278 lines
33 KiB
XML
278 lines
33 KiB
XML
<document ID-CLB-Dataset="24300" ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="001178297187810C9CCCBE7793A178FF" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole typhlos Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="99FD33D4-9BC1-4595-9231-1A85822A7B35" docUuidSource="ZooBank" docVersion="7" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1732649601836" updateUser="ExternalLinkService">
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<mods:mods id="42E2F318FB97783E7EC82A01E82DC943" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B0727644217C783323C7559B44253FD6">
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<mods:title id="15B1CFFBD77F8CE21770BB2186AFAC3A">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:name id="9DEE8C81F5BD4CCF5E1C02B92CD4215C" type="personal">
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<mods:namePart id="6B9C7835B371C5D74AA730469604DB6A">Salata, Sebastian</mods:namePart>
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<mods:name id="37468C626EC78C87F4D06ADEC2EC1F81" type="personal">
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<mods:roleTerm id="3F87EDCDC4BFEE110D92AD5552E08EE1">Author</mods:roleTerm>
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<mods:namePart id="0FFC39CFFD70C386312C417F0C526D4C">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="D2F3E7145B246344207258350B076A71">text</mods:typeOfResource>
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<mods:titleInfo id="F235E5088B2443308775B0633A8D9D99">
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<mods:title id="52012AC365FFC14BBAF91311C7A9BB26">ZooKeys</mods:title>
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<mods:part id="BA32FF075366D17A09BBBBA1DBBFCFBB">
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<mods:date id="3A062A1856D66574D38886613EFA447B">2020</mods:date>
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<mods:detail id="03EE4E210137BE86C2DDDE68D6F9B0F4" type="volume">
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<mods:number id="8D9A7BB1C7F78BE9DAFF1F01907254D5">905</mods:number>
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<mods:extent id="5BEDDDFF54A85D449F1609FA043BB95B" unit="page">
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<mods:start id="3F4E212E08609AFD2EDD58C7921911CA">1</mods:start>
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<mods:end id="0070261F5EE5C177E27BE78CD8FD1443">235</mods:end>
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<mods:location id="61FB849BDAE0E5BA07C9779215316344">
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<mods:url id="CB5778FD90761695699DA32C8312E324">http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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<mods:classification id="C1D8D0E6A684A2771D20F6F9AC03DCDB">journal article</mods:classification>
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<mods:identifier id="BF59A48734A7CEF71930CE917CA28E39" type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier id="261F6A7AD657886F654BCCF2549E7CAF" type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<treatment id="001178297187810C9CCCBE7793A178FF" ID-GBIF-Taxon="161776994" LSID="urn:lsid:zoobank.org:act:99FD33D4-9BC1-4595-9231-1A85822A7B35" httpUri="http://treatment.plazi.org/id/001178297187810C9CCCBE7793A178FF" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="08F9624F1C5CE01DD01DD185DB2CF4F4" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="FF2F2338CF197A9BF501E9A5590CF9BF" pageId="0" pageNumber="1">
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<taxonomicName id="7FDD2CB73C5DB3E9D64FB08A144F9F74" ID-CoL="CMM2N" LSID="http://zoobank.org/99FD33D4-9BC1-4595-9231-1A85822A7B35" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole typhlos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlos" status="sp. nov.">Pheidole typhlos</taxonomicName>
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<taxonomicNameLabel id="CCD2A5D06FA5769644F4838786E253AA" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="0093AC282130C7DE907EB27C547314A5" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">Figs 20A-F</figureCitation>
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<figureCitation id="6630CC99BA3AD6D485696EB528D2C7EE" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85X</figureCitation>
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<figureCitation id="CA54AA65D5971BEED3E04D2864766C0E" captionStart="Figure 88" captionStartId="F88" captionText="Figure 88. Distribution. Pheidole petax Forel (A). P. podargea sp. nov. (B). P. praegrandis sp. nov. (C). P. ranohirensis sp. nov. (D). P. rugocephala sp. nov. (E). P. rugofitarata sp. nov. (F). P. scabrata Forel (G). P. typhlos sp. nov. (H). P. vatovavensis sp. nov. (I). P. voasara sp. nov. (J). P. vohemarensis sp. nov. (K). P. zavamanira sp. nov. (L)." figureDoi="10.3897/zookeys.905.39592.figure88" httpUri="https://binary.pensoft.net/fig/373135" pageId="0" pageNumber="1">, 88H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="ED3DB08B1B8129F520F88E3B0CCA35E7" pageId="0" pageNumber="1" type="type material">
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<paragraph id="744D0EF5A586132765FA42F92F6F98AF" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="79F1815E9123F74D6615080703BBAA48" pageId="0" pageNumber="1">
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<emphasis id="6D240BDDAFB6E1CB6EFE16F3C1354075" bold="true" pageId="0" pageNumber="1">
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<emphasis id="0A32AC4CB4A343DCF52535A7780E5CDC" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Antsiranana; Galoko chain, Mont Galoko;
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<geoCoordinate id="6A22067F1C3854864894CF3CCECF135A" degrees="13.58487" direction="south" orientation="latitude" precision="1" value="-13.58487">-13.58487</geoCoordinate>
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,
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<geoCoordinate id="0D6F4BC51BBB8C78ABA94FBEBD9624D6" degrees="48.71818" direction="east" orientation="longitude" precision="1" value="48.71818">48.71818</geoCoordinate>
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; alt. 520 m; 19 Feb 2013; B.L. Fisher et al. leg.; BLF30851, CASENT0302953 (CASC).
