The true identity of Dixa modesta Johannsen (Diptera: Dixidae) resolved: synonymy of Dixa similis Johannsen, designation of the Dixa ubiquita species group, and description of three new eastern Nearctic species
Author
Moulton, John K.
text
Zootaxa
2017
4216
3
247
260
journal article
37359
10.5281/zenodo.231794
61dd696e-205c-47b2-b5b0-200b46743fc0
1175-5326
231794
4015302B-3141-4281-94E2-4D9966681F70
Dixa modesta
Johannsen, 1903
: 429
.
(
Figs 3–5
)
Dixa similis
Johannsen, 1923
: 57
.
syn. nov.
Type material.
D. modesta
:
LECTOTYPE
, ♂ labeled: “Ithaca, N.Y./ Apr.”; “OA Johannsen/ Lot 1304/ Sub slides, vial/ Cornell U.”; “HOLOTYPE/ Cornell U./ No. 2992” [red label]”; “HOLOTYPE/
Dixa
/
modesta Joh
[red-trimmed label]”; “Lectotype/
Dixa modesta
/ Johannsen/ Fixation by
T.M. Peters, 1968
/ (labeled:
J.K. Moulton, 2016
)” (CUIC).
D. similis
:
HOLOTYPE
, ♂ labeled: “Ithaca, N.Y.
28 Aug.
[18]’94”; “OA Johannsen/ Lot 1293 Sub vial/ Cornell U.”; “HOLOTYPE/
Dixa
/
similis
/ Johannsen [red-trimmed label]”; “HOLOTYPE/ Cornell U./ No. 2993”.
ALLOTYPE,
♂ labeled: “Ithaca, N.Y./
28 Aug.
[18]’94”; “OA Johannsen/ Lot 85/ Sub. 1147/ Cornell U.”; “ALLOTYPE/ Cornell U./ No. 2993”; “ALLOTYPE
Dixa similis Johannsen
” (CUIC).
Remarks.
In 1954, Alice Dietrich, Cornell University Insect Museum, accessioned
holotype
#2992 (
Figs 3–5
), the pinned “
holotype
” male of
Dixa modesta
(J. Liebherr, pers. comm.). This male, clearly having a striped scutum (
Figs 3–4
), corresponds to the adult successfully reared by
Johannsen (1903)
from a larva (and pupa, the exuviae of both included in the
syntype
series) partly serving as the basis for Johannsen ′s original description. Examination of the
lectotype
and additional material studied by Johannsen confirmed the presence of a mixed series. Examination of the primary
types
of
D. modesta
and
D. similis
,
including the badly faded terminalia of the former, confirmed they are of the same nominal species. Therefore,
D. similis
falls as an objective junior synonym of
D. modesta
. This species will be re-described in a future study.
Examination of Johannsen’s material, in addition to the
lectotype
, revealed one male (actually only partial terminalia in glycerin) each of
D. ubiquita
sp. nov.
from Ithaca, NY and Black Mtn, NC, one male of
Dixa brevis
(terminalia in glycerin), and one larva from Cascadilla Creek near Ithaca. Confirmation of
D
.
ubiquita
sp. nov.
from the Ithaca area fits nicely with the hypothesized, but unidentified, collection locality, Fall Creek (Flat Rock), a karst stream now largely contained within a Cornell Plantations Natural Area (Jason Dombroskie, pers. comm.). The larva from Cascadilla Creek could be either
D. ubiquita
sp. nov.
or
D
.
vockerothi
sp. nov.
based upon photographs of the stream, which has numerous staircase falls within a gorge.