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<document id="18461FDE440B300194E1AC38D2C62D9D" ID-CLB-Dataset="30255" ID-DOI="10.11646/zootaxa.4410.1.4" ID-GBIF-Dataset="f9d268e1-262b-4d9a-a97f-ffd1495e9eae" ID-ISSN="1175-5326" ID-Zenodo-Dep="1221257" ID-ZooBank="0944C1F0-1405-43E0-80B7-03438A19F334" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524145257255" checkinUser="plazi" docAuthor="Gąsiorek, Piotr &amp; Degma, Peter" docDate="2018" docId="001087910D06FF9AD399FE75FB44F80C" docLanguage="en" docName="zootaxa.4410.1.4.pdf" docOrigin="Zootaxa 4410 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bryodelphax instabilis Gąsiorek &amp; Degma 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="85" masterDocId="FC29FFE90D04FF92D30EFFD8FF9AFFEB" masterDocTitle="Three Echiniscidae species (Tardigrada: Heterotardigrada) new to the Polish fauna, with the description of a new gonochoristic Bryodelphax Thulin, 1928" masterLastPageNumber="96" masterPageNumber="77" pageNumber="79" updateTime="1698537978881" updateUser="ExternalLinkService">
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<mods:title id="49BCE65D29A7C301561319BCF9B9B7AD">Three Echiniscidae species (Tardigrada: Heterotardigrada) new to the Polish fauna, with the description of a new gonochoristic Bryodelphax Thulin, 1928</mods:title>
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<mods:namePart id="64AB588B4F2C2D6B537496FFF883E933">Gąsiorek, Piotr</mods:namePart>
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<mods:namePart id="A2CB222155340D9D6E00E1BDF719D102">Degma, Peter</mods:namePart>
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<mods:title id="293EEAD300884769784DC03C3B4BC658">Zootaxa</mods:title>
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<mods:date id="0C406307E835B34DA3A81F4C645B23B2">2018</mods:date>
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<mods:number id="3EED7C23AF3E5B08C4ACE5E24D601FAD">4410</mods:number>
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<treatment id="001087910D06FF9AD399FE75FB44F80C" ID-DOI="http://doi.org/10.5281/zenodo.5998748" ID-GBIF-Taxon="143605415" ID-Zenodo-Dep="5998748" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C" httpUri="http://treatment.plazi.org/id/001087910D06FF9AD399FE75FB44F80C" lastPageId="8" lastPageNumber="85" pageId="2" pageNumber="79">
<subSubSection id="C0A3650C0D06FF90D399FE75FE3AFE03" pageId="2" pageNumber="79" type="nomenclature">
<paragraph id="880636870D06FF90D399FE75FD90FE2C" blockId="2.[151,522,429,488]" box="[151,522,429,455]" pageId="2" pageNumber="79">
<heading id="D34E81EB0D06FF90D399FE75FD90FE2C" bold="true" box="[151,522,429,455]" fontSize="11" level="1" pageId="2" pageNumber="79" reason="1">
<emphasis id="BACDEA950D06FF90D399FE75FD90FE2C" bold="true" box="[151,522,429,455]" pageId="2" pageNumber="79">
<taxonomicName id="4FB94D040D06FF90D399FE75FE3CFE2C" ID-CoL="358bf5e4-073b-4ef9-ad5e-7f59779e5d57" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[151,422,429,455]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="79" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D06FF90D399FE75FE3CFE2C" bold="true" box="[151,422,429,455]" italics="true" pageId="2" pageNumber="79">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D06FF90D2A3FE75FD90FE2C" box="[429,522,429,455]" pageId="2" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="880636870D06FF90D399FE17FE3AFE03" blockId="2.[151,522,429,488]" box="[151,416,463,488]" pageId="2" pageNumber="79">
(
<figureCitation id="10822A020D06FF90D391FE17FE8BFE03" box="[159,273,463,488]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–9" captionStart-2="FIGURES10–12" captionStart-3="FIGURES13–16" captionStart-4="FIGURES 17–18" captionStart-5="FIGURES 19–20" captionStart-6="FIGURES21–23" captionStartId-0="4.[151,264,1781,1803]" captionStartId-1="5.[151,264,1301,1323]" captionStartId-2="6.[151,264,1846,1868]" captionStartId-3="7.[151,264,1209,1231]" captionStartId-4="7.[151,264,1936,1958]" captionStartId-5="8.[151,264,756,778]" captionStartId-6="8.[151,264,1721,1743]" captionTargetBox-0="[151,1436,195,1757]" captionTargetBox-1="[151,1436,193,1280]" captionTargetBox-2="[151,1436,193,1825]" captionTargetBox-3="[151,1436,193,1188]" captionTargetBox-4="[151,1436,1372,1913]" captionTargetBox-5="[151,1436,193,734]" captionTargetBox-6="[151,1436,873,1700]" captionTargetId-0="figure@4.[151,1436,195,1757]" captionTargetId-1="figure@5.[151,1436,193,1280]" captionTargetId-2="figure@6.[151,1436,193,1825]" captionTargetId-3="figure@7.[151,1436,193,1188]" captionTargetId-4="figure@7.[151,1436,1372,1913]" captionTargetId-5="figure@8.[151,1436,193,734]" captionTargetId-6="figure@8.[151,1436,858,1700]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="8" captionText-0="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." captionText-1="FIGURES 6–9. Bryodelphax instabilis sp. nov., habitus (SEM):6—female (paratype, dorsal view); 7—male (paratype,lateral view); 8—female (paratype, ventral view); 9—male (paratype, ventral view).Scale bars in µm." captionText-2="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." captionText-3="FIGURES13–16.Bryodelphax instabilissp.nov., dorsalplates (SEM):13—the scapular plate (paratype, male);14—posterior part of the median 2—median 3 (paratype, male), note almost complete absence of true pores; 15—the scapular plate (Slovakian female); 16—posterior part of the median 1—median 3 (Slovakian female), note numerous large true pores. Scale bars in µm." captionText-4="FIGURES 17–18. Bryodelphax instabilis sp. nov., male cephalic appendages (paratype, SEM): 17—cirrus A with clava, the arrowhead indicates the subterminal constriction; 18—enlarged clava. Scale bars inµm." captionText-5="FIGURES 19–20. Bryodelphax instabilis sp. nov., claws (SEM): 19—claws III; 20—claws IV with dentate collar. Scale bars in µm." captionText-6="FIGURES21–23. A semi-schematic illustration of the ontogenetic variability inthe ventral armature in Bryodelphax instabilis sp. nov.: 21—adult (mature female); 22—juvenile (second instar); 23—larva (first instar)." httpUri-0="https://zenodo.org/record/1221259/files/figure.png" httpUri-1="https://zenodo.org/record/1221261/files/figure.png" httpUri-2="https://zenodo.org/record/1221263/files/figure.png" httpUri-3="https://zenodo.org/record/1221265/files/figure.png" httpUri-4="https://zenodo.org/record/1221267/files/figure.png" httpUri-5="https://zenodo.org/record/1221269/files/figure.png" httpUri-6="https://zenodo.org/record/1221271/files/figure.png" pageId="2" pageNumber="79">Figs 1–23</figureCitation>
,
<tableCitation id="C53B033C0D06FF90D213FE17FE0DFE03" box="[285,407,463,488]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStart-2="TABLE 5" captionStartId-0="10.[151,239,1070,1093]" captionStartId-1="11.[151,239,152,175]" captionStartId-2="11.[151,239,1090,1113]" captionText-0="TABLE 3. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature females of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/ structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD— standard deviation)." captionText-1="TABLE 4. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature males and one larva of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD—standard deviation;?—unmeasurable structure)." captionText-2="TABLE 5. Statistical comparisons of mean pseudopore density per 100 µm 2 of selected dorsal plates between sexes of Bryodelphaxinstabilis sp. nov. (SD—standard deviation). Thep-values for the Welch t-test are provided and statistically significant differences are marked in bold (p&lt;0.050). 14 females and 8 males from the type series were compared." pageId="2" pageNumber="79">Tables 3–5</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C0A3650C0D06FF90D399FDC0FEA5FDBE" pageId="2" pageNumber="79" type="description">
<paragraph id="880636870D06FF90D399FDC0FEA5FDBE" blockId="2.[151,1437,535,2012]" pageId="2" pageNumber="79">
<materialsCitation id="38D13CDA0D06FF90D399FDC0FEA5FDBE" ID-GBIF-Occurrence="1838119989" country="Poland" elevation="595" latitude="49.404724" location="Pieniny Mts." longLatPrecision="18" longitude="20.547777" pageId="2" pageNumber="79" specimenCount="1">
<emphasis id="BACDEA950D06FF90D399FDC0FEA1FDDB" bold="true" box="[151,315,535,560]" italics="true" pageId="2" pageNumber="79">Locus typicus.</emphasis>
Moss on carbonate bedrock from Homole Ravine in the
<location id="8D66605C0D06FF90D0BBFDCFFBDFFDC4" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C:8D66605C0D06FF90D0BBFDCFFBDFFDC4" box="[949,1093,535,560]" country="Poland" latitude="49.404724" longLatPrecision="18" longitude="20.547777" name="Pieniny Mts." pageId="2" pageNumber="79">Pieniny Mts.</location>
(
<geoCoordinate id="ED8D50400D06FF90D75DFDC0FB43FDDB" box="[1107,1241,536,560]" direction="north" orientation="latitude" pageId="2" pageNumber="79" precision="15" value="49.404724">49°24'17''N</geoCoordinate>
,
<geoCoordinate id="ED8D50400D06FF90D7EAFDCFFAFDFDDB" box="[1252,1383,535,560]" direction="east" orientation="longitude" pageId="2" pageNumber="79" precision="15" value="20.547777">20°32'52''E</geoCoordinate>
;
<elevation id="0394D1B40D06FF90D67CFDCFFF4CFDBE" metricMagnitude="2" metricUnit="m" metricValue="5.95" pageId="2" pageNumber="79" unit="m" value="595.0">
<quantity id="4F419B620D06FF90D67CFDCFFF37FDBF" metricMagnitude="2" metricUnit="m" metricValue="5.95" pageId="2" pageNumber="79" unit="m" value="595.0">595 m</quantity>
asl
</elevation>
),
<collectingCountry id="F0AE76170D06FF90D3E4FDE4FEA1FDBE" box="[234,315,572,597]" name="Poland" pageId="2" pageNumber="79">Poland</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0A3650C0D06FF91D3C9FDBAFEECFA90" lastPageId="3" lastPageNumber="80" pageId="2" pageNumber="79" type="materials_examined">
<paragraph id="880636870D06FF90D3C9FDBAFE3BFC70" blockId="2.[151,1437,535,2012]" pageId="2" pageNumber="79">
<emphasis id="BACDEA950D06FF90D3C9FDBAFEEFFD90" bold="true" box="[199,373,610,635]" pageId="2" pageNumber="79">Type material.</emphasis>
Holotype (mature female, slide no. PL.189.19), coll. Maciej Barczyk,
<date id="FC0710470D06FF90D79FFDBAFAACFD90" box="[1169,1334,607,635]" pageId="2" pageNumber="79" value="2016-05-25">25th May 2016</date>
, allotype (mature male, slide no. PL.189.01), 30 paratypes (21 mature females, 7 males, one juvenile, and one larva; slides PL.189.02–32) and additional 15 paratypes (both juveniles and adults) on SEM stubs. Found together with numerous
<taxonomicName id="4FB94D040D06FF90D21EFD16FD31FD0C" authority="Thulin, 1928" authorityName="Thulin" authorityYear="1928" box="[272,683,718,743]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="79" phylum="Tardigrada" rank="species" species="parvulus">
<emphasis id="BACDEA950D06FF90D21EFD16FD91FD0C" box="[272,523,718,743]" italics="true" pageId="2" pageNumber="79">Bryodelphax parvulus</emphasis>
<bibRefCitation id="EC284B760D06FF90D118FD16FD31FD0C" box="[534,683,718,743]" pageId="2" pageNumber="79" refId="ref11774" refString="Thulin, G. (1928) Uber die Phylogenie und das System der Tardigraden. Hereditas, 11, 207 - 266. https: // doi. org / 10.1111 / j. 1601 - 5223.1928. tb 02488. x" type="journal article">Thulin, 1928</bibRefCitation>
</taxonomicName>
(
<figureCitation id="10822A020D06FF90D1B3FD16FC8EFD0C" box="[701,788,718,743]" captionStart="FIGURES 24–27" captionStartId="9.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@9.[151,1436,193,1825]" captionTargetPageId="9" captionText="FIGURES 24–27. Other Polish members of the genus Bryodelphax Thulin, 1928, and the comparison of sculpture of dorsal plates (PCM): 24—B. parvulus Thulin, 1928 (female, dorsolateral view); 25—B. tatrensis (węglarska, 1959) (male, dorsal view), note lack of the scapular plate faceting, the arrowhead indicates dentate collar on legs IV, and empty arrowheads point enlarged cephalic papillae; B. tatrensis, sexual dimorphism in the pore size: 26—female; 27—male.Note numerous true pores distributedon all plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221273/files/figure.png" pageId="2" pageNumber="79">Fig. 24</figureCitation>
),
<taxonomicName id="4FB94D040D06FF90D022FD16FA8CFD0C" authority="(Ehrenberg, 1853)" authorityName="Ehrenberg" authorityYear="1853" baseAuthorityName="Ehrenberg" baseAuthorityYear="1853" box="[812,1302,718,743]" class="Heterotardigrada" family="Echiniscidae" genus="Pseudechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="79" phylum="Tardigrada" rank="species" species="suillus">
<emphasis id="BACDEA950D06FF90D022FD16FBA3FD0C" box="[812,1081,718,743]" italics="true" pageId="2" pageNumber="79">Pseudechiniscus suillus</emphasis>
(
<bibRefCitation id="EC284B760D06FF90D745FD16FA94FD0C" box="[1099,1294,718,743]" pageId="2" pageNumber="79" refId="ref9857" refString="Ehrenberg, C. G. (1853) Diagnoses novarum formarum. Verhandlungen der Koniglich Preussische Akademie der Wissenschaften zu Berlin, 8, 526 - 533." type="journal article">Ehrenberg, 1853</bibRefCitation>
)
</taxonomicName>
, and single specimens of
<taxonomicName id="4FB94D040D06FF90D23FFD2AFCD7FCE0" authority="(Scourfield, 1897)" authorityName="Scourfield" authorityYear="1897" baseAuthorityName="Scourfield" baseAuthorityYear="1897" box="[305,845,754,779]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="79" phylum="Tardigrada" rank="species" species="spitsbergensis">
<emphasis id="BACDEA950D06FF90D23FFD2AFDEFFCE0" box="[305,629,754,779]" italics="true" pageId="2" pageNumber="79">Testechiniscus spitsbergensis</emphasis>
(
<bibRefCitation id="EC284B760D06FF90D18AFD2AFCDFFCE0" box="[644,837,754,779]" pageId="2" pageNumber="79" refId="ref11598" refString="Scourfield, D. J. (1897) Contributions to the non-marine fauna of Spitsbergen. Part I. Preliminary notes, and reports on the Rhizopoda, Tardigrada etc. Proceedings of the Zoological Society of London, 65 (3), 784 - 792." type="journal article">Scourfield, 1897</bibRefCitation>
)
</taxonomicName>
. All from the same moss sample. Holotype and most of paratypes deposited in the Institute of Zoology and Biomedical Research, Jagiellonian University, Kraków, Poland, two paratypes (slides PL.189.10–11) deposited in the Department of Zoology, Comenius University in Bratislava, Slovakia, and two paratypes (slides PL.189.22–23) deposited in the Zoological Museum, University of Copenhagen, Denmark.