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<emphasis id="3168D7B48CEE14974ACBF6B02F117BA1" bold="true" pageId="0" pageNumber="1">
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<emphasis id="7C988E0A6DE553DB25A90A9FF80CEE09" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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.
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</emphasis>
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Madagascar. •1 w.; same data as for holotype; CASENT0923217 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="165249AA21CE56C4B8B9A9598690292F" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="60975587050C8CFE6F27BEDD2D1C99C2" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="D8245E831243DB717608BE497F882B77" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="CEB1E01F29AE3B1E8228AF3A82A85937" bold="true" italics="true" pageId="0" pageNumber="1">Antananarivo</emphasis>
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: •6s.; Galoko chain, Mont Galoko;
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<geoCoordinate id="AA45EF502173642587A20F8DDED8BC7F" degrees="13.58487" direction="south" orientation="latitude" precision="1" value="-13.58487">-13.58487</geoCoordinate>
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,
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<geoCoordinate id="67817A896185F3BFFA40FD5278F0C252" degrees="48.71818" direction="east" orientation="longitude" precision="1" value="48.71818">48.71818</geoCoordinate>
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; alt. 520 m; 16 Feb 2013; B.L. Fisher et al. leg.; CASENT0305030 (CASC). •1w.; Galoko chain, Mont Galoko;
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<geoCoordinate id="FB7D979B2CFC14DE6264D7B9D30833CB" degrees="13.5888" direction="south" orientation="latitude" precision="5" value="-13.5888">-13.5888</geoCoordinate>
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,
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<geoCoordinate id="6683BC6983FE12B09549F046C23CF346" degrees="48.72864" direction="east" orientation="longitude" precision="1" value="48.72864">48.72864</geoCoordinate>
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; alt. 980 m; 22 Feb 2013; B.L. Fisher et al. leg.; CASENT0304748 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="5065D6027408F88701FBCB1854A67B9E" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="6CCA4387252AC5F80F8F9362C243EF4B" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="13771183DBD27827D8252E1273C951A5" pageId="0" pageNumber="1">
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<emphasis id="E14512870DB89B68F1CE9535F61E3D7A" bold="true" pageId="0" pageNumber="1">
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<emphasis id="CE2FFA7A4D441C35958DAE9A2F97F4EC" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Eyes small and reduced; body yellow; head rectangular, anterior of eyes relatively straight, posterior of eyes slightly convex; sides of the head with sparse, short, erect pilosity; occipital lobes, genae, and posterior part of frons shiny and smooth, sometimes with indistinct, very sparse, longitudinal rugulae; anterior part of frons and malar area shiny, with thick, very sparse, short, and interrupted rugae, interspaces smooth; propodeal spines short, triangular, with acute top; pronotum smooth and shiny, sometimes on lateral sides with indistinct and sparse rugulae; mesonotum and propodeum shiny, with fine, sparse to moderately sparse rugoreticulation; katepisternum indistinctly foveolate; anepisternum with thick rugoreticulation; inner hypostomal teeth distinct, moderately high, narrow, triangular, with rounded apex; outer hypostomal teeth distinct, triangular, and with rounded tops directed outward; outer hypostomal teeth are slightly lower than inner hypostomal teeth.
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<emphasis id="3B13E7691F1F5F22FFA0FC834D6F7178" bold="true" pageId="0" pageNumber="1">
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<emphasis id="019C7A2F4D34F015F1712E1524951571" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Eyes small and reduced; body yellow; head and mesosoma shiny and smooth, lateral sides of frons with short, indistinct, longitudinal rugulae; propodeal spines short and triangular.
