On Venezuelan pholcid spiders (Araneae, Pholcidae)
Author
Huber, Bernhard A.
33607F65-19BF-4DC9-94FD-4BB88CED455F
Zoological Research Museum Alexander Koenig, Bonn, Germany.
b.huber@leibniz-zfmk.de
Author
Villarreal, Osvaldo
679C385E-B068-4351-9D2F-97753E534C26
Museo del Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela. & Museu Nacional / UFRJ, Rio de Janeiro, Brazil.
osvaldovillarreal@gmail.com
text
European Journal of Taxonomy
2020
2020-10-01
718
1
317
journal article
10.5852/ejt.2020.718.1101
4069574
F9E9A91E-488C-4DB1-9361-E788E9AC5BC1
Systenita prasina
Simon, 1893
Figs
1008–1020
,
1021–1022
, 1065
Systenita prasina
Simon, 1893a: 318
.
Systenita prasina
–
Simon 1893b: 479–483
. —
Brignoli 1975: 36
, fig.
2g
. —
Huber 1997d: 608
, figs 28a–d, 29a–f. —
González-Sponga 2010: 23
, pl. 6, figs 1–9.
Misidentification
Systenita prasina
–
Caporiacco 1955: 299
, fig. 8 (see
Huber 1997d
).
Notes
Brignoli’s (1975)
drawing of the palp of this species was probably prepared from a
type
specimen in MNHN. He did
not
specify the origin of his specimen(s).
González-Sponga’s (2010)
material was reexamined and is correctly identified: 1 ♂,
1 ♀
, MIZA 105656 (MAGS 1347), La
Guaira
, “carretera El Junquito – Carayaca” (file card), “El Junquito – vía Carayaca” (publication) [approximately
10.445° N
,
67.147° W
],
11 Jan. 1992
(A.R. Delgado, M.A. González S.).
Diagnosis
Easily distinguished from similar pale six-eyed pholcids (
Mecolaesthus fallax
Huber
sp. nov.
,
Metagonia
spp.) by distinctive armature of male chelicerae (
Huber 1997d
: figs 29e–f; two large clubshaped hairs on each side), by very slender procursus partly lodged in groove of large bulbal process (
Huber 1997d
: fig. 29b), and by epigynum and female internal genitalia (Figs
1012–1020
; small light brown sclerite, tiny globular pore plates attached to median sclerite).
Type
material
VENEZUELA
–
Aragua
•
♂
lectotype
(designated in
Huber 1997d
),
8 ♂♂
,
16 ♀♀
paralectotypes
and
7 juvs
,
MNHN
(Ar 10527), Eugène Simon collection number 11023, Colonia Tovar [approximately
10.41° N
,
67.29° W
],
Jan.–Feb. 1888
(
E. Simon
); examined (see
Huber 1997d
).
New records
VENEZUELA
–
Aragua
•
9 ♂♂
,
15 ♀♀
,
ZFMK
(Ar 22137–38), and
2 ♂♂
,
2 ♀♀
,
3 juvs
in pure ethanol,
ZFMK
(Ven02/100-11),
forest above Colonia Tovar
(
10.414° N
,
67.301° W
), ~
2100 m
a.s.l.
,
26 Nov. 2002
(
B.A. Huber
)
•
9 ♂♂
,
11 ♀♀
,
ZFMK
(Ar 22139–40), and
1 ♂
,
5 ♀♀
in pure ethanol,
ZFMK
(Ven18-152), same locality,
2140 m
a.s.l.
,
8 Nov. 2018
(
B.A. Huber
,
O. Villarreal
M.)
•
9 ♂♂
,
7 ♀♀
,
ZFMK
(Ar 22141–43), and
1 ♂
,
2 ♀♀
,
1 juv.
in pure ethanol,
ZFMK
(Ven02/100-45),
near Colonia Tovar
,
forest at Cerro Picacho
(= Pico Codazzi) (
10.408° N
,
67.308° W
), ~
2200 m
a.s.l.
,
27 Nov. 2002
(
B.A. Huber
). –
La Guaira
•
1 ♂
,
2 ♀♀
,
ZFMK
(Ar 22144),
between Colonia Tovar and El Junquito
(
10.4230° N
,
67.2381° W
),
1960 m
a.s.l.
,
10 Nov. 2018
(
B.A. Huber
,
O. Villarreal
M.). –
Miranda
•
15 ♂♂
,
14 ♀♀
,
ZFMK
(Ar 22145–46),
El Ávila National Park
,
between Sabas Nieves and La Silla
(
10.529° N
,
66.854° W
), ~
2100 m
a.s.l.
,
25 Nov. 2002
(B.A.
Huber
)
•
21 ♂♂
,
6 ♀♀
,
2 juvs
,
ZFMK
(Ar 22147–48), and
3 ♂♂
,
1 ♀
in pure ethanol,
ZFMK
(Ven18-143), same locality (
10.5288° N
,
66.8546° W
),
1850 m
a.s.l.
,
7 Nov. 2018
(
B.A. Huber
,
O. Villarreal
M.)
•
2 ♀♀
,
1 juv.
,
MIZA 105805
(
MAGS 561
),
El Ávila National Park
,
El Paraíso
[
10.530° N
,
66.819° W
], ~
1500 m
a.s.l.
,
16 Aug. 1981
(
J.A. González
D.)
.
Redescription
(amendments, see
Huber 1997d
)
Male
Habitus as in Figs
1008–1009
. Eye measurements (male from Colonia Tovar): distance PME–PME
210 µm
; diameter PME
80 µm
; distance PME–ALE
70 µm
. Carapace monochromous whitish, clypeus variable, from pale grey to light brown to black; legs pale ochre yellow, femora 2 and 3 dorsally proximally with long dark mark, coxae 2 and 3 ventrally with dark mark; patellae and tibia-metatarsus joints dark; abdomen monochromous pale gray to pale bluish. Legs without curved hairs; retrolateral trichobothrium on tibia 1 at 1.5%; prolateral trichobothrium present on tibia 1. Tibia
1 in
56 newly examined males: 6.5–8.1 (mean 7.3).
Female
Colors as in male (Figs
1010–1011
), but femora 2 and 3 dorsally without dark mark and coxae 2 and 3 ventrally without dark mark; color of clypeus variable as in male. Tibia
1 in
50 newly examined females: 4.9–6.1 (mean 5.4). Epigynum (Figs
1012–1015
) very simple, ventral view variable depending on visibility of internal (often greenish or bluish) structures. Internal genitalia (Figs
1016–1020
) with tiny globular pore plates attached to median sclerite.
Distribution
Known from several localities in the Coastal Ranges (between
1850 and
2200
m a.s.l.) in the Venezuelan states
Aragua
, La
Guaira
, and
Miranda
(Fig. 1065).
Natural history
The spiders were collected from their relatively large webs attached to the undersides of leaves at ~
1–2 m
above the ground. They rested in an inverted position (dorsal side of the abdomen facing the underside of the leaf, frontal side of prosoma facing the forest floor; Figs 1008, 1010) at the apex of the domed web where it was closely attached to the leaf. When disturbed, the spiders moved away and bobbed or vibrated but were easy to catch. Egg sacs were often greenish (Figs
1010–1011
).