</paragraph>
<paragraph id="880636870D06FF90D3C9FC7EFBC7FB2A" blockId="2.[151,1437,535,2012]" pageId="2" pageNumber="79">
<materialsCitation id="38D13CDA0D06FF90D3C9FC7EFBC0FB2A" ID-GBIF-Occurrence="1838119990" collectingDate="2003-07-26" collectingDateMax="2016-09-24" collectingDateMin="2003-07-26" collectorName="Two &amp; Chuda Alp &amp; Piotr Gasiorek &amp; Peter Degma" country="Slovakia" county="Branisko Mts." elevation="1036" latitude="48.995" location="Tatra Mts." longLatPrecision="18" longitude="20.866388" municipality="Rajtopiky" pageId="2" pageNumber="79" specimenCount="278" specimenCount-female="43" specimenCount-juvenile="3" specimenCount-male="44" stateProvince="Eastern" typeStatus="holotype">
<emphasis id="BACDEA950D06FF90D3C9FC7EFDA3FC54" bold="true" box="[199,569,934,959]" pageId="2" pageNumber="79">Additional material examined.</emphasis>
<collectorName id="254C53510D06FF90D14CFC70FDEEFC54" box="[578,628,936,959]" pageId="2" pageNumber="79">Two</collectorName>
specimens in a moss sample from
<collectorName id="254C53510D06FF90D703FC7EFB10FC54" box="[1037,1162,934,959]" pageId="2" pageNumber="79">Chuda Alp</collectorName>
in the
<location id="8D66605C0D06FF90D7D1FC70FACFFC54" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C:8D66605C0D06FF90D7D1FC70FACFFC54" box="[1247,1365,936,959]" country="Slovakia" county="Branisko Mts." latitude="48.995" longLatPrecision="18" longitude="20.866388" municipality="Rajtopiky" name="Tatra Mts." pageId="2" pageNumber="79" stateProvince="Eastern">Tatra Mts.</location>
(slide PL.258.06), coll.
<collectorName id="254C53510D06FF90D251FC15FD9CFC0D" box="[351,518,973,998]" pageId="2" pageNumber="79">Piotr Gąsiorek</collectorName>
,
<collectingDate id="EC43E9AF0D06FF90D11DFC16FC9AFC0D" box="[531,768,970,998]" pageId="2" pageNumber="79" value="2016-09-24">
<date id="FC0710470D06FF90D11DFC16FC9AFC0D" box="[531,768,970,998]" pageId="2" pageNumber="79" value="2016-09-24">24th September 2016</date>
</collectingDate>
, deposited together with
<typeStatus id="570288250D06FF90D72FFC16FBCBFC0D" box="[1057,1105,974,998]" pageId="2" pageNumber="79">type</typeStatus>
material.
<specimenCount id="9EBFFD0E0D06FF90D7C8FC16FAF4FC0D" box="[1222,1390,973,998]" pageId="2" pageNumber="79" type="generic">128 specimens</specimenCount>
(
<specimenCount id="9EBFFD0E0D06FF90D671FC16FF41FBE1" pageId="2" pageNumber="79" type="male">44 males</specimenCount>
,
<specimenCount id="9EBFFD0E0D06FF90D3E8FC2AFEF8FBE1" box="[230,354,1009,1034]" pageId="2" pageNumber="79" type="female">43 females</specimenCount>
,
<specimenCount id="9EBFFD0E0D06FF90D263FC2AFE70FBE1" box="[365,490,1009,1034]" pageId="2" pageNumber="79" type="juvenile">3 juveniles</specimenCount>
, and
<specimenCount id="9EBFFD0E0D06FF90D12AFC2AFD21FBE1" box="[548,699,1009,1034]" pageId="2" pageNumber="79" type="generic">38 specimens</specimenCount>
of unidentified sex due to unsuitable orientation of a specimen on a slide; slides 390/7–10, 390/18, 390/20, 390/23, 390/25, 390/27–30, and 390/32; deposited in the
<location id="8D66605C0D06FF90D7F6FBCEFF6DFBB9" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C:8D66605C0D06FF90D7F6FBCEFF6DFBB9" country="Slovakia" county="Branisko Mts." latitude="48.995" longLatPrecision="18" longitude="20.866388" municipality="Rajtopiky" name="Department of Zoology" pageId="2" pageNumber="79" stateProvince="Eastern">Department of Zoology</location>
,
<location id="8D66605C0D06FF90D20BFBE1FD0EFBB9" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C:8D66605C0D06FF90D20BFBE1FD0EFBB9" box="[261,660,1081,1106]" country="Slovakia" county="Branisko Mts." latitude="48.995" longLatPrecision="18" longitude="20.866388" municipality="Rajtopiky" name="Comenius University in Bratislava" pageId="2" pageNumber="79" stateProvince="Eastern">Comenius University in Bratislava</location>
, except for
<specimenCount id="9EBFFD0E0D06FF90D011FBE2FC20FBB9" box="[799,954,1081,1106]" pageId="2" pageNumber="79" type="generic">13 specimens</specimenCount>
deposited in the
<location id="8D66605C0D06FF90D78AFBE1FAF0FBB9" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C:8D66605C0D06FF90D78AFBE1FAF0FBB9" box="[1156,1386,1081,1106]" country="Slovakia" county="Branisko Mts." latitude="48.995" longLatPrecision="18" longitude="20.866388" municipality="Rajtopiky" name="Institute of Zoology" pageId="2" pageNumber="79" stateProvince="Eastern">Institute of Zoology</location>
and
<location id="8D66605C0D06FF90D399FB85FE16FB9D" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C:8D66605C0D06FF90D399FB85FE16FB9D" box="[151,396,1117,1142]" country="Slovakia" county="Branisko Mts." latitude="48.995" longLatPrecision="18" longitude="20.866388" municipality="Rajtopiky" name="Biomedical Research" pageId="2" pageNumber="79" stateProvince="Eastern">Biomedical Research</location>
,
<location id="8D66605C0D06FF90D294FB85FD39FB9D" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C:8D66605C0D06FF90D294FB85FD39FB9D" box="[410,675,1117,1142]" country="Slovakia" county="Branisko Mts." latitude="48.995" longLatPrecision="18" longitude="20.866388" municipality="Rajtopiky" name="Jagiellonian University" pageId="2" pageNumber="79" stateProvince="Eastern">Jagiellonian University</location>
,
<location id="8D66605C0D06FF90D1BFFB85FC91FB9D" LSID="urn:lsid:plazi:treatment:001087910D06FF9AD399FE75FB44F80C:8D66605C0D06FF90D1BFFB85FC91FB9D" box="[689,779,1117,1142]" country="Slovakia" county="Branisko Mts." latitude="48.995" longLatPrecision="18" longitude="20.866388" municipality="Rajtopiky" name="Krakow" pageId="2" pageNumber="79" stateProvince="Eastern">Kraków</location>
<emphasis id="BACDEA950D06FF90D005FB85FCBDFB9D" bold="true" box="[779,807,1117,1142]" pageId="2" pageNumber="79">—</emphasis>
slide 390/18) and
<specimenCount id="9EBFFD0E0D06FF90D0F4FB86FB1DFB9D" box="[1018,1159,1117,1142]" pageId="2" pageNumber="79" type="generic">9 specimens</specimenCount>
used for SEM analyses (not deposited) in a moss sample on a rock from
<collectingMunicipality id="6862ACFD0D06FF90D1C0FB59FCA6FB71" box="[718,828,1153,1178]" pageId="2" pageNumber="79">Rajtopíky</collectingMunicipality>
hill, the
<collectingCounty id="61674E0B0D06FF90D0ABFB59FBDCFB72" box="[933,1094,1153,1178]" pageId="2" pageNumber="79">Branisko Mts.</collectingCounty>
(
<geoCoordinate id="ED8D50400D06FF90D759FB59FB47FB71" box="[1111,1245,1153,1178]" direction="north" orientation="latitude" pageId="2" pageNumber="79" precision="15" value="48.995">48°59'42''N</geoCoordinate>
,
<geoCoordinate id="ED8D50400D06FF90D7E4FB59FAF7FB71" box="[1258,1389,1153,1178]" direction="east" orientation="longitude" pageId="2" pageNumber="79" precision="15" value="20.866388">20°51'59''E</geoCoordinate>
;
<emphasis id="BACDEA950D06FF90D674FB5AFA0EFB72" box="[1402,1428,1154,1177]" italics="true" pageId="2" pageNumber="79">ca</emphasis>
.
<elevation id="0394D1B40D06FF90D399FB70FE88FB2A" box="[151,274,1192,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.036" pageId="2" pageNumber="79" unit="m" value="1036.0">
<quantity id="4F419B620D06FF90D399FB70FF76FB2B" box="[151,236,1192,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.036" pageId="2" pageNumber="79" unit="m" value="1036.0">1036 m</quantity>
asl
</elevation>
<emphasis id="BACDEA950D06FF90D21CFB70FEB4FB2A" bold="true" box="[274,302,1192,1217]" pageId="2" pageNumber="79">—</emphasis>
on the hill top area),
<collectingRegion id="4A7DF8650D06FF90D119FB71FDF1FB2B" box="[535,619,1193,1216]" country="American Samoa" name="Eastern" pageId="2" pageNumber="79">Eastern</collectingRegion>
<collectingCountry id="F0AE76170D06FF90D17CFB70FD4DFB2A" box="[626,727,1192,1217]" name="Slovakia" pageId="2" pageNumber="79">Slovakia</collectingCountry>
, coll.
<collectorName id="254C53510D06FF90D014FB71FC34FB2A" box="[794,942,1193,1217]" pageId="2" pageNumber="79">Peter Degma</collectorName>
,
<collectingDate id="EC43E9AF0D06FF90D0B7FB70FBC0FB2A" box="[953,1114,1188,1217]" pageId="2" pageNumber="79" value="2003-07-26">
<date id="FC0710470D06FF90D0B7FB70FBC0FB2A" box="[953,1114,1188,1217]" pageId="2" pageNumber="79" value="2003-07-26">26th July 2003</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="880636870D06FF90D3C9FB14FD62FB0E" blockId="2.[151,1437,535,2012]" box="[199,760,1228,1253]" pageId="2" pageNumber="79">
Short description of
<typeStatus id="570288250D06FF90D2A2FB14FD94FB0E" box="[428,526,1228,1253]" pageId="2" pageNumber="79" type="holotype">holotype</typeStatus>
(
<figureCitation id="10822A020D06FF90D113FB14FDFEFB0E" box="[541,612,1228,1253]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="2" pageNumber="79">Figs 1</figureCitation>
,
<figureCitation id="10822A020D06FF90D17CFB15FD0BFB0E" box="[626,657,1229,1253]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="2" pageNumber="79">10</figureCitation>
;
<tableCitation id="C53B033C0D06FF90D193FB14FD75FB0E" box="[669,751,1228,1253]" captionStart="TABLE 3" captionStartId="10.[151,239,1070,1093]" captionText="TABLE 3. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature females of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/ structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD— standard deviation)." pageId="2" pageNumber="79">Table 3</tableCitation>
)
</paragraph>
<paragraph id="880636870D06FF90D3C9FB28FD06FA9E" blockId="2.[151,1437,535,2012]" pageId="2" pageNumber="79">
Adult female; colour greyish; eyes not visible (after preparation); body length 137 µm (
<emphasis id="BACDEA950D06FF90D796FB28FB58FAE2" box="[1176,1218,1264,1289]" italics="true" pageId="2" pageNumber="79">685</emphasis>
); cephalic cirri and papillae, and clava not enlarged; cirrus A 26.5 µm (
<emphasis id="BACDEA950D06FF90D1FAFACDFCA9FAC6" box="[756,819,1301,1325]" italics="true" pageId="2" pageNumber="79">132.5</emphasis>
); claws I–III of similar size, claw IV longer; dentate collar present, with three teeth; small papilla on leg IV. Dorsal plates with sparse pseudopores/pores; ventral plates not obvious. Female gonopore rosette-shaped.