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</paragraph>
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</subSubSection>
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<subSubSection id="9C6310AB789A2B837A6151DE3B64935C" pageId="0" pageNumber="1" type="description">
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<paragraph id="6DA47E54B5E521AAE0E4ABF8200010B3" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="E30E937F11EA38FD819B1898A554E2FE" pageId="0" pageNumber="1">
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<emphasis id="B583DE93F6EFC242BE5F72FE1EFD8748" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="CBDB55F58CCCC620A377B037BA9D9D59" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 8): HL: 1.2-1.34 (1.28); HW: 0.96-1.07 (1.01); SL: 0.49-0.52 (0.5); EL: 0.08-0.12 (0.1); WL: 0.79-0.91 (0.84); PSL: 0.15-0.19 (0.16); MTL: 0.48-0.52 (0.51); PNW: 0.48-0.55 (0.51); PTW: 0.14-0.16 (0.15); PPW: 0.37-0.43 (0.4); CI: 75.1-82.0 (79.5); SI: 48.0-52.3 (49.8); PSLI: 11.9-14.3 (12.7); PPI: 36.3-39.9 (37.9); PNI: 47.4-55.3 (50.6); MTI: 46.9-52.9 (49.8).
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<emphasis id="580304905888D10ABD583C0DA35CC147" bold="true" pageId="0" pageNumber="1">
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<emphasis id="9DF86613925767CAB5BBAE16B299173A" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view elongate, anterior of eyes relatively straight, posterior of eyes slightly convex (Fig.
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<figureCitation id="7486DB66E32F1C27B2E01BD40F093E80" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces relatively flat; inner hypostomal teeth invisible. Antennal scrobes absent. Sides of the head with sparse, short, erect pilosity; whole head with dense, short, suberect to erect pilosity. Occipital lobes, genae, and posterior part of frons shiny and smooth, sometimes with indistinct, very sparse, longitudinal rugulae; anterior part of frons and malar area shiny, with thick, very sparse, short, and interrupted rugae, interspaces smooth. Clypeus shiny and smooth, with a few indistinct, longitudinal rugae on the lateral sides; median notch present, wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching the midlength of head; pilosity decumbent to erect (Fig.
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<figureCitation id="F7C682EB7D35EE592046243A41C62A47" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20B, D</figureCitation>
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). Inner hypostomal teeth distinct, moderately high, narrow, triangular, with rounded apex; outer hypostomal teeth distinct, triangular, and with rounded tops directed outward; outer hypostomal teeth are slightly lower than inner hypostomal teeth (Fig.
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<figureCitation id="15C7CB8ACCC2C5517C3F501DF71D0F95" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85X</figureCitation>
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).
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<emphasis id="95DA4C9C51CD4CFC02118A5DD705315F" bold="true" pageId="0" pageNumber="1">
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<emphasis id="FB66C8328BED77128C9D41CE9F98D4D6" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum convex, with indistinct tubercle-like projection; promesonotal groove absent; metanotal groove absent or very indistinct; propodeal spines short, triangular, with acute top; humeral area with wide and flat tubercles (Fig.
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<figureCitation id="2D4EC05245453015D9B63E2CF2031141" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D</figureCitation>
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). Pronotum smooth and shiny, sometimes on lateral sides with indistinct and sparse rugulae; mesonotum and propodeum shiny, with fine, sparse to moderately sparse rugoreticulation; katepisternum indistinctly foveolate; anepisternum with thick rugoreticulation. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation id="1B39B324F307978E6AA350A310B38001" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
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).
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<emphasis id="B254E52B546F2FAE9A76CFB923C40A96" bold="true" pageId="0" pageNumber="1">
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<emphasis id="535B9BFDF34E05E2272C3D9C76E735C9" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny; peduncle finely foveolate, without horizontal lobes on its basal part; node smooth, moderately high, with convex apex, in rear view node dorsoventrally concave; pilosity moderately dense and erect (Fig.
