Revision of the Mesoamerican species of Calolydella Townsend (Diptera: Tachinidae) and description of twenty-three new species reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica Author Fleming, AJ Author Wood, D. Monty Author Smith, M. Alex Author Hallwachs, Winnie Author Janzen, Daniel H text Biodiversity Data Journal 2018 6 11223 11223 http://dx.doi.org/10.3897/BDJ.6.e11223 journal article http://dx.doi.org/10.3897/BDJ.6.e11223 1314-2828--11223 Calolydella discalis Fleming & Wood sp. n. Materials Type status: Holotype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017774 ; recordedBy: D.H. Janzen, W. Hallwachs & Androny Valdelomar ; individualID: DHJPAR0017774; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-196, BOLD:AAI6844, ASTAR485-07; Taxon: scientificName: Calolydelladiscalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: discalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Rio Blanco Abajo; verbatimElevation: 500 ; verbatimLatitude: 10.9004; verbatimLongitude: -85.3725; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Turuptianaobliqua ; verbatimEventDate: 04-Feb-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017773 ; recordedBy: D.H. Janzen, W. Hallwachs & Androny Valdelomar ; individualID: DHJPAR0017773; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-211, BOLD:AAI6844, ASTAR484-07; Taxon: scientificName: Calolydelladiscalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: discalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Rio Blanco Abajo; verbatimElevation: 500 ; verbatimLatitude: 10.9004; verbatimLongitude: -85.3725; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Turuptianaobliqua ; verbatimEventDate: 03-Feb-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Description Male (Fig. 13a, b, c). Length: 8-9mm. Head (Fig. 13b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, with a single row of short setulae outside of frontal setae; parafacial at least 50% silver pollinose. Thorax (Fig. 13a, c): gold on both dorsal and lateral surfaces; outermost two thoracic vittae twice as wide as innermost two; scutellar discal setae situated as wide apart as subapical scutellar setae; postpronotum with three setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 3:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft. Wing vein R4+5 with 4-5 small setulae dorsally at base. Abdomen (Fig. 13a): ground color dark brown-orange with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, and with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae pair and two pairs of discal setae; T4 with two pairs of discal setae. Terminalia (Fig. 14): sternite 5 (Fig. 14c) with two small lobes and a wide U-shaped median cleft, 0.37X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite covered with short setae, all of equal length. Cerci (Fig. 14b), in dorsal view, narrow and tapered, separated by a narrow gap widening at apex; cercus long, slender and straight when viewed laterally, very slightly tapered from its already narrow base; setose along its basal half. Surstylus (Fig. 14a) subequal to length to cercus, slender and digitiform when viewed laterally; with short setae along entire length; tip of surstylus not curved inwards when viewed dorsally. Female: not known at this time. Diagnosis Calolydella discalis can be distinguished from all other species of Calolydella by the following combination of traits: parafacial mostly gold, fronto-orbital plate with a single row of small black setulae outside of the frontal setae, anatergite with a small tuft of three or more hair-like setae, and T3 and T4 each with two pairs of discal setae. Etymology The specific epithet refers to the two pairs of discal setae on T3 and T4. Distribution Costa Rica, ACG, Alajuela Province, Rio Blanco Abajo, 500m. Ecology Calolydella discalis has been reared twice from Turuptiana obliqua Walker, 1869 ( Lepidoptera : Erebidae : Arctiinae ), in rain forest.