</paragraph>
<paragraph id="880636870D06FF90D3C9FA58FD6AFA72" blockId="2.[151,1437,535,2012]" box="[199,752,1408,1433]" pageId="2" pageNumber="79">
Short description of
<typeStatus id="570288250D06FF90D2A2FA58FD9CFA72" box="[428,518,1408,1433]" pageId="2" pageNumber="79" type="allotype">allotype</typeStatus>
(
<figureCitation id="10822A020D06FF90D11BFA58FDC4FA72" box="[533,606,1408,1433]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="2" pageNumber="79">Figs 2</figureCitation>
,
<figureCitation id="10822A020D06FF90D164FA59FD13FA72" box="[618,649,1409,1433]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="2" pageNumber="79">12</figureCitation>
;
<tableCitation id="C53B033C0D06FF90D19BFA58FD73FA72" box="[661,745,1408,1433]" captionStart="TABLE 4" captionStartId="11.[151,239,152,175]" captionText="TABLE 4. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature males and one larva of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD—standard deviation;?—unmeasurable structure)." pageId="2" pageNumber="79">Table 4</tableCitation>
)
</paragraph>
<paragraph id="880636870D06FF90D3C9FA7CFA07F9C2" blockId="2.[151,1437,535,2012]" pageId="2" pageNumber="79">
Adult male; colour greyish; eyes not visible (after preparation); body length 123 µm (
<emphasis id="BACDEA950D06FF90D79AFA7CFB24FA56" box="[1172,1214,1444,1469]" italics="true" pageId="2" pageNumber="79">628</emphasis>
); cephalic cirri and papillae, and clava enlarged; cirrus A 20.7 µm (103.0); claws I–III of similar size, claw IV longer; dentate collar present, with three teeth; small papilla on leg IV not visible. Dorsal plates with sparse pseudopores/pores; suggestion of deeper faceting on scapular and caudal plates; ventral plates present. Male gonopore simple, circular.
</paragraph>
<paragraph id="880636870D06FF90D3C9F9ECFE4BF99A" blockId="2.[151,1437,535,2012]" pageId="2" pageNumber="79">
Details of the new species. Adults (
<emphasis id="BACDEA950D06FF90D150F9EDFD1BF9A7" box="[606,641,1589,1612]" italics="true" pageId="2" pageNumber="79">i.e.</emphasis>
from the third instar onwards; measurements and statistics for ♀♀ in
<tableCitation id="C53B033C0D06FF90D399F980FF76F99A" box="[151,236,1624,1649]" captionStart="TABLE 3" captionStartId="10.[151,239,1070,1093]" captionText="TABLE 3. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature females of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/ structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD— standard deviation)." pageId="2" pageNumber="79">Table 3</tableCitation>
, for ♂♂ in
<tableCitation id="C53B033C0D06FF90D263F980FE59F99A" box="[365,451,1624,1649]" captionStart="TABLE 4" captionStartId="11.[151,239,152,175]" captionText="TABLE 4. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature males and one larva of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD—standard deviation;?—unmeasurable structure)." pageId="2" pageNumber="79">Table 4</tableCitation>
):
</paragraph>
<paragraph id="880636870D06FF91D3C9F9A4FC37FCB4" blockId="2.[151,1437,535,2012]" lastBlockId="3.[151,1437,151,2015]" lastPageId="3" lastPageNumber="80" pageId="2" pageNumber="79">
Body greyish; eyes absent or not visible after preparation. Mouth opening surrounded by a ring of 10 papulae (visible in SEM only). Cephalic papillae greatly enlarged in males (compare
<figureCitation id="10822A020D06FF90D70FF978FB34F952" box="[1025,1198,1696,1721]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="2" pageNumber="79">Figs 1 and 2, 4</figureCitation>
), and clavae are also more prominent (
<figureCitation id="10822A020D06FF90D250F91CFE22F936" box="[350,440,1732,1757]" captionStart="FIGURES13–16" captionStartId="7.[151,264,1209,1231]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure@7.[151,1436,193,1188]" captionTargetPageId="7" captionText="FIGURES13–16.Bryodelphax instabilissp.nov., dorsalplates (SEM):13—the scapular plate (paratype, male);14—posterior part of the median 2—median 3 (paratype, male), note almost complete absence of true pores; 15—the scapular plate (Slovakian female); 16—posterior part of the median 1—median 3 (Slovakian female), note numerous large true pores. Scale bars in µm." httpUri="https://zenodo.org/record/1221265/files/figure.png" pageId="2" pageNumber="79">Figs 13</figureCitation>
,
<figureCitation id="10822A020D06FF90D2C8F91DFD8AF936" box="[454,528,1733,1757]" captionStart="FIGURES 17–18" captionStartId="7.[151,264,1936,1958]" captionTargetBox="[151,1436,1372,1913]" captionTargetId="figure@7.[151,1436,1372,1913]" captionTargetPageId="7" captionText="FIGURES 17–18. Bryodelphax instabilis sp. nov., male cephalic appendages (paratype, SEM): 17—cirrus A with clava, the arrowhead indicates the subterminal constriction; 18—enlarged clava. Scale bars inµm." httpUri="https://zenodo.org/record/1221267/files/figure.png" pageId="2" pageNumber="79">17–18</figureCitation>
). Internal cirri much shorter than external (
<tableCitation id="C53B033C0D06FF90D70DF91CFB19F936" box="[1027,1155,1732,1757]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="10.[151,239,1070,1093]" captionStartId-1="11.[151,239,152,175]" captionText-0="TABLE 3. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature females of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/ structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD— standard deviation)." captionText-1="TABLE 4. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature males and one larva of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD—standard deviation;?—unmeasurable structure)." pageId="2" pageNumber="79">Tables 3–4</tableCitation>
,
<figureCitation id="10822A020D06FF90D79FF91CFB6DF936" box="[1169,1271,1732,1757]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="2" pageNumber="79">Figs 1–4</figureCitation>
,
<figureCitation id="10822A020D06FF90D608F91DFAAEF936" box="[1286,1332,1733,1757]" captionStart="FIGURES 6–9" captionStartId="5.[151,264,1301,1323]" captionTargetBox="[151,1436,193,1280]" captionTargetId="figure@5.[151,1436,193,1280]" captionTargetPageId="5" captionText="FIGURES 6–9. Bryodelphax instabilis sp. nov., habitus (SEM):6—female (paratype, dorsal view); 7—male (paratype,lateral view); 8—female (paratype, ventral view); 9—male (paratype, ventral view).Scale bars in µm." httpUri="https://zenodo.org/record/1221261/files/figure.png" pageId="2" pageNumber="79">7–9</figureCitation>
). Cirri A thinner at the tip (visible only under SEM;
<figureCitation id="10822A020D06FF90D176F930FD50F8EA" box="[632,714,1768,1793]" captionStart="FIGURES 17–18" captionStartId="7.[151,264,1936,1958]" captionTargetBox="[151,1436,1372,1913]" captionTargetId="figure@7.[151,1436,1372,1913]" captionTargetPageId="7" captionText="FIGURES 17–18. Bryodelphax instabilis sp. nov., male cephalic appendages (paratype, SEM): 17—cirrus A with clava, the arrowhead indicates the subterminal constriction; 18—enlarged clava. Scale bars inµm." httpUri="https://zenodo.org/record/1221267/files/figure.png" pageId="2" pageNumber="79">Fig. 17</figureCitation>
). Dorsal plates covered with sparsely distributed pseudopores or pores of large and intermediate size (visible in
<collectionCode id="EEA8AE420D06FF90D1BBF8D5FD6AF8CE" box="[693,752,1805,1829]" country="Poland" httpUri="http://grbio.org/cool/f3v1-41fg" name="Polish Collection of Microorganisms" pageId="2" pageNumber="79">PCM</collectionCode>
as bright dots with margins either blurred (pseudopores) or obvious (pores);
<figureCitation id="10822A020D06FF90D254F8E8FE5AF8A2" box="[346,448,1840,1865]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="2" pageNumber="79">Figs 1–2</figureCitation>
,
<figureCitation id="10822A020D06FF90D2C0F8E9FD80F8A2" box="[462,538,1841,1865]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="2" pageNumber="79">10, 12</figureCitation>
), true pores occur infrequently, being restricted to the posterior portion of the cephalic plate and the anterior portion of the scapular plate (
<figureCitation id="10822A020D06FF90D05FF88CFC3EF886" box="[849,932,1876,1901]" captionStart="FIGURES13–16" captionStartId="7.[151,264,1209,1231]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure@7.[151,1436,193,1188]" captionTargetPageId="7" captionText="FIGURES13–16.Bryodelphax instabilissp.nov., dorsalplates (SEM):13—the scapular plate (paratype, male);14—posterior part of the median 2—median 3 (paratype, male), note almost complete absence of true pores; 15—the scapular plate (Slovakian female); 16—posterior part of the median 1—median 3 (Slovakian female), note numerous large true pores. Scale bars in µm." httpUri="https://zenodo.org/record/1221265/files/figure.png" pageId="2" pageNumber="79">Fig. 13</figureCitation>
) in the proximal body part, to the posterior parts of segmental plates (
<figureCitation id="10822A020D06FF90D2B0F8A0FD88F87A" box="[446,530,1912,1937]" captionStart="FIGURES13–16" captionStartId="7.[151,264,1209,1231]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure@7.[151,1436,193,1188]" captionTargetPageId="7" captionText="FIGURES13–16.Bryodelphax instabilissp.nov., dorsalplates (SEM):13—the scapular plate (paratype, male);14—posterior part of the median 2—median 3 (paratype, male), note almost complete absence of true pores; 15—the scapular plate (Slovakian female); 16—posterior part of the median 1—median 3 (Slovakian female), note numerous large true pores. Scale bars in µm." httpUri="https://zenodo.org/record/1221265/files/figure.png" pageId="2" pageNumber="79">Fig. 14</figureCitation>
), and the anterior margin of the caudal plate in the distal body part. Pores rarely present on all plates (
<figureCitation id="10822A020D06FF90D29EF844FE72F85E" box="[400,488,1948,1973]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="2" pageNumber="79">Figs 11</figureCitation>
,
<figureCitation id="10822A020D06FF90D2F9F844FDDAF85E" box="[503,576,1948,1973]" captionStart="FIGURES13–16" captionStartId="7.[151,264,1209,1231]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure@7.[151,1436,193,1188]" captionTargetPageId="7" captionText="FIGURES13–16.Bryodelphax instabilissp.nov., dorsalplates (SEM):13—the scapular plate (paratype, male);14—posterior part of the median 2—median 3 (paratype, male), note almost complete absence of true pores; 15—the scapular plate (Slovakian female); 16—posterior part of the median 1—median 3 (Slovakian female), note numerous large true pores. Scale bars in µm." httpUri="https://zenodo.org/record/1221265/files/figure.png" pageId="2" pageNumber="79">15–16</figureCitation>
), but are often absent (
<figureCitation id="10822A020D06FF90D043F844FC2CF85E" box="[845,950,1948,1973]" captionStart="FIGURES 6–9" captionStartId="5.[151,264,1301,1323]" captionTargetBox="[151,1436,193,1280]" captionTargetId="figure@5.[151,1436,193,1280]" captionTargetPageId="5" captionText="FIGURES 6–9. Bryodelphax instabilis sp. nov., habitus (SEM):6—female (paratype, dorsal view); 7—male (paratype,lateral view); 8—female (paratype, ventral view); 9—male (paratype, ventral view).Scale bars in µm." httpUri="https://zenodo.org/record/1221261/files/figure.png" pageId="2" pageNumber="79">Figs 6–7</figureCitation>
). Pseudopores/pores density similar on all dorsal plates, approximate mean density 10–13 pseudopores/pores per 100 µm
<quantity id="4F419B620D06FF90D702F867FBA8F837" box="[1036,1074,1983,2012]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="79" unit="in" value="2.0">2 in</quantity>
females and 8–10 pseudopores/ pores per 100 µm
<quantity id="4F419B620D07FF91D26BFF4FFE17FF58" box="[357,397,151,179]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="80" unit="in" value="2.0">2 in</quantity>
males (
<tableCitation id="C53B033C0D07FF91D2E5FF42FDDFFF58" box="[491,581,154,179]" captionStart="TABLE 5" captionStartId="11.[151,239,1090,1113]" captionText="TABLE 5. Statistical comparisons of mean pseudopore density per 100 µm 2 of selected dorsal plates between sexes of Bryodelphaxinstabilis sp. nov. (SD—standard deviation). Thep-values for the Welch t-test are provided and statistically significant differences are marked in bold (p&lt;0.050). 14 females and 8 males from the type series were compared." pageId="3" pageNumber="80">Table 5</tableCitation>
). Intracuticular pillars visible as fine dark dots under
<collectionCode id="EEA8AE420D07FF91D7B2FF43FB6DFF58" box="[1212,1271,155,179]" country="Poland" httpUri="http://grbio.org/cool/f3v1-41fg" name="Polish Collection of Microorganisms" pageId="3" pageNumber="80">PCM</collectionCode>
(
<figureCitation id="10822A020D07FF91D604FF42FA0AFF58" box="[1290,1424,154,179]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">Figs 10–12</figureCitation>
). Scapular plate typically distinctly faceted with a median longitudinal fold and three smaller transverse folds (
<figureCitation id="10822A020D07FF91D664FF66FF3DFF10" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 2</figureCitation>
,
<figureCitation id="10822A020D07FF91D3BCFF3BFF4BFF10" box="[178,209,227,251]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">12</figureCitation>
). In females, sometimes only pseudopore/pore rows mark borders of faint facets (
<figureCitation id="10822A020D07FF91D753FF3AFB3EFF10" box="[1117,1188,226,251]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 1</figureCitation>
,
<figureCitation id="10822A020D07FF91D7BFFF3BFB63FF10" box="[1201,1273,227,251]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">10–11</figureCitation>
). Paired plates divided into two unequal anterior and posterior parts by a transverse stripe (
<figureCitation id="10822A020D07FF91D71DFEDEFB02FEF4" box="[1043,1176,262,287]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">Figs 10–12</figureCitation>
,
<figureCitation id="10822A020D07FF91D7A9FEDFFB6EFEF4" box="[1191,1268,262,287]" captionStart="FIGURES13–16" captionStartId="7.[151,264,1209,1231]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure@7.[151,1436,193,1188]" captionTargetPageId="7" captionText="FIGURES13–16.Bryodelphax instabilissp.nov., dorsalplates (SEM):13—the scapular plate (paratype, male);14—posterior part of the median 2—median 3 (paratype, male), note almost complete absence of true pores; 15—the scapular plate (Slovakian female); 16—posterior part of the median 1—median 3 (Slovakian female), note numerous large true pores. Scale bars in µm." httpUri="https://zenodo.org/record/1221265/files/figure.png" pageId="3" pageNumber="80">14, 16</figureCitation>
). Caudal plate faceted with two evident longitudinal folds (
<figureCitation id="10822A020D07FF91D195FEF2FC99FEA8" box="[667,771,298,323]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 1–2</figureCitation>
,
<figureCitation id="10822A020D07FF91D01CFEF2FCB8FEA8" box="[786,802,298,323]" captionStart="FIGURES 6–9" captionStartId="5.[151,264,1301,1323]" captionTargetBox="[151,1436,193,1280]" captionTargetId="figure@5.[151,1436,193,1280]" captionTargetPageId="5" captionText="FIGURES 6–9. Bryodelphax instabilis sp. nov., habitus (SEM):6—female (paratype, dorsal view); 7—male (paratype,lateral view); 8—female (paratype, ventral view); 9—male (paratype, ventral view).Scale bars in µm." httpUri="https://zenodo.org/record/1221261/files/figure.png" pageId="3" pageNumber="80">6</figureCitation>
,
<figureCitation id="10822A020D07FF91D03FFEF3FCE0FEA8" box="[817,890,299,323]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">10–12</figureCitation>
), most often the central portion of this plate is subdivided into two parts by a transverse V-suture (
<figureCitation id="10822A020D07FF91D1EFFE96FCB1FE8C" box="[737,811,334,359]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 2</figureCitation>
,
<figureCitation id="10822A020D07FF91D036FE96FCD2FE8C" box="[824,840,334,359]" captionStart="FIGURES 6–9" captionStartId="5.[151,264,1301,1323]" captionTargetBox="[151,1436,193,1280]" captionTargetId="figure@5.[151,1436,193,1280]" captionTargetPageId="5" captionText="FIGURES 6–9. Bryodelphax instabilis sp. nov., habitus (SEM):6—female (paratype, dorsal view); 7—male (paratype,lateral view); 8—female (paratype, ventral view); 9—male (paratype, ventral view).Scale bars in µm." httpUri="https://zenodo.org/record/1221261/files/figure.png" pageId="3" pageNumber="80">6</figureCitation>
,
<figureCitation id="10822A020D07FF91D05BFE97FCEEFE8C" box="[853,884,335,359]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">12</figureCitation>
). Median plates 1 and 2 with transverse division into two unequal parts (compare
<figureCitation id="10822A020D07FF91D116FEAAFD00FE60" box="[536,666,370,395]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">Figs 10–12</figureCitation>
,
<figureCitation id="10822A020D07FF91D1A7FEABFD6FFE60" box="[681,757,370,395]" captionStart="FIGURES13–16" captionStartId="7.[151,264,1209,1231]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure@7.[151,1436,193,1188]" captionTargetPageId="7" captionText="FIGURES13–16.Bryodelphax instabilissp.nov., dorsalplates (SEM):13—the scapular plate (paratype, male);14—posterior part of the median 2—median 3 (paratype, male), note almost complete absence of true pores; 15—the scapular plate (Slovakian female); 16—posterior part of the median 1—median 3 (Slovakian female), note numerous large true pores. Scale bars in µm." httpUri="https://zenodo.org/record/1221265/files/figure.png" pageId="3" pageNumber="80">14, 16</figureCitation>
). Median plate 3 with faint transverse suture, triangular in shape, and with a roundish posterior edge (
<figureCitation id="10822A020D07FF91D17BFE4EFD6FFE44" box="[629,757,406,431]" captionStart="FIGURES13–16" captionStartId="7.[151,264,1209,1231]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure@7.[151,1436,193,1188]" captionTargetPageId="7" captionText="FIGURES13–16.Bryodelphax instabilissp.nov., dorsalplates (SEM):13—the scapular plate (paratype, male);14—posterior part of the median 2—median 3 (paratype, male), note almost complete absence of true pores; 15—the scapular plate (Slovakian female); 16—posterior part of the median 1—median 3 (Slovakian female), note numerous large true pores. Scale bars in µm." httpUri="https://zenodo.org/record/1221265/files/figure.png" pageId="3" pageNumber="80">Figs 14, 16</figureCitation>
). Poorly developed supplementary lateral platelets present at the levels of median plates (three pairs of platelets on each body side: a pair between scapular plate and first pair of the segmental plates, a pair between paired plates, and a pair between second pair of segmental plates and caudal plate;
<figureCitation id="10822A020D07FF91D3D2FDDAFEBBFDF0" box="[220,289,514,539]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 1</figureCitation>
,
<figureCitation id="10822A020D07FF91D221FDDBFEE2FDF0" box="[303,376,515,539]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">10–12</figureCitation>
), devoid of pores/pseudopores (
<figureCitation id="10822A020D07FF91D1D3FDDAFCB8FDF0" box="[733,802,514,539]" captionStart="FIGURES 6–9" captionStartId="5.[151,264,1301,1323]" captionTargetBox="[151,1436,193,1280]" captionTargetId="figure@5.[151,1436,193,1280]" captionTargetPageId="5" captionText="FIGURES 6–9. Bryodelphax instabilis sp. nov., habitus (SEM):6—female (paratype, dorsal view); 7—male (paratype,lateral view); 8—female (paratype, ventral view); 9—male (paratype, ventral view).Scale bars in µm." httpUri="https://zenodo.org/record/1221261/files/figure.png" pageId="3" pageNumber="80">Fig. 7</figureCitation>
). Ventral cuticle typically with weakly developed plates. Ventral intracuticular pillars either absent or present and well-visible (compare
<figureCitation id="10822A020D07FF91D72DFDFEFB12FDD4" box="[1059,1160,550,575]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 4–5</figureCitation>
) at 100x oil immersion, and always less evident than dorsal (compare
<figureCitation id="10822A020D07FF91D197FD92FD44FD88" box="[665,734,586,611]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 5</figureCitation>
and
<figureCitation id="10822A020D07FF91D01BFD93FCA8FD88" box="[789,818,587,611]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">11</figureCitation>
). Ventral plate configuration: VII/IX:(2)-(1)-2/4-2-2/4- 2-2-2-1. Two small subcephalic longitudinal plates just below the mouth opening and one subpharyngeal plate weakly outlined and usually only visible under SEM (compare
<figureCitation id="10822A020D07FF91D082FD4AFC42FD40" box="[908,984,658,683]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 5</figureCitation>
,
<figureCitation id="10822A020D07FF91D0E4FD4BFB8DFD40" box="[1002,1047,659,683]" captionStart="FIGURES 6–9" captionStartId="5.[151,264,1301,1323]" captionTargetBox="[151,1436,193,1280]" captionTargetId="figure@5.[151,1436,193,1280]" captionTargetPageId="5" captionText="FIGURES 6–9. Bryodelphax instabilis sp. nov., habitus (SEM):6—female (paratype, dorsal view); 7—male (paratype,lateral view); 8—female (paratype, ventral view); 9—male (paratype, ventral view).Scale bars in µm." httpUri="https://zenodo.org/record/1221261/files/figure.png" pageId="3" pageNumber="80">8–9</figureCitation>
). Typically, rows III–VIII/IX are discernible (
<figureCitation id="10822A020D07FF91D22FFD6EFE1CFD24" box="[289,390,694,719]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 4–5</figureCitation>
,
<figureCitation id="10822A020D07FF91D29CFD6FFE24FD24" box="[402,446,695,719]" captionStart="FIGURES 6–9" captionStartId="5.[151,264,1301,1323]" captionTargetBox="[151,1436,193,1280]" captionTargetId="figure@5.[151,1436,193,1280]" captionTargetPageId="5" captionText="FIGURES 6–9. Bryodelphax instabilis sp. nov., habitus (SEM):6—female (paratype, dorsal view); 7—male (paratype,lateral view); 8—female (paratype, ventral view); 9—male (paratype, ventral view).Scale bars in µm." httpUri="https://zenodo.org/record/1221261/files/figure.png" pageId="3" pageNumber="80">8–9</figureCitation>
,
<figureCitation id="10822A020D07FF91D2C4FD6FFE73FD24" box="[458,489,695,719]" captionStart="FIGURES21–23" captionStartId="8.[151,264,1721,1743]" captionTargetBox="[151,1436,873,1700]" captionTargetId="figure@8.[151,1436,858,1700]" captionTargetPageId="8" captionText="FIGURES21–23. A semi-schematic illustration of the ontogenetic variability inthe ventral armature in Bryodelphax instabilis sp. nov.: 21—adult (mature female); 22—juvenile (second instar); 23—larva (first instar)." httpUri="https://zenodo.org/record/1221271/files/figure.png" pageId="3" pageNumber="80">21</figureCitation>
). Rarely, ventral plates invisible under
<collectionCode id="EEA8AE420D07FF91D0AFFD6FFC46FD24" box="[929,988,695,719]" country="Poland" httpUri="http://grbio.org/cool/f3v1-41fg" name="Polish Collection of Microorganisms" pageId="3" pageNumber="80">PCM</collectionCode>
(
<emphasis id="BACDEA950D07FF91D0E3FD60FB8FFD24" box="[1005,1045,696,719]" italics="true" pageId="3" pageNumber="80">e.g.</emphasis>
<typeStatus id="570288250D07FF91D712FD6EFB19FD24" box="[1052,1155,694,719]" pageId="3" pageNumber="80" type="holotype">holotype</typeStatus>
). Ventral granules absent or restricted to the plate surface (
<figureCitation id="10822A020D07FF91D10BFD02FDD2FD18" box="[517,584,730,755]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Fig. 5</figureCitation>
). Papilla on legs I absent, minute papilla on legs IV present (
<figureCitation id="10822A020D07FF91D7E2FD02FAB7FD18" box="[1260,1325,730,755]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Fig. 1</figureCitation>
). Dentate collar on legs IV with 3–6 teeth (
<figureCitation id="10822A020D07FF91D102FD26FDEBFCFC" box="[524,625,766,791]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 1–2</figureCitation>
,
<figureCitation id="10822A020D07FF91D173FD27FD06FCFC" box="[637,668,767,791]" captionStart="FIGURES 19–20" captionStartId="8.[151,264,756,778]" captionTargetBox="[151,1436,193,734]" captionTargetId="figure@8.[151,1436,193,734]" captionTargetPageId="8" captionText="FIGURES 19–20. Bryodelphax instabilis sp. nov., claws (SEM): 19—claws III; 20—claws IV with dentate collar. Scale bars in µm." httpUri="https://zenodo.org/record/1221269/files/figure.png" pageId="3" pageNumber="80">20</figureCitation>
, arrowheads). External claws of all legs smooth, internal claws with tiny spurs pointing strongly downwards with very small gap between spur and claw base, making them almost invisible under
<collectionCode id="EEA8AE420D07FF91D24BFC9FFE1AFCB4" box="[325,384,839,863]" country="Poland" httpUri="http://grbio.org/cool/f3v1-41fg" name="Polish Collection of Microorganisms" pageId="3" pageNumber="80">PCM</collectionCode>
(however, always clear under SEM;
<figureCitation id="10822A020D07FF91D011FC9EFC3BFCB4" box="[799,929,838,863]" captionStart="FIGURES 19–20" captionStartId="8.[151,264,756,778]" captionTargetBox="[151,1436,193,734]" captionTargetId="figure@8.[151,1436,193,734]" captionTargetPageId="8" captionText="FIGURES 19–20. Bryodelphax instabilis sp. nov., claws (SEM): 19—claws III; 20—claws IV with dentate collar. Scale bars in µm." httpUri="https://zenodo.org/record/1221269/files/figure.png" pageId="3" pageNumber="80">Figs 19–20</figureCitation>
).
