Contributions to the systematics of the genera Dipseudopsis, Hyalopsyche and Pseudoneureclipsis (Trichoptera: Dipseudopsidae), with descriptions of 19 new species from the Oriental Region.
Author
Oláh, János
Author
Johanson, Kjell Arne
text
Zootaxa
2010
2658
1
37
journal article
10.5281/zenodo.198974
9be137a0-e57e-4b73-9dfc-681b65a34c5a
1175-5326
198974
Hyalopsyche
Ulmer
Hyalopsyche
Ulmer, 1904
: 357
.
Type
species:
Hyalopsyche palpata
Ulmer
(monotypic);
type
locality: "
French Congo
" (Ogowe Faktorei).
Hyalopsychodes
Betten, 1909
: 237–238.
Type
species:
H. rivalis
Betten
(monotypic);
type
locality:
India
(
East Bengal
). Synonym according to
Martynov, 1914
: 20.
Hyalopsychella
Ulmer, 1930
: 422
.
Type
species:
H. winkleri
Ulmer
(monotypic);
type
locality:
Malaysia
(Borneo).
New Synonym
.
This genus is known from the Palaearctic, Oriental, Australasian and Afrotropical Biogeographical Regions. All species have similar cephalic, thoracic and genital characters.
Adult. The cephalic and thoracic groove and setal wart patterns are very stable and persistent in all examined species and we present the head structures of
Hyalopsyche sachalinica
Martynov
to represent the generic characteristics.
Head: Tentorium without dorsal arms and vestigial structures; rounded broadened flange present mesoventrally on middle of anterior arms. Frontogenal septa (i.e., internal folds of frontogenal and clypeogenal grooves) well-developed, with dorsal frontogenal arms extending dorsad from anterior tentorial pits to margin of circumantennal sclerites; posterior tentorial arms very short, extending and giving support to internal occipital ridges; tentorial bridge shifted posterad to posterior tentorial pits. Maxillary palps reduced in size (i.e., abbreviated to length of facial area) and adhering to frontogenal and clypeogenal surfaces in resting position; segment 5 of each maxillary palp shortened and annulations reduced. Laciniae reduced in size. Labial palps absent. Labrum with elongate, freely hanging apex. Single facial groove (short and fused frontal groove) visible only on face, its fusion being mediated by enlargement of scapes pressing interantennal area into narrow strip; coronal groove well-developed and visible along almost entire vertex. Central clypeal wart and labral posterior central warts weakly discernible on face and covered only scarcely with minute setae. Vertexal lateroantennal, occipital and postgenal compact setose warts present and visible on head dorsum. Vertexal medioantennal wart visible in limited area compressed by antennal groove and circumantennal sclerites of enlarged scapes.
Thorax: Pronotum and mesonotum variable. Spur formula 3(2), 4, 4. Number of apical forks in the forewings and hind wings variable among species.
Remarks.
The description of
Hyalopsyche
was based originally on
H. palpata
Ulmer
on the basis of having reduced wing venation, with reduced number of apical forks in the forewing and absence of hind wing forks I and IV.
Marlier (1962)
reported that forewing fork I and hind wing forks I, III and IV are absent in
H. palpata
Ulmer. New
monotypic genera related to
Hyalopsyche
have been established based primarily on the presence or absence of apical wing forks. The genus
Hyalopsychodes
was described by
Betten (1909)
based on presence of forewing fork I. This genus was synonymised with
Hyalopsyche
by
Martynov (1914)
. The genus
Hyalopsychella
was distinguished by the presence of a short forewing discoidal cell and the absence of forewing fork I and hind wing forks I and III. We observed a range in the length of the discoidal cell among species in the genus
Hyalopsyche
, being longest in
H. trunga
,
new species
and shortest in
H. winkleri
(Ulmer)
. Below we describe
H. trunga
,
new species
from
Vietnam
, having only hind wing forks II and V present, and with a long discoidal cell in the forewings. The loss of apical forks, especially in the hind wings, occurs in some species, but these species are otherwise similar to species with intact wing forks, particularly in male genitalia. As the character states used for separating
Hyalopsychella
and
Hyalopsyche
are found also in
Hyalopsyche
, the genus
Hyalopsychella
is synonymised with
Hyalopsyche
.