Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae)
Author
Constant, Jérôme
6E6072A1-9415-4C8D-8E60-2504444DB290
Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier Street 29, B- 1000 Brussels, Belgium.
jerome.constant@naturalsciences.be
Author
Semeraro, Linda
DCAAD2E7-80F7-4EA9-98D9-167FE8B97C58
Department of Jobs, Precincts and Regions, Agriculture Victoria, Agribio, Centre for Agribioscience, 5 Ring Road, Bundoora, 3083, Victoria, Australia.
linda.semeraro@agriculture.vic.gov.au
Author
Moir, Melinda L.
8361DF9D-50F4-4DD0-906B-7E7D3F7983B9
Department of Primary Industries and Regional Development, South Perth, Western Australia 6151, Australia.
melinda.moir@dpird.wa.gov.au
text
European Journal of Taxonomy
2022
2022-09-07
836
66
95
http://zoobank.org/132e46b2-b8f1-48c3-affd-2d0e48e2606d
journal article
139208
10.5852/ejt.2022.836.1917
ce64819a-d53f-4a40-9cd5-d4efc3d251d1
2118-9773
7061621
132E46B2-B8F1-48C3-AFFD-2D0E48E2606D
Tribe
Cleotychini
Emeljanov, 1997
Diagnosis
Habitus reminiscent of
Caliscelidae
(
Hemiptera
: Fulgoromorpha), with pyriform body shape and the anterior part of body slightly compressed. Head with well delimited trigons (triangular area delimited by carinae at anterior angles of vertex). Frons approximately rectangular, parallel-sided, much longer than wide. Vertex subquadrate, longer than wide. Apical segment of the labium half as long as subapical segment. Pronotum elongate (slightly shorter than vertex along mid-line), dorsally tricarinate, with a single lateral keel on each side and posterior margin nearly straight. Mesonotum smooth, carinae nearly obsolete. Tegmina strongly reduced, without carina; clavus not defined. Tegulae absent. Dorsum of abdomen without carinae. Procoxae with outer carina extended into triangular apical lobe. Profemora and protibiae flattened and dilated, often foliate. Metatibiae with two lateral spines, one near tibiofemoral joint and one in distal half of tibia. Apex of metatibiae with 6 teeth separated in two groups: 2 teeth on internal side and 4 on external side. First and second segments of metatarsus with platellae instead of spines, except large lateral common teeth without setae. Arolium with one pair of chetoid sensilla. Claws with 3 setae. Sternite VIII of males fused with pygofer. Pygofer with broad base, narrowed backwards in ventral view. Gonostyli without dorsal teeth (such as in
Orthopagus
Uhler, 1897 species
for example) but with dorsal lobe bearing a dorsolateral hook. Connective fused with lower wall of pygofer, largely connected with base of gonostyli. Lateral lobe of gonoplacs (third valvae of ovipositor) without appendix; medial lobe of gonapophyses VIII (first valvae of ovipositor) with two nervures; lateral lobes with bidentate apex.
Remark
The tribe
Cleotychini
is here conservatively retained within the family
Dictyopharidae
until more detailed phylogenetic studies confirm or refute the familial placement (see also discussion below).