</paragraph>
<paragraph id="880636870D07FF91D3C9FCB2FE43FC04" blockId="3.[151,1437,151,2015]" pageId="3" pageNumber="80">
Remarks on the sexual dimorphism: Sex differences in the new species are well-marked and embrace: longer, tubbier primary and secondary
<specimenCount id="9EBFFD0E0D07FF91D2F2FC48FD8BFC4C" box="[508,529,912,935]" pageId="3" pageNumber="80" type="male">♂</specimenCount>
clavae (compare
<figureCitation id="10822A020D07FF91D1D2FC56FCA5FC4C" box="[732,831,910,935]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 1–2</figureCitation>
and
<tableCitation id="C53B033C0D07FF91D077FC56FC63FC4C" box="[889,1017,910,935]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="10.[151,239,1070,1093]" captionStartId-1="11.[151,239,152,175]" captionText-0="TABLE 3. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature females of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/ structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD— standard deviation)." captionText-1="TABLE 4. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature males and one larva of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD—standard deviation;?—unmeasurable structure)." pageId="3" pageNumber="80">Tables 3–4</tableCitation>
); stronger faceting of the
<specimenCount id="9EBFFD0E0D07FF91D62FFC48FAACFC4C" box="[1313,1334,912,935]" pageId="3" pageNumber="80" type="male">♂</specimenCount>
scapular plate (compare
<figureCitation id="10822A020D07FF91D248FC6AFD83FC20" box="[326,537,946,971]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="3" pageNumber="80">Figs 10–11 and 12</figureCitation>
); significantly lower pseudopore/pore density on unpaired plates in
<specimenCount id="9EBFFD0E0D07FF91D618FC6BFAB1FC21" box="[1302,1323,947,970]" pageId="3" pageNumber="80" type="male">♂</specimenCount>
(compare ranges presented in
<tableCitation id="C53B033C0D07FF91D278FC0EFE51FC04" box="[374,459,982,1007]" captionStart="TABLE 5" captionStartId="11.[151,239,1090,1113]" captionText="TABLE 5. Statistical comparisons of mean pseudopore density per 100 µm 2 of selected dorsal plates between sexes of Bryodelphaxinstabilis sp. nov. (SD—standard deviation). Thep-values for the Welch t-test are provided and statistically significant differences are marked in bold (p&lt;0.050). 14 females and 8 males from the type series were compared." pageId="3" pageNumber="80">Table 5</tableCitation>
).
</paragraph>
<paragraph id="880636870D07FF91D3C9FC22FC0DFBF8" blockId="3.[151,1437,151,2015]" box="[199,919,1018,1043]" pageId="3" pageNumber="80">
Juveniles (
<emphasis id="BACDEA950D07FF91D231FC24FEF8FBF8" box="[319,354,1020,1043]" italics="true" pageId="3" pageNumber="80">i.e.</emphasis>
the second instar, four-clawed without gonopore):
</paragraph>
<paragraph id="880636870D07FF91D3C9FBF8FBE4FB48" blockId="3.[151,1437,151,2015]" pageId="3" pageNumber="80">
In appearance like adults, but clearly smaller (75 µm) and with indiscernible ventral plates (
<figureCitation id="10822A020D07FF91D7F1FBC6FACDFBDC" box="[1279,1367,1054,1079]" captionStart="FIGURES21–23" captionStartId="8.[151,264,1721,1743]" captionTargetBox="[151,1436,873,1700]" captionTargetId="figure@8.[151,1436,858,1700]" captionTargetPageId="8" captionText="FIGURES21–23. A semi-schematic illustration of the ontogenetic variability inthe ventral armature in Bryodelphax instabilis sp. nov.: 21—adult (mature female); 22—juvenile (second instar); 23—larva (first instar)." httpUri="https://zenodo.org/record/1221271/files/figure.png" pageId="3" pageNumber="80">Fig. 22</figureCitation>
). It is impossible to state firmly whether the species exhibits ontogenetic shifts in ventral armature or the lack of ventral plates in the first two life stages is inconstant. This issue certainly requires more studies, since changes in ventral armature occurring during development were recently detected by
<bibRefCitation id="EC284B760D07FF91D08FFB52FBE0FB48" box="[897,1146,1162,1187]" pageId="3" pageNumber="80" refId="ref10101" refString="Gasiorek, P., Stec, D., Morek, w., Marnissi, J. &amp; Michalczyk, L. (2017) The tardigrade fauna of Tunisia, with an integrative description of Bryodelphax maculatus sp. nov. (Heterotardigrada: Echiniscidae). African Zoology, 52 (2), 77 - 89. https: // doi. org / 10.1080 / 15627020.2017.1297688" type="journal article">
Gąsiorek
<emphasis id="BACDEA950D07FF91D0E3FB54FB84FB48" box="[1005,1054,1162,1187]" italics="true" pageId="3" pageNumber="80">et al</emphasis>
. (2017)
</bibRefCitation>
.
</paragraph>
<paragraph id="880636870D07FF91D3C9FB68FBE7FB2C" blockId="3.[151,1437,151,2015]" box="[199,1149,1198,1224]" pageId="3" pageNumber="80">
Larvae (
<emphasis id="BACDEA950D07FF91D22BFB68FEDDFB2C" box="[293,327,1200,1223]" italics="true" pageId="3" pageNumber="80">i.e.</emphasis>
the first instar, two-clawed without gonopore; measurements in
<tableCitation id="C53B033C0D07FF91D717FB76FBF5FB2C" box="[1049,1135,1198,1223]" captionStart="TABLE 4" captionStartId="11.[151,239,152,175]" captionText="TABLE 4. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature males and one larva of Bryodelphax instabilis sp. nov. mounted in Hoyer’s medium (N—number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD—standard deviation;?—unmeasurable structure)." pageId="3" pageNumber="80">Table 4</tableCitation>
):
</paragraph>
<paragraph id="880636870D07FF91D3C9FB0AFD09FABC" blockId="3.[151,1437,151,2015]" pageId="3" pageNumber="80">
Median plates with poorly delineated margins, lacking pseudopores/pores; supplementary lateral platelets and pedal (leg) plates undeveloped. Pores/pseudopores present only on the anterior and posterior margins of the cephalic, scapular, segmental, and caudal plates (
<figureCitation id="10822A020D07FF91D1C7FAC2FC95FAD8" box="[713,783,1306,1331]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Fig. 3</figureCitation>
). Claws with spurs formed as in adults. Legs IV without dentate collar. Ventral plates absent (
<figureCitation id="10822A020D07FF91D13CFAE6FD1CFABC" box="[562,646,1342,1367]" captionStart="FIGURES21–23" captionStartId="8.[151,264,1721,1743]" captionTargetBox="[151,1436,873,1700]" captionTargetId="figure@8.[151,1436,858,1700]" captionTargetPageId="8" captionText="FIGURES21–23. A semi-schematic illustration of the ontogenetic variability inthe ventral armature in Bryodelphax instabilis sp. nov.: 21—adult (mature female); 22—juvenile (second instar); 23—larva (first instar)." httpUri="https://zenodo.org/record/1221271/files/figure.png" pageId="3" pageNumber="80">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="880636870D07FF91D3C9FABBFEECFA90" blockId="3.[151,1437,151,2015]" box="[199,374,1378,1403]" pageId="3" pageNumber="80">Eggs unknown.</paragraph>
</subSubSection>
<subSubSection id="C0A3650C0D07FF91D3C9FA5EFBCBFA0C" pageId="3" pageNumber="80" type="etymology">
<paragraph id="880636870D07FF91D3C9FA5EFBCBFA0C" blockId="3.[151,1437,151,2015]" pageId="3" pageNumber="80">
<emphasis id="BACDEA950D07FF91D3C9FA5EFECBFA74" bold="true" box="[199,337,1414,1439]" pageId="3" pageNumber="80">Etymology:</emphasis>
The name
<taxonomicName id="4FB94D040D07FF91D2D5FA5EFDD8FA74" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[475,578,1414,1439]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="instabilis">
<emphasis id="BACDEA950D07FF91D2D5FA5EFDD8FA74" box="[475,578,1414,1439]" italics="true" pageId="3" pageNumber="80">instabilis</emphasis>
</taxonomicName>
underlines the variability in the number of ventral plates (see Remarks), which is unusual for
<taxonomicName id="4FB94D040D07FF91D28DFA72FD88FA28" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[387,530,1450,1475]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="genus">
<emphasis id="BACDEA950D07FF91D28DFA72FD88FA28" box="[387,530,1450,1475]" italics="true" pageId="3" pageNumber="80">Bryodelphax</emphasis>
</taxonomicName>
. Moreover, often only some of the ventral plates rows are visible under the light microscope, thus an animal’s venter seems to be devoid of plates in its proximal part.
</paragraph>
</subSubSection>
<subSubSection id="C0A3650C0D07FF91D3C9FA2AFBD2F9B8" pageId="3" pageNumber="80" type="discussion">
<paragraph id="880636870D07FF91D3C9FA2AFBD2F9B8" blockId="3.[151,1437,151,2015]" pageId="3" pageNumber="80">
<emphasis id="BACDEA950D07FF91D3C9FA2AFEA1F9E0" bold="true" box="[199,315,1522,1547]" pageId="3" pageNumber="80">Remarks.</emphasis>
Polish and Slovak populations have same ventral plates configuration but in three mature females from the Slovak population, row V has two additional, smaller ventral plates placed in a more marginal position, and two of the specimens also have such additional plates in row III (
<figureCitation id="10822A020D07FF91D090F9E2FC7FF9B8" box="[926,997,1594,1619]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 5</figureCitation>
and
<figureCitation id="10822A020D07FF91D715F9E3FBA0F9B8" box="[1051,1082,1595,1619]" captionStart="FIGURES21–23" captionStartId="8.[151,264,1721,1743]" captionTargetBox="[151,1436,873,1700]" captionTargetId="figure@8.[151,1436,858,1700]" captionTargetPageId="8" captionText="FIGURES21–23. A semi-schematic illustration of the ontogenetic variability inthe ventral armature in Bryodelphax instabilis sp. nov.: 21—adult (mature female); 22—juvenile (second instar); 23—larva (first instar)." httpUri="https://zenodo.org/record/1221271/files/figure.png" pageId="3" pageNumber="80">21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C0A3650C0D07FF91D3C9F986FEC4F8EC" pageId="3" pageNumber="80" type="diagnosis">
<paragraph id="880636870D07FF91D3C9F986FEC4F8EC" blockId="3.[151,1437,151,2015]" pageId="3" pageNumber="80">
<emphasis id="BACDEA950D07FF91D3C9F986FE54F99C" bold="true" box="[199,462,1630,1655]" pageId="3" pageNumber="80">Differential diagnosis.</emphasis>
<taxonomicName id="4FB94D040D07FF91D2DAF986FD48F99C" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[468,722,1630,1655]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D07FF91D2DAF986FD48F99C" box="[468,722,1630,1655]" italics="true" pageId="3" pageNumber="80">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D07FF91D1D7F986FCA9F99C" box="[729,819,1630,1655]" pageId="3" pageNumber="80" rank="species">
<emphasis id="BACDEA950D07FF91D1D7F986FCA9F99C" bold="true" box="[729,819,1630,1655]" pageId="3" pageNumber="80">sp. nov.</emphasis>
</taxonomicNameLabel>
can easily be distinguished from the other members of the
<taxonomicName id="4FB94D040D07FF91D3CDF95AFED9F970" baseAuthorityName="Pilato" baseAuthorityYear="1972" box="[195,323,1666,1691]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="BACDEA950D07FF91D3CDF95AFED9F970" box="[195,323,1666,1691]" italics="true" pageId="3" pageNumber="80">weglarskae</emphasis>
</taxonomicName>
group on the basis of the scapular plate faceting and being gonochorous. Nevertheless, the new species should be compared with four most similar taxa having last six ventral plates rows configuration same as the new species,
<emphasis id="BACDEA950D07FF91D26FF914FEE7F908" box="[353,381,1740,1763]" italics="true" pageId="3" pageNumber="80">i.e</emphasis>
. 2-2-2-2-2-1 or 2-4-2-2-2-1 (due to the instability in ventral plate arrangement). It differs specifically from:
</paragraph>
</subSubSection>
<subSubSection id="C0A3650C0D07FF97D3C9F8CAFBF8F824" lastPageId="5" lastPageNumber="82" pageId="3" pageNumber="80" type="reference_group">
<paragraph id="880636870D07FF91D3C9F8CAFA15F898" blockId="3.[151,1437,151,2015]" pageId="3" pageNumber="80">
<taxonomicName id="4FB94D040D07FF91D3C9F8CAFD2CF8C0" ID-CoL="NHZL" authority="Bertolani et al., 1996" authorityName="Bertolani et al." authorityYear="1996" box="[199,694,1810,1835]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="iohannis">
<emphasis id="BACDEA950D07FF91D3C9F8CAFE27F8C0" box="[199,445,1810,1835]" italics="true" pageId="3" pageNumber="80">Bryodelphax iohannis</emphasis>
<bibRefCitation id="EC284B760D07FF91D2CBF8CAFD2CF8C0" box="[453,694,1810,1835]" pageId="3" pageNumber="80" refId="ref9715" refString="Bertolani, R., Guidi, A. &amp; Rebecchi, L. (1996) Tardigradi della Sardegna e di alcune piccole isole circum-sarde. Biogeographia, 18, 229 - 247." type="journal article">
Bertolani
<emphasis id="BACDEA950D07FF91D13AF8CCFDFCF8C0" box="[564,614,1810,1835]" italics="true" pageId="3" pageNumber="80">et al</emphasis>
., 1996
</bibRefCitation>
</taxonomicName>
, by a different ventral plate configuration (VII/IX:(2)-(1)-2/4-2-2/ 4-2-2-2-
<quantity id="4F419B620D07FF91D3F6F8EFFEBAF8A4" box="[248,288,1846,1871]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="80" unit="in" value="1.0">1 in</quantity>
the new species
<emphasis id="BACDEA950D07FF91D2EFF8E0FE62F8A4" box="[481,504,1848,1871]" italics="true" pageId="3" pageNumber="80">vs</emphasis>
X: 2-1-1-5-2-4-2-2-2-
<quantity id="4F419B620D07FF91D1F7F8EEFCBBF8A4" box="[761,801,1846,1871]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="80" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4FB94D040D07FF91D027F8E0FC33F8A4" ID-CoL="NHZL" box="[809,937,1846,1871]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="iohannis">
<emphasis id="BACDEA950D07FF91D027F8E0FC33F8A4" box="[809,937,1846,1871]" italics="true" pageId="3" pageNumber="80">B. iohannis</emphasis>
</taxonomicName>
), by the presence of lateral platelets (absent in
<taxonomicName id="4FB94D040D07FF91D3BAF884FEAEF898" ID-CoL="NHZL" box="[180,308,1882,1907]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="iohannis">
<emphasis id="BACDEA950D07FF91D3BAF884FEAEF898" box="[180,308,1882,1907]" italics="true" pageId="3" pageNumber="80">B. iohannis</emphasis>
</taxonomicName>
) and by longer teeth on the dentate collar (compare
<figureCitation id="10822A020D07FF91D070F882FC44F898" box="[894,990,1882,1907]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Figs 1–2</figureCitation>
with
<figureCitation id="10822A020D07FF91D711F882FBEEF898" box="[1055,1140,1882,1907]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="3" pageNumber="80">Fig. 2A</figureCitation>
in
<bibRefCitation id="EC284B760D07FF91D796F882FA1BF898" box="[1176,1409,1882,1907]" pageId="3" pageNumber="80" refId="ref9715" refString="Bertolani, R., Guidi, A. &amp; Rebecchi, L. (1996) Tardigradi della Sardegna e di alcune piccole isole circum-sarde. Biogeographia, 18, 229 - 247." type="journal article">
Bertolani
<emphasis id="BACDEA950D07FF91D609F884FAA2F898" box="[1287,1336,1882,1907]" italics="true" pageId="3" pageNumber="80">et al</emphasis>
. 1996
</bibRefCitation>
);
</paragraph>
<paragraph id="880636870D07FF91D3C9F8A6FC35F834" blockId="3.[151,1437,151,2015]" pageId="3" pageNumber="80">
<taxonomicName id="4FB94D040D07FF91D3C9F8A6FC96F87C" ID-CoL="2bce9d79-e03c-4d8d-b511-a57dac2a476e" authority="Kaczmarek et al., 2012" authorityName="Kaczmarek et al." authorityYear="2012" box="[199,780,1918,1943]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="parvuspolaris">
<emphasis id="BACDEA950D07FF91D3C9F8A6FE60F87C" box="[199,506,1918,1943]" italics="true" pageId="3" pageNumber="80">Bryodelphax parvuspolaris</emphasis>
<bibRefCitation id="EC284B760D07FF91D10FF8A6FC96F87C" box="[513,780,1918,1943]" pageId="3" pageNumber="80" refId="ref10449" refString="Kaczmarek, L., Zawierucha, K., Smykla, J. &amp; Michalczyk, L. (2012) Tardigrada of the Revdalen (Spitsbergen) with the descriptions of two new species: Bryodelphax parvuspolaris (Heterotardigrada) and Isohypsibius coulsoni (Eutardigrada). Polar Biology, 35, 1013 - 1026. https: // doi. org / 10.1007 / s 00300 - 011 - 1149 - 0" type="journal article">
Kaczmarek
<emphasis id="BACDEA950D07FF91D187F858FD21F87C" box="[649,699,1918,1943]" italics="true" pageId="3" pageNumber="80">et al</emphasis>
., 2012
</bibRefCitation>
</taxonomicName>
, by a different ventral plate configuration (VII/IX:(2)-(1)- 2/4-2-2/4-2-2-2-
<quantity id="4F419B620D07FF91D25DF87BFEE0F850" box="[339,378,1954,1979]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="80" unit="in" value="1.0">1 in</quantity>
the new species
<emphasis id="BACDEA950D07FF91D134F87CFDCBF850" box="[570,593,1956,1979]" italics="true" pageId="3" pageNumber="80">vs</emphasis>
VIII: 1-1-2-2-2-2-2-
<quantity id="4F419B620D07FF91D04EF87BFCF2F850" box="[832,872,1954,1979]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="80" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4FB94D040D07FF91D061F87CFBB1F850" ID-CoL="2bce9d79-e03c-4d8d-b511-a57dac2a476e" box="[879,1067,1954,1979]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="parvuspolaris">
<emphasis id="BACDEA950D07FF91D061F87CFBB1F850" box="[879,1067,1954,1979]" italics="true" pageId="3" pageNumber="80">B. parvuspolaris</emphasis>
</taxonomicName>
), and by well-developed dentate collar (dentate collar with poorly developed teeth in
<taxonomicName id="4FB94D040D07FF91D1EDF810FC04F834" ID-CoL="2bce9d79-e03c-4d8d-b511-a57dac2a476e" box="[739,926,1990,2015]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="80" phylum="Tardigrada" rank="species" species="parvuspolaris">
<emphasis id="BACDEA950D07FF91D1EDF810FC04F834" box="[739,926,1990,2015]" italics="true" pageId="3" pageNumber="80">B. parvuspolaris</emphasis>
</taxonomicName>
);
</paragraph>
<caption id="DCC6660F0D00FF96D399F92DFD81F84E" httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="4" pageNumber="81" startId="4.[151,264,1781,1803]" targetBox="[151,1436,195,1757]" targetPageId="4">
<paragraph id="880636870D00FF96D399F92DFD81F84E" blockId="4.[151,1436,1781,1957]" pageId="4" pageNumber="81">
<emphasis id="BACDEA950D00FF96D399F92DFEA4F8E0" bold="true" box="[151,318,1781,1803]" pageId="4" pageNumber="81">FIGURES 1–5.</emphasis>
<taxonomicName id="4FB94D040D00FF96D246F92DFDAAF8E0" ID-CoL="358bf5e4-073b-4ef9-ad5e-7f59779e5d57" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[328,560,1781,1803]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="81" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D00FF96D246F92DFDAAF8E0" box="[328,560,1781,1803]" italics="true" pageId="4" pageNumber="81">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D00FF96D134F92DFD14F8E0" box="[570,654,1781,1803]" pageId="4" pageNumber="81" rank="species">
<emphasis id="BACDEA950D00FF96D134F92DFD14F8E0" bold="true" box="[570,654,1781,1803]" pageId="4" pageNumber="81">sp. nov.</emphasis>
</taxonomicNameLabel>
, habitus (all but Fig. 3 in PCM):
<emphasis id="BACDEA950D00FF96D0F0F92DFBBFF8E0" bold="true" box="[1022,1061,1781,1803]" pageId="4" pageNumber="81">1—</emphasis>
female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets);
<emphasis id="BACDEA950D00FF96D02EF8CCFCDCF8C1" bold="true" box="[800,838,1812,1834]" pageId="4" pageNumber="81">2—</emphasis>
male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV);
<emphasis id="BACDEA950D00FF96D0AEF8EBFC5CF8A2" bold="true" box="[928,966,1843,1865]" pageId="4" pageNumber="81">3—</emphasis>
larva (paratype, dorsal view, slide PL.189.32, DIC);
<emphasis id="BACDEA950D00FF96D3D3F889FE99F88C" bold="true" box="[221,259,1873,1895]" pageId="4" pageNumber="81">4—</emphasis>
male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows);
<emphasis id="BACDEA950D00FF96D7F6F889FA84F88C" bold="true" box="[1272,1310,1873,1895]" pageId="4" pageNumber="81">5—</emphasis>
female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows). Scale bars in µm.
</paragraph>
</caption>
<caption id="DCC6660F0D01FF97D399FACDFBC1FAA1" httpUri="https://zenodo.org/record/1221261/files/figure.png" pageId="5" pageNumber="82" startId="5.[151,264,1301,1323]" targetBox="[151,1436,193,1280]" targetPageId="5">
<paragraph id="880636870D01FF97D399FACDFBC1FAA1" blockId="5.[151,1435,1301,1354]" pageId="5" pageNumber="82">
<emphasis id="BACDEA950D01FF97D399FACDFEA0FAC0" bold="true" box="[151,314,1301,1323]" pageId="5" pageNumber="82">FIGURES 6–9.</emphasis>
<taxonomicName id="4FB94D040D01FF97D24EFACDFDBFFAC0" ID-CoL="358bf5e4-073b-4ef9-ad5e-7f59779e5d57" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[320,549,1301,1323]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D01FF97D24EFACDFDBFFAC0" box="[320,549,1301,1323]" italics="true" pageId="5" pageNumber="82">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D01FF97D125FACDFDE1FAC0" box="[555,635,1301,1323]" pageId="5" pageNumber="82" rank="species">
<emphasis id="BACDEA950D01FF97D125FACDFDE1FAC0" bold="true" box="[555,635,1301,1323]" pageId="5" pageNumber="82">sp. nov.</emphasis>
</taxonomicNameLabel>
, habitus (SEM):
<emphasis id="BACDEA950D01FF97D024FACDFCCAFAC0" bold="true" box="[810,848,1301,1323]" pageId="5" pageNumber="82">6—</emphasis>
female (paratype, dorsal view);
<emphasis id="BACDEA950D01FF97D79DFACDFB23FAC0" bold="true" box="[1171,1209,1301,1323]" pageId="5" pageNumber="82">7—</emphasis>
male (paratype, lateral view);
<emphasis id="BACDEA950D01FF97D3D0FAECFE99FAA1" bold="true" box="[222,259,1332,1354]" pageId="5" pageNumber="82">8—</emphasis>
female (paratype, ventral view);
<emphasis id="BACDEA950D01FF97D141FAECFDEEFAA1" bold="true" box="[591,628,1332,1354]" pageId="5" pageNumber="82">9—</emphasis>
male (paratype, ventral view). Scale bars in µm.
</paragraph>
</caption>
<paragraph id="880636870D01FF97D3C9FAAEFD68FA3C" blockId="5.[151,1436,1398,1999]" pageId="5" pageNumber="82">
<taxonomicName id="4FB94D040D01FF97D3C9FAAEFDC3FA64" ID-CoL="NHZQ" authority="(Pilato, 1974)" authorityName="Pilato" authorityYear="1974" baseAuthorityName="Pilato" baseAuthorityYear="1974" box="[199,601,1398,1423]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="sinensis">
<emphasis id="BACDEA950D01FF97D3C9FAAEFE2CFA65" box="[199,438,1398,1423]" italics="true" pageId="5" pageNumber="82">Bryodelphax sinensis</emphasis>
(
<bibRefCitation id="EC284B760D01FF97D2CAFAAEFDCBFA64" box="[452,593,1398,1423]" pageId="5" pageNumber="82" refId="ref11190" refString="Pilato, G. (1974) Tre nuove specie di Tardigradi muscicoli di Cina. Animalia, 1, 59 - 68." type="journal article">Pilato, 1974</bibRefCitation>
)
</taxonomicName>
, by a different ventral plate configuration (VII/IX:(2)-(1)-2/4-2-2/4-2-2-2-
<quantity id="4F419B620D01FF97D399FA43FF59FA58" box="[151,195,1434,1459]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="82" unit="in" value="1.0">1 in</quantity>
the new species
<emphasis id="BACDEA950D01FF97D28AFA44FE01FA58" box="[388,411,1436,1459]" italics="true" pageId="5" pageNumber="82">vs</emphasis>
VII:2-2-2-2-2-2-
<quantity id="4F419B620D01FF97D16CFA43FD10FA58" box="[610,650,1434,1459]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="82" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4FB94D040D01FF97D19CFA44FC91FA58" ID-CoL="NHZQ" baseAuthorityName="Pilato" baseAuthorityYear="1974" box="[658,779,1436,1459]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="sinensis">
<emphasis id="BACDEA950D01FF97D19CFA44FC91FA58" box="[658,779,1436,1459]" italics="true" pageId="5" pageNumber="82">B. sinensis</emphasis>
</taxonomicName>
), and by the presence of well-developed dentate collar on legs IV in adults (dentate collar absent in
<taxonomicName id="4FB94D040D01FF97D167FA67FD7BFA3D" ID-CoL="NHZQ" baseAuthorityName="Pilato" baseAuthorityYear="1974" box="[617,737,1471,1494]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="sinensis">
<emphasis id="BACDEA950D01FF97D167FA67FD7BFA3D" box="[617,737,1471,1494]" italics="true" pageId="5" pageNumber="82">B. sinensis</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="880636870D01FF97D3C9FA3AFE7EF9A8" blockId="5.[151,1436,1398,1999]" pageId="5" pageNumber="82">
<taxonomicName id="4FB94D040D01FF97D3C9FA3AFD18FA10" ID-CoL="NHZS" authority="(Pilato, 1972)" authorityName="Pilato" authorityYear="1972" baseAuthorityName="Pilato" baseAuthorityYear="1972" box="[199,642,1506,1531]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="BACDEA950D01FF97D3C9FA3AFE44FA10" box="[199,478,1506,1531]" italics="true" pageId="5" pageNumber="82">Bryodelphax weglarskae</emphasis>
(
<bibRefCitation id="EC284B760D01FF97D2E3FA3AFDE1FA10" box="[493,635,1506,1531]" pageId="5" pageNumber="82" refId="ref11155" refString="Pilato, G. (1972) Prime osservazioni sui tardigradi delle Isole Egadi. Bollettino delle sedute dell'Accademia gioenia di scienze naturale in Catania, 11 (5), 111 - 124." type="journal article">Pilato, 1972</bibRefCitation>
)
</taxonomicName>
, by a different ventral plate configuration (VII/IX:(2)-(1)-2/4-2-2/4-2- 2-2-
<quantity id="4F419B620D01FF97D3C7F9DFFF69F9F4" box="[201,243,1542,1567]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="82" unit="in" value="1.0">1 in</quantity>
the new species
<emphasis id="BACDEA950D01FF97D2B4F9DFFE4BF9F5" box="[442,465,1543,1566]" italics="true" pageId="5" pageNumber="82">vs</emphasis>
IX:2-1-5-2-4-2-2-2-
<quantity id="4F419B620D01FF97D1B1F9DEFD73F9F4" box="[703,745,1542,1567]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="82" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4FB94D040D01FF97D1FCF9DFFC09F9F4" ID-CoL="NHZS" baseAuthorityName="Pilato" baseAuthorityYear="1972" box="[754,915,1542,1567]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="BACDEA950D01FF97D1FCF9DFFC09F9F4" box="[754,915,1542,1567]" italics="true" pageId="5" pageNumber="82">B. weglarskae</emphasis>
</taxonomicName>
), and by non-bifurcated cephalic appendages (bifurcated in
<taxonomicName id="4FB94D040D01FF97D23BF9F4FE49F9A8" ID-CoL="NHZS" baseAuthorityName="Pilato" baseAuthorityYear="1972" box="[309,467,1578,1603]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="BACDEA950D01FF97D23BF9F4FE49F9A8" box="[309,467,1578,1603]" italics="true" pageId="5" pageNumber="82">B. weglarskae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="880636870D01FF97D3C9F997FBF8F824" blockId="5.[151,1436,1398,1999]" pageId="5" pageNumber="82">
Moreover,
<taxonomicName id="4FB94D040D01FF97D24DF996FDD8F98C" ID-CoL="358bf5e4-073b-4ef9-ad5e-7f59779e5d57" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[323,578,1614,1639]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D01FF97D24DF996FDD8F98C" box="[323,578,1614,1639]" italics="true" pageId="5" pageNumber="82">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D01FF97D144F996FD3EF98C" box="[586,676,1614,1639]" pageId="5" pageNumber="82" rank="species">
<emphasis id="BACDEA950D01FF97D144F996FD3EF98C" bold="true" box="[586,676,1614,1639]" pageId="5" pageNumber="82">sp. nov.</emphasis>
</taxonomicNameLabel>
must be primarily compared with the first described gonochoristic representative of the genus, namely
<taxonomicName id="4FB94D040D01FF97D147F9AAFBBDF960" ID-CoL="NHZR" authority="(Weglarska, 1959)" authorityName="Weglarska" authorityYear="1959" baseAuthorityName="Weglarska" baseAuthorityYear="1959" box="[585,1063,1650,1675]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="tatrensis">
<emphasis id="BACDEA950D01FF97D147F9AAFCDEF960" box="[585,836,1650,1675]" italics="true" pageId="5" pageNumber="82">Bryodelphax tatrensis</emphasis>
(Węglarska, 1959)
</taxonomicName>
because it resembles the latter species when the ventral armature is faint.
<taxonomicName id="4FB94D040D01FF97D179F94EFCEEF944" ID-CoL="358bf5e4-073b-4ef9-ad5e-7f59779e5d57" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[631,884,1686,1711]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D01FF97D179F94EFCEEF944" box="[631,884,1686,1711]" italics="true" pageId="5" pageNumber="82">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D01FF97D072F94EFC4FF944" box="[892,981,1686,1711]" pageId="5" pageNumber="82" rank="species">
<emphasis id="BACDEA950D01FF97D072F94EFC4FF944" bold="true" box="[892,981,1686,1711]" pageId="5" pageNumber="82">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from
<taxonomicName id="4FB94D040D01FF97D7DEF94FFAD5F945" ID-CoL="NHZR" baseAuthorityName="Weglarska" baseAuthorityYear="1959" box="[1232,1359,1687,1710]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="tatrensis">
<emphasis id="BACDEA950D01FF97D7DEF94FFAD5F945" box="[1232,1359,1687,1710]" italics="true" pageId="5" pageNumber="82">B. tatrensis</emphasis>
</taxonomicName>
on the basis of: (1) the average pseudopore/pore size is larger in the new species in comparison with the minute pores in
<taxonomicName id="4FB94D040D01FF97D68AF964FF6DF91D" ID-CoL="NHZR" baseAuthorityName="Weglarska" baseAuthorityYear="1959" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="tatrensis">
<emphasis id="BACDEA950D01FF97D68AF964FF6DF91D" italics="true" pageId="5" pageNumber="82">B. tatrensis</emphasis>
</taxonomicName>
(the difference is especially obvious between females of both species, compare
<figureCitation id="10822A020D01FF97D780F906FA96F91C" box="[1166,1292,1758,1783]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="5" pageNumber="82">Figs 10–11</figureCitation>
and
<figureCitation id="10822A020D01FF97D648F906FAFFF91C" box="[1350,1381,1758,1783]" captionStart="FIGURES 24–27" captionStartId="9.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@9.[151,1436,193,1825]" captionTargetPageId="9" captionText="FIGURES 24–27. Other Polish members of the genus Bryodelphax Thulin, 1928, and the comparison of sculpture of dorsal plates (PCM): 24—B. parvulus Thulin, 1928 (female, dorsolateral view); 25—B. tatrensis (węglarska, 1959) (male, dorsal view), note lack of the scapular plate faceting, the arrowhead indicates dentate collar on legs IV, and empty arrowheads point enlarged cephalic papillae; B. tatrensis, sexual dimorphism in the pore size: 26—female; 27—male.Note numerous true pores distributedon all plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221273/files/figure.png" pageId="5" pageNumber="82">26</figureCitation>
), (2) more pronounced faceting of the scapular plate (clearly visible median suture and typically well-developed facets in the new species
<emphasis id="BACDEA950D01FF97D27EF8FFFE1DF8D5" box="[368,391,1831,1854]" italics="true" pageId="5" pageNumber="82">vs</emphasis>
only faint median suture, which can be absent in
<taxonomicName id="4FB94D040D01FF97D0CCF8FFFBD8F8D5" ID-CoL="NHZR" baseAuthorityName="Weglarska" baseAuthorityYear="1959" box="[962,1090,1831,1854]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="tatrensis">
<emphasis id="BACDEA950D01FF97D0CCF8FFFBD8F8D5" box="[962,1090,1831,1854]" italics="true" pageId="5" pageNumber="82">B. tatrensis</emphasis>
</taxonomicName>
; compare
<figureCitation id="10822A020D01FF97D7B5F8FEFAA3F8D4" box="[1211,1337,1830,1855]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="5" pageNumber="82">
Figs
<date id="FC0710470D01FF97D7FAF8FFFAA3F8D4" box="[1268,1337,1831,1855]" pageId="5" pageNumber="82" value="1927-11-12" valueMax="1927-11-26" valueMin="1927-11-12">11–12</date>
</figureCitation>
and
<figureCitation id="10822A020D01FF97D67CF8FEFF2CF888" captionStart="FIGURES 24–27" captionStartId="9.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@9.[151,1436,193,1825]" captionTargetPageId="9" captionText="FIGURES 24–27. Other Polish members of the genus Bryodelphax Thulin, 1928, and the comparison of sculpture of dorsal plates (PCM): 24—B. parvulus Thulin, 1928 (female, dorsolateral view); 25—B. tatrensis (węglarska, 1959) (male, dorsal view), note lack of the scapular plate faceting, the arrowhead indicates dentate collar on legs IV, and empty arrowheads point enlarged cephalic papillae; B. tatrensis, sexual dimorphism in the pore size: 26—female; 27—male.Note numerous true pores distributedon all plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221273/files/figure.png" pageId="5" pageNumber="82">26– 27</figureCitation>
), and (3) the caudal plate consisting of four facets in males of the new species instead of three in
<taxonomicName id="4FB94D040D01FF97D612F894FA01F888" ID-CoL="NHZR" baseAuthorityName="Weglarska" baseAuthorityYear="1959" box="[1308,1435,1868,1891]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="tatrensis">
<emphasis id="BACDEA950D01FF97D612F894FA01F888" box="[1308,1435,1868,1891]" italics="true" pageId="5" pageNumber="82">B. tatrensis</emphasis>
</taxonomicName>
males (compare
<figureCitation id="10822A020D01FF97D25CF8B6FE01F86C" box="[338,411,1902,1927]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1781,1803]" captionTargetBox="[151,1436,195,1757]" captionTargetId="figure@4.[151,1436,195,1757]" captionTargetPageId="4" captionText="FIGURES 1–5. Bryodelphax instabilis sp. nov., habitus (all but Fig. 3 in PCM): 1—female (holotype, dorsolateral view, incised arrowheads indicate supplementary lateral platelets); 2—male (allotype, dorsal view, an empty arrowhead points enlarged cephalic papillae; arrowheads indicate dentate collar on legs IV); 3—larva (paratype, dorsal view, slide PL.189.32, DIC); 4—male (paratype, ventral view, slide PL.189.05, Roman numerals indicate ventral plates rows); 5—female with aberrant additional ventral plates in rows III and V (Slovakian specimen, ventral view, slide 390/32, Roman numerals indicate ventral plates rows).Scale bars in µm." httpUri="https://zenodo.org/record/1221259/files/figure.png" pageId="5" pageNumber="82">Figs 2</figureCitation>
,
<figureCitation id="10822A020D01FF97D2A6F8B7FE5EF86C" box="[424,452,1903,1927]" captionStart="FIGURES10–12" captionStartId="6.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@6.[151,1436,193,1825]" captionTargetPageId="6" captionText="FIGURES10–12. Bryodelphax instabilis sp. nov., dorsal plates(PCM):10—female(holotype, dorsolateral view); 11—female (Slovakian specimen, dorsal view, slide no. 390/32); 12—male (allotype, dorsal view), note deep faceting of the scapular and caudal plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="5" pageNumber="82">12</figureCitation>
and
<figureCitation id="10822A020D01FF97D2F3F8B6FDD2F86C" box="[509,584,1902,1927]" captionStart="FIGURES 24–27" captionStartId="9.[151,264,1846,1868]" captionTargetBox="[151,1436,193,1825]" captionTargetId="figure@9.[151,1436,193,1825]" captionTargetPageId="9" captionText="FIGURES 24–27. Other Polish members of the genus Bryodelphax Thulin, 1928, and the comparison of sculpture of dorsal plates (PCM): 24—B. parvulus Thulin, 1928 (female, dorsolateral view); 25—B. tatrensis (węglarska, 1959) (male, dorsal view), note lack of the scapular plate faceting, the arrowhead indicates dentate collar on legs IV, and empty arrowheads point enlarged cephalic papillae; B. tatrensis, sexual dimorphism in the pore size: 26—female; 27—male.Note numerous true pores distributedon all plates.Scale bars in µm." httpUri="https://zenodo.org/record/1221273/files/figure.png" pageId="5" pageNumber="82">25, 27</figureCitation>
). Additional discriminative criteria (
<emphasis id="BACDEA950D01FF97D0E8F8B7FB95F86D" box="[998,1039,1903,1926]" italics="true" pageId="5" pageNumber="82">e.g.</emphasis>
morphometric) could be presented when measurements of a large population of
<taxonomicName id="4FB94D040D01FF97D1A4F84CFCB1F840" ID-CoL="NHZR" baseAuthorityName="Weglarska" baseAuthorityYear="1959" box="[682,811,1940,1963]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="tatrensis">
<emphasis id="BACDEA950D01FF97D1A4F84CFCB1F840" box="[682,811,1940,1963]" italics="true" pageId="5" pageNumber="82">B. tatrensis</emphasis>
</taxonomicName>
become available. Unfortunately, general rarity of
<taxonomicName id="4FB94D040D01FF97D68AF84CFF6DF825" ID-CoL="NHZR" baseAuthorityName="Weglarska" baseAuthorityYear="1959" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="82" phylum="Tardigrada" rank="species" species="tatrensis">
<emphasis id="BACDEA950D01FF97D68AF84CFF6DF825" italics="true" pageId="5" pageNumber="82">B. tatrensis</emphasis>
</taxonomicName>
hinders the redescription of this taxon (
<bibRefCitation id="EC284B760D01FF97D1BDF86EFBCDF824" box="[691,1111,1974,1999]" pageId="5" pageNumber="82" refId="ref9805" refString="Dastych, H. (1988) The Tardigrada of Poland. Monografie Fauny Polski, 16, 1 - 255." type="journal article">Dastych 1988, personal observations</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DCC6660F0D02FF94D399F8EEFE4DF861" httpUri="https://zenodo.org/record/1221263/files/figure.png" pageId="6" pageNumber="83" startId="6.[151,264,1846,1868]" targetBox="[151,1436,193,1825]" targetPageId="6">
<paragraph id="880636870D02FF94D399F8EEFE4DF861" blockId="6.[151,1436,1846,1930]" pageId="6" pageNumber="83">
<emphasis id="BACDEA950D02FF94D399F8EEFEC9F8A7" bold="true" box="[151,339,1846,1868]" pageId="6" pageNumber="83">FIGURES 10–12.</emphasis>
<taxonomicName id="4FB94D040D02FF94D257F8EEFDA7F8A7" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[345,573,1846,1868]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="83" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D02FF94D257F8EEFDA7F8A7" box="[345,573,1846,1868]" italics="true" pageId="6" pageNumber="83">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D02FF94D14DF8EEFD08F8A7" box="[579,658,1846,1868]" pageId="6" pageNumber="83" rank="species">
<emphasis id="BACDEA950D02FF94D14DF8EEFD08F8A7" bold="true" box="[579,658,1846,1868]" pageId="6" pageNumber="83">sp. nov.</emphasis>
</taxonomicNameLabel>
, dorsal plates (PCM):
<emphasis id="BACDEA950D02FF94D079F8EEFC33F8A7" bold="true" box="[887,937,1846,1868]" pageId="6" pageNumber="83">10—</emphasis>
female (holotype, dorsolateral view);
<emphasis id="BACDEA950D02FF94D628F8EEFACFF8A7" bold="true" box="[1318,1365,1846,1868]" pageId="6" pageNumber="83">11—</emphasis>
female (Slovakian specimen, dorsal view, slide no. 390/32);
<emphasis id="BACDEA950D02FF94D1B9F88DFD73F880" bold="true" box="[695,745,1877,1899]" pageId="6" pageNumber="83">12—</emphasis>
male (allotype, dorsal view), note deep faceting of the scapular and caudal plates. Scale bars in µm.
</paragraph>
</caption>
<caption id="DCC6660F0D03FF95D399FB61FE90FAC0" httpUri="https://zenodo.org/record/1221265/files/figure.png" pageId="7" pageNumber="84" startId="7.[151,264,1209,1231]" targetBox="[151,1436,193,1188]" targetPageId="7">
<paragraph id="880636870D03FF95D399FB61FE90FAC0" blockId="7.[151,1436,1209,1323]" pageId="7" pageNumber="84">
<emphasis id="BACDEA950D03FF95D399FB61FEC9FB24" bold="true" box="[151,339,1209,1231]" pageId="7" pageNumber="84">FIGURES 13–16.</emphasis>
<taxonomicName id="4FB94D040D03FF95D256FB61FDA7FB24" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[344,573,1209,1231]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="84" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D03FF95D256FB61FDA7FB24" box="[344,573,1209,1231]" italics="true" pageId="7" pageNumber="84">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D03FF95D14CFB61FD08FB24" box="[578,658,1209,1231]" pageId="7" pageNumber="84" rank="species">
<emphasis id="BACDEA950D03FF95D14CFB61FD08FB24" bold="true" box="[578,658,1209,1231]" pageId="7" pageNumber="84">sp. nov.</emphasis>
</taxonomicNameLabel>
, dorsal plates (SEM):
<emphasis id="BACDEA950D03FF95D07BFB61FC3DFB24" bold="true" box="[885,935,1209,1231]" pageId="7" pageNumber="84">13—</emphasis>
the scapular plate (paratype, male);
<emphasis id="BACDEA950D03FF95D601FB61FADBFB24" bold="true" box="[1295,1345,1209,1231]" pageId="7" pageNumber="84">14—</emphasis>
posterior part of the median 2
<emphasis id="BACDEA950D03FF95D274FB00FE09FB05" bold="true" box="[378,403,1240,1262]" pageId="7" pageNumber="84">—</emphasis>
median 3 (paratype, male), note almost complete absence of true pores;
<emphasis id="BACDEA950D03FF95D7A2FB00FB44FB05" bold="true" box="[1196,1246,1240,1262]" pageId="7" pageNumber="84">15—</emphasis>
the scapular plate (Slovakian female);
<emphasis id="BACDEA950D03FF95D269FB2EFE03FAE7" bold="true" box="[359,409,1270,1292]" pageId="7" pageNumber="84">16—</emphasis>
posterior part of the median 1
<emphasis id="BACDEA950D03FF95D1C5FB2EFD7EFAE7" bold="true" box="[715,740,1270,1292]" pageId="7" pageNumber="84">—</emphasis>
median 3 (Slovakian female), note numerous large true pores. Scale bars in µm.
</paragraph>
</caption>
<caption id="DCC6660F0D03FF95D399F848FB88F82F" httpUri="https://zenodo.org/record/1221267/files/figure.png" pageId="7" pageNumber="84" startId="7.[151,264,1936,1958]" targetBox="[151,1436,1372,1913]" targetPageId="7">
<paragraph id="880636870D03FF95D399F848FB88F82F" blockId="7.[151,1436,1936,1988]" pageId="7" pageNumber="84">
<emphasis id="BACDEA950D03FF95D399F848FECFF84D" bold="true" box="[151,341,1936,1958]" pageId="7" pageNumber="84">FIGURES 17–18.</emphasis>
<taxonomicName id="4FB94D040D03FF95D252F848FDD8F84D" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[348,578,1936,1958]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="84" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D03FF95D252F848FDD8F84D" box="[348,578,1936,1958]" italics="true" pageId="7" pageNumber="84">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D03FF95D144F848FD06F84D" box="[586,668,1936,1958]" pageId="7" pageNumber="84" rank="species">
<emphasis id="BACDEA950D03FF95D144F848FD06F84D" bold="true" box="[586,668,1936,1958]" pageId="7" pageNumber="84">sp. nov.</emphasis>
</taxonomicNameLabel>
, male cephalic appendages (paratype, SEM):
<emphasis id="BACDEA950D03FF95D778F848FB32F84D" bold="true" box="[1142,1192,1936,1958]" pageId="7" pageNumber="84">17—</emphasis>
cirrus A with clava, the arrowhead indicates the subterminal constriction;
<emphasis id="BACDEA950D03FF95D19EF876FD58F82F" bold="true" box="[656,706,1966,1988]" pageId="7" pageNumber="84">18—</emphasis>
enlarged clava. Scale bars in µm.
</paragraph>
</caption>
<caption id="DCC6660F0D0CFF9AD399FD2CFF43FCC3" httpUri="https://zenodo.org/record/1221269/files/figure.png" pageId="8" pageNumber="85" startId="8.[151,264,756,778]" targetBox="[151,1436,193,734]" targetPageId="8">
<paragraph id="880636870D0CFF9AD399FD2CFF43FCC3" blockId="8.[151,1435,756,808]" pageId="8" pageNumber="85">
<emphasis id="BACDEA950D0CFF9AD399FD2CFECFFCE1" bold="true" box="[151,341,756,778]" pageId="8" pageNumber="85">FIGURES 19–20.</emphasis>
<taxonomicName id="4FB94D040D0CFF9AD255FD2CFDDBFCE1" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[347,577,756,778]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D0CFF9AD255FD2CFDDBFCE1" box="[347,577,756,778]" italics="true" pageId="8" pageNumber="85">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D0CFF9AD149FD2CFD03FCE1" box="[583,665,756,778]" pageId="8" pageNumber="85" rank="species">
<emphasis id="BACDEA950D0CFF9AD149FD2CFD03FCE1" bold="true" box="[583,665,756,778]" pageId="8" pageNumber="85">sp. nov.</emphasis>
</taxonomicNameLabel>
, claws (SEM):
<emphasis id="BACDEA950D0CFF9AD034FD2CFCF6FCE1" bold="true" box="[826,876,756,778]" pageId="8" pageNumber="85">19—</emphasis>
claws III;
<emphasis id="BACDEA950D0CFF9AD0DBFD2CFB92FCE1" bold="true" box="[981,1032,756,778]" pageId="8" pageNumber="85">20—</emphasis>
claws IV with dentate collar. Scale bars in µm.
</paragraph>
</caption>
<caption id="DCC6660F0D0CFF9AD399F961FBB3F905" httpUri="https://zenodo.org/record/1221271/files/figure.png" pageId="8" pageNumber="85" startId="8.[151,264,1721,1743]" targetBox="[151,1436,873,1700]" targetPageId="8">
<paragraph id="880636870D0CFF9AD399F961FBB3F905" blockId="8.[151,1436,1721,1774]" pageId="8" pageNumber="85">
<emphasis id="BACDEA950D0CFF9AD399F961FEC9F924" bold="true" box="[151,339,1721,1743]" pageId="8" pageNumber="85">FIGURES 21–23.</emphasis>
A semi-schematic illustration of the ontogenetic variability in the ventral armature in
<taxonomicName id="4FB94D040D0CFF9AD7B6F961FA06F924" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[1208,1436,1721,1743]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D0CFF9AD7B6F961FA06F924" box="[1208,1436,1721,1743]" italics="true" pageId="8" pageNumber="85">Bryodelphax instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D0CFF9AD399F900FF7DF905" box="[151,231,1752,1774]" pageId="8" pageNumber="85" rank="species">
<emphasis id="BACDEA950D0CFF9AD399F900FF7DF905" bold="true" box="[151,231,1752,1774]" pageId="8" pageNumber="85">sp. nov.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="BACDEA950D0CFF9AD3FBF900FEBDF905" bold="true" box="[245,295,1752,1774]" pageId="8" pageNumber="85">21—</emphasis>
adult (mature female);
<emphasis id="BACDEA950D0CFF9AD11EF900FDD9F905" bold="true" box="[528,579,1752,1774]" pageId="8" pageNumber="85">22—</emphasis>
juvenile (second instar);
<emphasis id="BACDEA950D0CFF9AD030F900FCEAF905" bold="true" box="[830,880,1752,1774]" pageId="8" pageNumber="85">23—</emphasis>
larva (first instar).
</paragraph>
</caption>
<subSubSection id="C0A3650C0D0CFF9AD3C9F8C3FB44F80C" pageId="8" pageNumber="85" type="description">
<paragraph id="880636870D0CFF9AD3C9F8C3FB44F80C" blockId="8.[151,1436,1819,2024]" pageId="8" pageNumber="85">
<emphasis id="BACDEA950D0CFF9AD3C9F8C3FE5CF8DF" bold="true" box="[199,454,1819,1844]" pageId="8" pageNumber="85">General remarks on</emphasis>
<taxonomicName id="4FB94D040D0CFF9AD2DAF8C3FB2BF8DF" authority="in Polish Pieniny Mts. and Tatra Mts. There" authorityName="in Polish Pieniny Mts. and Tatra Mts. There" box="[468,1201,1819,1844]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="genus">
<emphasis id="BACDEA950D0CFF9AD2DAF8C3FBF8F8DF" bold="true" box="[468,1122,1819,1844]" pageId="8" pageNumber="85">
<emphasis id="BACDEA950D0CFF9AD2DAF8C3FDF2F8DF" bold="true" box="[468,616,1819,1844]" italics="true" pageId="8" pageNumber="85">Bryodelphax</emphasis>
in Polish Pieniny Mts. and Tatra Mts.
</emphasis>
There
</taxonomicName>
are four sympatric
<taxonomicName id="4FB94D040D0CFF9AD399F8E6FEBCF8BC" box="[151,294,1854,1879]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="genus">
<emphasis id="BACDEA950D0CFF9AD399F8E6FEBCF8BC" box="[151,294,1854,1879]" italics="true" pageId="8" pageNumber="85">Bryodelphax</emphasis>
</taxonomicName>
spp. that inhabit
<collectingCountry id="F0AE76170D0CFF9AD2E9F8E6FDAFF8BC" box="[487,565,1854,1879]" name="Poland" pageId="8" pageNumber="85">Poland</collectingCountry>
and occur in the Pieniny Mts. (
<bibRefCitation id="EC284B760D0CFF9AD09DF8E6FBAEF8BC" box="[915,1076,1854,1879]" pageId="8" pageNumber="85" refId="ref9805" refString="Dastych, H. (1988) The Tardigrada of Poland. Monografie Fauny Polski, 16, 1 - 255." type="journal article">Dastych 1988</bibRefCitation>
); three (with the exception of
<taxonomicName id="4FB94D040D0CFF9AD68AF898FE8DF897" baseAuthorityName="Pilato" baseAuthorityYear="1972" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="BACDEA950D0CFF9AD68AF898FE8DF897" italics="true" pageId="8" pageNumber="85">B. weglarskae</emphasis>
</taxonomicName>
) are also present in the Tatra Mts. Our new discovery confirms an eucalciphil and subalpine preference for these Central European members of the genus. In
<collectingCountry id="F0AE76170D0CFF9AD01BF85EFCFCF874" box="[789,870,1926,1951]" name="Poland" pageId="8" pageNumber="85">Poland</collectingCountry>
,
<taxonomicName id="4FB94D040D0CFF9AD07BF85EFB9EF874" box="[885,1028,1926,1951]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="genus">
<emphasis id="BACDEA950D0CFF9AD07BF85EFB9EF874" box="[885,1028,1926,1951]" italics="true" pageId="8" pageNumber="85">Bryodelphax</emphasis>
</taxonomicName>
spp. are frequently found in close proximity. For example, in the Pieniny Mts.
<taxonomicName id="4FB94D040D0CFF9AD19EF874FC88F82F" authorityName="Thulin" authorityYear="1928" box="[656,786,1963,1988]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="species" species="parvulus">
<emphasis id="BACDEA950D0CFF9AD19EF874FC88F82F" box="[656,786,1963,1988]" italics="true" pageId="8" pageNumber="85">B. parvulus</emphasis>
</taxonomicName>
was found in the same moss cushions as
<taxonomicName id="4FB94D040D0CFF9AD7E2F874FAE9F82F" authority="Gąsiorek &amp; Degma, 2018" authorityName="Gąsiorek &amp; Degma" authorityYear="2018" box="[1260,1395,1963,1988]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="species" species="instabilis" status="sp. nov.">
<emphasis id="BACDEA950D0CFF9AD7E2F874FAE9F82F" box="[1260,1395,1963,1988]" italics="true" pageId="8" pageNumber="85">B. instabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1FE57EE0D0CFF9AD674F873FF52F803" pageId="8" pageNumber="85" rank="species">
<emphasis id="BACDEA950D0CFF9AD674F873FF52F803" bold="true" pageId="8" pageNumber="85">sp. nov.</emphasis>
</taxonomicNameLabel>
or
<taxonomicName id="4FB94D040D0CFF9AD3E3F808FE16F803" baseAuthorityName="Pilato" baseAuthorityYear="1972" box="[237,396,1999,2024]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="BACDEA950D0CFF9AD3E3F808FE16F803" box="[237,396,1999,2024]" italics="true" pageId="8" pageNumber="85">B. weglarskae</emphasis>
</taxonomicName>
; similarly,
<taxonomicName id="4FB94D040D0CFF9AD104F808FD11F803" authorityName="Thulin" authorityYear="1928" box="[522,651,1999,2024]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="species" species="parvulus">
<emphasis id="BACDEA950D0CFF9AD104F808FD11F803" box="[522,651,1999,2024]" italics="true" pageId="8" pageNumber="85">B. parvulus</emphasis>
</taxonomicName>
and
<taxonomicName id="4FB94D040D0CFF9AD1CCF808FCDAF80C" baseAuthorityName="Weglarska" baseAuthorityYear="1959" box="[706,832,2000,2023]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="85" phylum="Tardigrada" rank="species" species="tatrensis">
<emphasis id="BACDEA950D0CFF9AD1CCF808FCDAF80C" box="[706,832,2000,2023]" italics="true" pageId="8" pageNumber="85">B. tatrensis</emphasis>
</taxonomicName>
co-occur in mosses in the Tatra Mts.
</paragraph>
</subSubSection>
</treatment>
</document>
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