The true identity of Dixa modesta Johannsen (Diptera: Dixidae) resolved: synonymy of Dixa similis Johannsen, designation of the Dixa ubiquita species group, and description of three new eastern Nearctic species
Author
Moulton, John K.
text
Zootaxa
2017
4216
3
247
260
journal article
37359
10.5281/zenodo.231794
61dd696e-205c-47b2-b5b0-200b46743fc0
1175-5326
231794
4015302B-3141-4281-94E2-4D9966681F70
Dixa ubiquita
species group
Diagnosis.
Adults of the
D
.
ubiquita
group and its closest relatives (i.e.,
Meringodixa
Nowell
spp.,
D. rudis
Garrett
and allied species, and all
Nearctic
congeners except
D
.
hegemonica
Dyar & Shannon
,
D
.
modesta
Johannsen
,
D
.
neohegemonica
Peters
,
D
.
notata
Loew
, and
D
.
terna
Loew
, are readily distinguished from all other
Dixidae
by the solid or indistinctly brown striped (
D
.
brevis
,
D
.
rhathyme
Dyar & Shannon
, and
D
.
lobata
sensu Peters
, particularly females) scutum and presence of katepisternal (= pre-episternum II of authors) setae.
Similarly
colored brown species with basal flagellomeres more cylindrical than fusiform (e.g.,
Dixella indiana
(Dyar)
,
D
.
nigra
(Staeger
)
,
and related species) have a bare katepisternum.
Historically
used characters, i.e., “fusiform vs. cylindrical” basal flagellomeres and “long versus short” apical gonocoxal lobes do not reliably separate adult
Dixa
and
Dixella
Dyar & Shannon
species worldwide, especially in the
Nearctic Region
where nonconforming species exist in each genus.
Within
the aforementioned group having solid or indistinctly striped brown scutal coloration,
D
.
ubiquita
group members are distinguished morphologically from all of these species, except
D. brevis
, by the narrow, falcate/lanceolate gonostylus (in lateral view).
They
differ from
D. brevis
in lacking strong spines (as seen laterally) on the apex of the basal gonocoxal lobe and the cercus being sinuous, posterodorsally directly, and similar in length (i.e., 1.25X) to lateral proctiger width (versus straight, posteroventrally directed, and
2X
lateral width of proctiger).
Extensive
collection data suggest non-overlap between
D
.
brevis
and members of the
D
.
ubiquita
group (
JK Moulton
, unpublished).