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<figureCitation id="A403CF1FAB99CB0256CC8F258A565242" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
|
|
).
|
|
<emphasis id="06059FF0E9882F1D15F2491ABE87EC3B" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="36775CD2E03E9A1EE97B1073D8A4AD14" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Finely foveolate and shiny; in dorsal view sides with moderately long, acute, and triangular projections; pilosity moderately long, moderately dense, and erect (Fig.
|
|
<figureCitation id="98276EF1950887C1514259B49C0EA2F2" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
|
|
).
|
|
<emphasis id="7428555E3D9F628F46239A9E3B6C8BBF" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="0BC8BB5B1A16B6E3D1B2C0C47202521E" italics="true" pageId="0" pageNumber="1">Gaster</emphasis>
|
|
.
|
|
</emphasis>
|
|
Smooth and shiny; pilosity dense, short, and erect (Fig.
|
|
<figureCitation id="AF90CB11A53DFC2A5A5DB6D6AC5D6434" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
|
|
).
|
|
<emphasis id="2F261907B0BFFFDB22E1719BC587ED62" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="DE0BE81DC43832E5A85754F884564542" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Yellow, malar area and lower part of frons dark yellow to bright brown (Fig.
|
|
<figureCitation id="19A210D7E103B785FE31C079328D1785" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
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).
|
|
</paragraph>
|
|
<caption id="EF636DB36D6DD21D548FFE2D10AA63F8" doi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1" start="Figure 20" startId="F20">
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|
<paragraph id="B5018A3E9E9112C603F63A3A5DCDE194" pageId="0" pageNumber="1">
|
|
<emphasis id="A2525255B1554AE31B7BAF0898528F01" bold="true" pageId="0" pageNumber="1">Figure 20.</emphasis>
|
|
<taxonomicName id="F9090CF22CA39479B4AFA1130E40F72C" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole typhlos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlos">
|
|
<emphasis id="24CC08014D07D6A59038856E3993432B" italics="true" pageId="0" pageNumber="1">Pheidole typhlos</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., full-face view (
|
|
<emphasis id="2C263BD27820BB1759BB60FEB7842A7D" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), profile (
|
|
<emphasis id="2AA3B06DE844A2D6DF72CFF3E4AC6669" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), and dorsal view (
|
|
<emphasis id="0790FEEEAD7C67DAEEE1B039AFCF0F3E" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
) of paratype minor worker (CASENT0923217) and full-face view (
|
|
<emphasis id="47543B2F351829B56BDE941394CFD016" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), profile (
|
|
<emphasis id="6ACB2EC2F97396229391457741D7147D" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
), and dorsal view (
|
|
<emphasis id="284A0C06AECC80F5E99F259979F84447" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
) of holotype major worker (CASENT0302953).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="DC9B7ABB1FE15F6F09DA3EC2727C9DE5" pageId="0" pageNumber="1">
|
|
<emphasis id="3401319975B87B980D958CD07A6DD5AA" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
|
Measurements (
|
|
<emphasis id="3BA45BF7F02FD1B4281C7FE26A0A5196" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 1): HL: 0.55; HW: 0.46; SL: 0.45; EL: 0.05; WL: 0.62; PSL: 0.09; MTL: 0.35; PNW: 0.32; PTW: 0.09; PPW: 0.14; CI: 83.9; SI: 97.4; PSLI: 16.4; PPI: 61.2; PNI: 69.8; MTI: 76.7.
|
|
<emphasis id="03D83D93673F0EEE59F7E3FB8E3A726D" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="BC3CE1EB470815D19EEF27A38232DC53" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|
.
|
|
</emphasis>
|
|
In full-face view rectangular, posterior and anterior of eyes slightly convex, occipital margin straight or indistinctly concave; occipital carina absent (Fig.
|
|
<figureCitation id="BA9927DF1B15C235C71172D8AFD91A2E" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20A</figureCitation>
|
|
). Pilosity moderately dense, short, subdecumbent to suberect. Whole head shiny and smooth, only lateral sides of frons with short, indistinct, longitudinal rugulae; antennal sockets with sparse carinae curved outward. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, reaching posterior head margin; pilosity suberect to erect (Fig.
|
|
<figureCitation id="2DF5256E2771639F3F071ECDFDA54876" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20A, C</figureCitation>
|
|
).
|
|
<emphasis id="574FB2D1006D8A526895E64E86FDAD42" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="01F0E7478C47508CC81E95CC1A5044BC" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
|
.
|
|
</emphasis>
|
|
In lateral view, promesonotum box-like; promesonotal groove absent; metanotal groove distinct; propodeal spines short and triangular, apex acute (Fig.
|
|
<figureCitation id="9D53754C5C033E99386319559E815245" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C</figureCitation>
|
|
). Sculpture smooth and shiny, sometimes lateral sides with indistinct, irregular, and sparse rugae. Pilosity moderately sparse, short, and erect (Fig.
|
|
<figureCitation id="CAF51F22D32245D289EC86156F3FF7EB" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C, E</figureCitation>
|
|
).
