CBF9F8FA23D5EE" box="[1224,1436,1663,1688]" italics="true" pageId="22" pageNumber="173">Euphaea ochracea</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB59CF923FF43D5CA" box="[159,252,1700,1725]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17e</figureCitation>
|
||
) by having a well-developed triangular ventral lobe at the base (
|
||
<figureCitation id="10FD2A40FFF3EC0EB6CFF923FB98D5CB" box="[972,1063,1700,1725]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17j</figureCitation>
|
||
).
|
||
<taxonomicName id="4FC64D46FFF3EC0EB138F923FAA7D5CA" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[1083,1304,1700,1725]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB138F923FAA7D5CA" box="[1083,1304,1700,1725]" italics="true" pageId="22" pageNumber="173">Euphaea sanguinea</emphasis>
|
||
</taxonomicName>
|
||
also differs by the relatively narrow area of reddish orange in HW (
|
||
<figureCitation id="10FD2A40FFF3EC0EB606F940FCDFD596" box="[773,864,1735,1760]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17f</figureCitation>
|
||
) compared to both topotypical material (which has wings almost completely unicolourous) and compared to material of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB6A0F96AFB94D473" authorityName=": Selys" authorityYear="1859" box="[931,1067,1772,1797]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB6A0F96AFB94D473" box="[931,1067,1772,1797]" italics="true" pageId="22" pageNumber="173">E. ochracea</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F0D17655FFF3EC0EB171F96BFB6ED473" box="[1138,1233,1772,1797]" name="Vietnam" pageId="22" pageNumber="173">Vietnam</collectingCountry>
|
||
(
|
||
<figureCitation id="10FD2A40FFF3EC0EB1E2F96BFA80D472" box="[1249,1343,1772,1797]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17a</figureCitation>
|
||
) and by the clear-cut reddish pattern on S2–6 compared to the specimens of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB6A1F896FB94D45E" authorityName=": Selys" authorityYear="1859" box="[930,1067,1807,1832]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB6A1F896FB94D45E" box="[930,1067,1807,1832]" italics="true" pageId="22" pageNumber="173">E. ochracea</emphasis>
|
||
</taxonomicName>
|
||
occurring in
|
||
<collectingCountry id="F0D17655FFF3EC0EB1C5F897FB43D451" box="[1222,1276,1808,1831]" name="Laos" pageId="22" pageNumber="173">Laos</collectingCountry>
|
||
and
|
||
<collectingCountry id="F0D17655FFF3EC0EB035F888FA27D45E" box="[1334,1432,1807,1832]" name="Vietnam" pageId="22" pageNumber="173">Vietnam</collectingCountry>
|
||
, although topotypical material has distinctly redder abdomen, if not as clear-cut (
|
||
<figureCitation id="10FD2A40FFF3EC0EB126F8B3FB20D43B" box="[1061,1183,1844,1869]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17a, f</figureCitation>
|
||
). Females of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB03BF8B3FEB5D406" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB03BF8B3FEB5D406" italics="true" pageId="22" pageNumber="173">Euphaea sanguinea</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from those of
|
||
<taxonomicName id="4FC64D46FFF3EC0EB79DF8DEFC9AD406" authorityName=": Selys" authorityYear="1859" box="[670,805,1879,1904]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB79DF8DEFC9AD406" box="[670,805,1879,1904]" italics="true" pageId="22" pageNumber="173">E. ochracea</emphasis>
|
||
</taxonomicName>
|
||
by the pattern of their abdominal yellow markings with much wider stripes on S
|
||
<quantity id="4F3E9B20FFF3EC0EB4ABF8FBFE4FD4E3" box="[424,496,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="15.24" metricValueMin="2.54" pageId="22" pageNumber="173" unit="in" value="3.5" valueMax="6.0" valueMin="1.0">1–6 in</quantity>
|
||
the former (
|
||
<figureCitation id="10FD2A40FFF3EC0EB77DF8FBFD60D4E2" box="[638,735,1916,1941]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">
|
||
Fig.
|
||
<quantity id="4F3E9B20FFF3EC0EB7B0F8FBFD60D4E2" box="[691,735,1916,1940]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="22" pageNumber="173" unit="g" value="17.0">17g</quantity>
|
||
</figureCitation>
|
||
), but narrower stripes on S
|
||
<quantity id="4F3E9B20FFF3EC0EB112F8FBFBE6D4E3" box="[1041,1113,1916,1941]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.778" metricValueMin="0.254" pageId="22" pageNumber="173" unit="in" value="4.0" valueMax="7.0" valueMin="1.0">1–7 in</quantity>
|
||
the latter species (
|
||
<figureCitation id="10FD2A40FFF3EC0EB02EF8FBFA31D4E3" box="[1325,1422,1916,1941]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="22" pageNumber="173">Fig. 17b</figureCitation>
|
||
), although this is to some extent an age-dependant character and as such should be treated with care. Although
|
||
<bibRefCitation id="EC574B34FFF3EC0EB055F826FF5ED4AA" author="Fraser" pageId="22" pageNumber="173" refString="Fraser, F. C. (1934) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. II. Taylor and Francis, London, xxiii + 398 pp., col. Pls. I - IV excl." type="book" year="1934">Fraser (1934)</bibRefCitation>
|
||
and Asahina (1986) mention a yellow stripe across the antefrons for
|
||
<taxonomicName id="4FC64D46FFF3EC0EB12FF843FABAD4AB" authorityName=": Selys" authorityYear="1859" box="[1068,1285,1988,2013]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF3EC0EB12FF843FABAD4AB" box="[1068,1285,1988,2013]" italics="true" pageId="22" pageNumber="173">Euphaea ochracea</emphasis>
|
||
</taxonomicName>
|
||
, at least the specimens from
|
||
<collectingCountry id="F0D17655FFF2EC0FB45BFF10FE08D3C6" box="[344,439,151,176]" name="Vietnam" pageId="23" pageNumber="174">Vietnam</collectingCountry>
|
||
we were able to inspect have this stripe interrupted, although not as widely as in
|
||
<taxonomicName id="4FC64D46FFF2EC0FB087FF1EFEB5D3A2" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB087FF1EFEB5D3A2" italics="true" pageId="23" pageNumber="174">E. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="10FD2A40FFF2EC0FB419FF3BFE30D3A3" box="[282,399,188,213]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="23" pageNumber="174">Fig. 17d, i</figureCitation>
|
||
). To some extent the width of the gap may also be age-dependant, but it appears that female
|
||
<taxonomicName id="4FC64D46FFF2EC0FB594FF58FEC6D38E" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[151,377,223,248]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB594FF58FEC6D38E" box="[151,377,223,248]" italics="true" pageId="23" pageNumber="174">Euphaea sanguinea</emphasis>
|
||
</taxonomicName>
|
||
generally has relatively less yellow marking on the frons. The structure of the prothorax appears to be similar for the females of both
|
||
<taxonomicName id="4FC64D46FFF2EC0FB78FFE83FCD5D26A" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[652,874,260,285]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB78FFE83FCD5D26A" box="[652,874,260,285]" italics="true" pageId="23" pageNumber="174">Euphaea sanguinea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4FC64D46FFF2EC0FB6A3FE82FB98D26B" authorityName=": Selys" authorityYear="1859" box="[928,1063,260,285]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB6A3FE82FB98D26B" box="[928,1063,260,285]" italics="true" pageId="23" pageNumber="174">E. ochracea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="887936C5FFF2EC0FB5C4FEA0FA80D1BB" blockId="23.[151,1436,151,789]" pageId="23" pageNumber="174">
|
||
Earlier,
|
||
<bibRefCitation id="EC574B34FFF2EC0FB420FEA0FE66D236" author="Kosterin" box="[291,473,295,320]" pageId="23" pageNumber="174" refString="Kosterin, O. E. (2016) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2016. International Dragonfly Fund Report. Journal of the International Dragonfly Fund, 98, 1 - 85." type="journal article" year="2016">Kosterin (2016)</bibRefCitation>
|
||
provided photos of damselflies identified as ‘
|
||
<taxonomicName id="4FC64D46FFF2EC0FB6F1FEA0FB77D236" authorityName=": Selys" authorityYear="1859" box="[1010,1224,295,320]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB6F1FEA0FB77D236" box="[1010,1224,295,320]" italics="true" pageId="23" pageNumber="174">Euphaea ochracea</emphasis>
|
||
</taxonomicName>
|
||
’ from
|
||
<collectingRegion id="4A02F827FFF2EC0FB01FFEA0FA24D236" box="[1308,1435,295,320]" country="Cambodia" name="Mondolkiri" pageId="23" pageNumber="174">Mondulkiri</collectingRegion>
|
||
Prov.,
|
||
<collectingCountry id="F0D17655FFF2EC0FB5DCFECBFEEAD213" box="[223,341,332,357]" name="Cambodia" pageId="23" pageNumber="174">Cambodia</collectingCountry>
|
||
, and pointed out that the five collected female specimens had "entirely black postclypeus and the pale marking on the anteclypeus broadly interrupted." This was erroneous wording. Proper wording should have been "entirely black clypeus and pale marking on the antefrons broadly interrupted" (Kosterin,
|
||
<emphasis id="BAB2EAD7FFF2EC0FB1BAFE12FAFED2DA" box="[1209,1345,405,428]" italics="true" pageId="23" pageNumber="174">pers. comm.</emphasis>
|
||
), which is in line with our own material from
|
||
<collectingCountry id="F0D17655FFF2EC0FB735FE30FD26D2A6" box="[566,665,439,464]" name="Vietnam" pageId="23" pageNumber="174">Vietnam</collectingCountry>
|
||
. He put “?ssp.” after the species name to indicate that those specimen could represent an undescribed subspecies, but refrained from nominating the Cambodian specimens as a new taxon. The abdomen pattern of distinctly red S2–6 displayed in the photos of Cambodian
|
||
<emphasis id="BAB2EAD7FFF2EC0FB19AFE78FA3FD16E" box="[1177,1408,511,536]" italics="true" pageId="23" pageNumber="174">
|
||
‘
|
||
<taxonomicName id="4FC64D46FFF2EC0FB19DFE78FAB8D16E" authorityName="Selys" authorityYear="1840" box="[1182,1287,511,536]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="genus">Euphaea</taxonomicName>
|
||
ochracea’
|
||
</emphasis>
|
||
is identical to that of
|
||
<taxonomicName id="4FC64D46FFF2EC0FB46FFDA2FE40D14A" authority="Phan & Kompier & Karube & Hayashi, 2018" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[364,511,549,572]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="sanguinea" status="sp. nov.">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB46FFDA2FE40D14A" box="[364,511,549,572]" italics="true" pageId="23" pageNumber="174">E. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A18157ACFFF2EC0FB705FDA3FDDFD14B" box="[518,608,548,573]" pageId="23" pageNumber="174" rank="species">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB705FDA3FDDFD14B" bold="true" box="[518,608,548,573]" pageId="23" pageNumber="174">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and the same holds true of the HW coloration (
|
||
<figureCitation id="10FD2A40FFF2EC0FB183FDA3FB66D14B" box="[1152,1241,548,573]" captionStart="FIGURE 17" captionStartId="25.[151,250,1366,1388]" captionTargetBox="[248,1338,210,1339]" captionTargetId="figure@25.[217,1369,193,1345]" captionTargetPageId="25" captionText="FIGURE 17. Euphaea spp. [17a–d] E. ochracea, Mang Den, Kon Tum & [17e], Penang, Malaysia; [17f–j] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/3096113/files/figure.png" pageId="23" pageNumber="174">Fig. 17f</figureCitation>
|
||
in this paper and Fig. 25, p.
|
||
<quantity id="4F3E9B20FFF2EC0FB417FDC0FEF1D116" box="[276,334,583,608]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="23" pageNumber="174" unit="in" value="25.0">25 in</quantity>
|
||
<bibRefCitation id="EC574B34FFF2EC0FB455FDC0FE43D116" author="Kosterin" box="[342,508,583,608]" pageId="23" pageNumber="174" refString="Kosterin, O. E. (2016) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2016. International Dragonfly Fund Report. Journal of the International Dragonfly Fund, 98, 1 - 85." type="journal article" year="2016">Kosterin 2016</bibRefCitation>
|
||
). Therefore, it can be safely assumed that
|
||
<emphasis id="BAB2EAD7FFF2EC0FB6DEFDC0FB7BD116" box="[989,1220,583,608]" italics="true" pageId="23" pageNumber="174">
|
||
‘
|
||
<taxonomicName id="4FC64D46FFF2EC0FB6E2FDC0FBF5D116" authorityName="Selys" authorityYear="1840" box="[993,1098,583,608]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="genus">Euphaea</taxonomicName>
|
||
ochracea’
|
||
</emphasis>
|
||
from
|
||
<collectingCountry id="F0D17655FFF2EC0FB009FDC0FAC2D116" box="[1290,1405,583,608]" name="Cambodia" pageId="23" pageNumber="174">Cambodia</collectingCountry>
|
||
in fact were
|
||
<taxonomicName id="4FC64D46FFF2EC0FB40BFDEAFE24D1F2" authority="Phan & Kompier & Karube & Hayashi, 2018" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[264,411,621,644]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="sanguinea" status="sp. nov.">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB40BFDEAFE24D1F2" box="[264,411,621,644]" italics="true" pageId="23" pageNumber="174">E. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A18157ACFFF2EC0FB4A0FDEBFE41D1F3" box="[419,510,620,645]" pageId="23" pageNumber="174" rank="species">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB4A0FDEBFE41D1F3" bold="true" box="[419,510,620,645]" pageId="23" pageNumber="174">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
. According to M. Hämäläinen (
|
||
<emphasis id="BAB2EAD7FFF2EC0FB668FDEAFC17D1F3" box="[875,936,620,645]" italics="true" pageId="23" pageNumber="174">in litt</emphasis>
|
||
.) the specimens from
|
||
<collectingRegion id="4A02F827FFF2EC0FB1A5FDEAFA9FD1F3" box="[1190,1312,621,645]" country="Vietnam" name="Lam Dong" pageId="23" pageNumber="174">Lam Dong</collectingRegion>
|
||
(Bao Loc) and
|
||
<collectingRegion id="4A02F827FFF2EC0FB5C9FD17FE84D1DE" box="[202,315,655,680]" country="Vietnam" name="Dong Nai" pageId="23" pageNumber="174">Dong Nai</collectingRegion>
|
||
(Cat Tien National Park) recorded as
|
||
<taxonomicName id="4FC64D46FFF2EC0FB7FFFD08FC6DD1DE" authorityName=": Selys" authorityYear="1859" box="[764,978,655,680]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB7FFFD08FC6DD1DE" box="[764,978,655,680]" italics="true" pageId="23" pageNumber="174">Euphaea ochracea</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EC574B34FFF2EC0FB107FD17FB5DD1DE" author="Do" box="[1028,1250,656,680]" pageId="23" pageNumber="174" refString="Do, M. C. & Dang, T. T. H. (2007) Checklist of dragonfly from Vietnam. Vietnam National University Publisher, Hanoi, 182 pp." type="book" year="2007">Do & Dang (2007)</bibRefCitation>
|
||
also refer to
|
||
<taxonomicName id="4FC64D46FFF2EC0FB087FD16FEB5D1BA" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB087FD16FEB5D1BA" italics="true" pageId="23" pageNumber="174">E. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
; Hämäläinen himself has also collected specimens of
|
||
<taxonomicName id="4FC64D46FFF2EC0FB66BFD33FBF9D1BA" authorityName="Phan & Kompier & Karube & Hayashi" authorityYear="2018" box="[872,1094,692,717]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB66BFD33FBF9D1BA" box="[872,1094,692,717]" italics="true" pageId="23" pageNumber="174">Euphaea sanguinea</emphasis>
|
||
</taxonomicName>
|
||
from these provinces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0DC654EFFF2EC0FB5C4FD50FEEED063" pageId="23" pageNumber="174" type="distribution">
|
||
<paragraph id="887936C5FFF2EC0FB5C4FD50FEEED063" blockId="23.[151,1436,151,789]" pageId="23" pageNumber="174">
|
||
<emphasis id="BAB2EAD7FFF2EC0FB5C4FD50FEDFD186" bold="true" box="[199,352,727,752]" pageId="23" pageNumber="174">Distribution.</emphasis>
|
||
<collectingCountry id="F0D17655FFF2EC0FB46FFD50FE74D186" box="[364,459,727,752]" name="Vietnam" pageId="23" pageNumber="174">Vietnam</collectingCountry>
|
||
(
|
||
<collectingRegion id="4A02F827FFF2EC0FB4DEFD50FDF8D186" box="[477,583,727,752]" country="Vietnam" name="Dak Lak" pageId="23" pageNumber="174">Dak Lak</collectingRegion>
|
||
,
|
||
<collectingRegion id="4A02F827FFF2EC0FB754FD5FFD6AD186" box="[599,725,728,752]" country="Vietnam" name="Lam Dong" pageId="23" pageNumber="174">Lam Dong</collectingRegion>
|
||
and
|
||
<collectingRegion id="4A02F827FFF2EC0FB617FD5FFC39D186" box="[788,902,727,752]" country="Vietnam" name="Dong Nai" pageId="23" pageNumber="174">Dong Nai</collectingRegion>
|
||
Provinces),
|
||
<collectingCountry id="F0D17655FFF2EC0FB11FFD50FB30D186" box="[1052,1167,727,752]" name="Cambodia" pageId="23" pageNumber="174">Cambodia</collectingCountry>
|
||
(
|
||
<collectingRegion id="4A02F827FFF2EC0FB1A1FD50FA2BD186" box="[1186,1428,727,752]" country="Cambodia" name="Mondolkiri" pageId="23" pageNumber="174">Mondulkiri Province</collectingRegion>
|
||
) (
|
||
<bibRefCitation id="EC574B34FFF2EC0FB59DFD7BFEFAD063" author="Kosterin" box="[158,325,764,789]" pageId="23" pageNumber="174" refString="Kosterin, O. E. (2016) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2016. International Dragonfly Fund Report. Journal of the International Dragonfly Fund, 98, 1 - 85." type="journal article" year="2016">Kosterin 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |