Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group Author Golovatch, Sergei I. Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru sgol@orc.ru Author Geoffroy, Jean-Jacques Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr geoffroy@mnhn.fr Author Mauriès, Jean-Paul Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr collmill@mnhn.fr Author Spiegel, Didier Van Den Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be text Zoosystema 2007 29 3 417 456 journal article 10.5281/zenodo.5393630 1638-9387 5393630 Glyphiulus obliteratoides n. sp. ( Figs 9-11 ) TYPE MATERIAL . — China . Ghizhou Prov. , Anshun County , Luangshuijing , Tian Xian Dong Cave (aquatic), 22.II.2004 , leg. S. Prevorčnik & B. Sket , holotype ♂ ( MNHN GA 047 ) ; paratypes 1 ♂ , 1 ♀ , 4 juvs ( MNHN GA 047 ) ; 1 ♂ , 1 ♀ , 2 juvs ( ZMUM ) ; 1 ♀ ( OBBFUL ) ; 1 ♀ ( SEM ) ; 1 subad. ♀ ( ZMUC ). — Same cave , 23.II.2004 , leg. B. Sket & S. Prevorčnik , 1 juv. ( OBBFUL ) . FIG. 10. — Glyphiulus obliteratoides n. sp. , paratype ♀: A , telson, ventral view; B , midbody segment section; C , claw of midbody leg; D , claw. Scale bars: A, 0.2 mm; B, 0.5 mm; C, 0.05 mm; D, 0.02 mm. ETYMOLOGY. — To emphasize the very close resemblance to G. obliteratus n. sp. DIAGNOSIS. — Differs from congeners by the pigmented ocelli and body, coupled with the more distinctly undulate collum, the undivided crests, etc. (see also key below). DESCRIPTION Holotype with 59p+1a+ T , about 32 mm long and 1.9 mm wide; adult paratypes ( ♂ , ♀ ) with 45-60p+2- 1a+ T , length 22-35 mm , width 1.3-1.9 mm . Coloration entirely pallid in young juvenile stadia, becoming marbled red-brown in adults, tergal crests dark castaneous brown; antennae, legs and venter light red-brown in adults. Adults with 7-9 blackish ocelli in a subtriangular eye patch. Antennal sensilla, segment shape ( Fig. 10B ), tergal crests (including ozoporiferous ones) ( Figs 9 ; 10B ), epiproct ( Fig. 9E, F ), paraprocts ( Fig. 9E ), male legs 2 and 3 ( Fig. 11D, E ), penes ( Fig. 11D ), gonopods ( Fig. 11G, H ) generally as in G. obliteratus n. sp. , but antennae ( Figs 9A ; 11A ) slightly shorter, gnathochilarium (mentum divided, n = 3) ( Fig. 11B ) more polytrichous; collum ( Fig. 9A, B ) with 1a-4a+ma+4a-1a undulations at caudal margin, undulation ma being shortest; hypoproct ( Fig. 10A ) very faintly emarginate caudomedially, conspicuous pleurosternal crests on segments 2-4 wanting ( Fig. 9A ); tergal crests similarly low and undivided, even adding another lateral crest below ozoporiferous one, which is barely traceable as a low anterior tubercle, but anterior end of all dorsal crests broadened ( Fig. 9 ); carinotaxy pattern thus 1/0+1+I/i+3+I/i+1+1/0 ( Figs 9C, D ; 10B ); a small accessory spine present at base of claw on legs 2 and 3 only ( Fig. 11D ), onward nearly missing ( Figs 10B ; 11F ); legs about as long as body diameter ( Fig. 10B ); male leg 1 with 5-segmented but shortened telopodites ( Fig. 11C ). FIG. 11. — Glyphiulus obliteratoides n. sp. , paratype ♂: A , antenna; B , gnathochilarium; C , leg 1, frontal view; D , leg 2, caudal view; E , leg 3; F , leg 9; G , anterior gonopods, caudal view; H , right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm. Head width = segment 2 = 6 <collum = 7(8)> segment 3 = 5 <7 <8 <9 <10 <11 = midbody segments; body gradually tapering toward telson. Postcollar constriction evident ( Fig. 9B ). REMARKS The samples contain five haplopodous juveniles with 30-33p+1-3a+T, each with the ozoporiferous tubercles fully developed only on segments 5 and 6. In general, even juveniles of later, diplopodous, stadia are easily recognised due to the ozoporiferous cones being especially well developed on segments 5 and 6 only. In the absence of any troglomorphic traits, this species can only be considered troglophilic at most.