|
|
<emphasis id="6AADEFD1281513A96FDD2411367EAF0C" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="304D8008E1952B4160108C3BC7233E3F" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Peduncle short with ventral face slightly convex; node globular; with few short, erect setae (Fig.
|
|
<figureCitation id="95AB51911735C96018BDA61F038571DD" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C</figureCitation>
|
|
,
|
|
<figureCitation id="8C5E373A65DD9395D210AFB3E87B6DBC" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20E</figureCitation>
|
|
).
|
|
<emphasis id="AAC2D2613B0778E4BE6D81186BF5E807" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="B0136A8CD3E8F76A604EED53FC66FE41" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Short, low, and convex; with few short, erect setae (Fig.
|
|
<figureCitation id="23B6C6B20FDF2FB568E7784714819EC7" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C, E</figureCitation>
|
|
).
|
|
<emphasis id="83FD3B2FFFCDCBBF470E9981850EB281" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="F0FED274EE44C4E04A136A93CCCF4B82" italics="true" pageId="0" pageNumber="1">Gaster</emphasis>
|
|
.
|
|
</emphasis>
|
|
Pilosity moderately dense, erect, and short (Fig.
|
|
<figureCitation id="804251F5769B810D4074EF1914C51EC5" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C, E</figureCitation>
|
|
).
|
|
<emphasis id="6E73DFEF23599542C6AF1BA02D477855" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="344639FCF694B7F40E8E600636BDAA81" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Unicolourous, yellow (Fig.
|
|
<figureCitation id="1C41BF90312413DAC3B8C11C76F6B1F5" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="57260EDCA7CC3879D6607C8A3A6269C9" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="B51C95ACF7C05B35615AC94B86A733ED" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="19E6433FF62778DC94F48643D2636B88" pageId="0" pageNumber="1">Greek for blind [τυφλός], in reference to the reduced eyes.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7AB67B33C8732CEB8C5B8FBEEC11501F" pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph id="B47184F42DC1C0FE376903DCB853599E" pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph id="D17BF65DA39C0C310F18F54076300EAA" pageId="0" pageNumber="1">The species was collected at elevation between 520-980 m, in rainforest, in montane forest. Nesting preferences unknown.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6654036E3AB6375F1F11EA79C5F387C2" pageId="0" pageNumber="1" type="comments">
|
|
<paragraph id="84993043229760810AFA0FFFC5C3C31B" pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph id="64449DBB2D4C64AB1A8931CD3524CE43" pageId="0" pageNumber="1">
|
|
This species is most similar to
|
|
<taxonomicName id="6B63BCF66389FADF1CD9E7F3F776CD97" lsidName="P. parviocula" pageId="0" pageNumber="1" rank="species" species="parviocula">
|
|
<emphasis id="4B390C72CE7ACD50037668D8B6ED93F2" italics="true" pageId="0" pageNumber="1">P. parviocula</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis id="7C30013BCA95D414A6CE6A573B303912" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="37D61E6D5EE42470BD1225D54D0FFC82" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName id="93C125233A5560B93F774BDE4CFF7578" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole typhlos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlos">
|
|
<emphasis id="E60458AAB0C121858EEC27233E0E0177" italics="true" pageId="0" pageNumber="1">Pheidole typhlos</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName id="F394E9997192266C994755B3CB0AAA94" lsidName="P. parviocula" pageId="0" pageNumber="1" rank="species" species="parviocula">
|
|
<emphasis id="FFDF597BA4BF96C7D3D173DBDC3CA3D7" italics="true" pageId="0" pageNumber="1">P. parviocula</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by sparse and short pilosity on sides of the head, presence of distinct, triangular inner hypostomal teeth and presence of short, triangular propodeal spines.
|
|
<emphasis id="F671488012C9962DA0BB00EDC5B5F43D" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="8367BCBED1A60C60F0973325E1FD474B" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName id="230F17F54531399EA8123D4BC9833CB8" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole typhlos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlos">
|
|
<emphasis id="049F0D9232F9D1541EB07CF341F24098" italics="true" pageId="0" pageNumber="1">Pheidole typhlos</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName id="833FFDBBBCFBB933F7980E683F37B23E" lsidName="P. parviocula" pageId="0" pageNumber="1" rank="species" species="parviocula">
|
|
<emphasis id="B025A22BF8E3878B5E2F76E8420C1E19" italics="true" pageId="0" pageNumber="1">P. parviocula</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by box-like promesonotum, absence of promesonotal groove and presence of short and triangular propodeal spines.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |