The first fossil spider cricket (Orthoptera: Gryllidae: Phalangopsinae): 20 million years of troglobiomorphosis or exaptation in the dark? Author Heads, Sam W. text Zoological Journal of the Linnean Society 2010 2010-01-31 158 1 56 65 http://dx.doi.org/10.1111/j.1096-3642.2009.00587.x journal article 10.1111/j.1096-3642.2009.00587.x 0024-4082 4720036 TRIBE LUZARINI HEBARD, 1928 Luzarae Hebard, 1928: 1 . Luzarini Hebard, 1928: 1 . Luzarinae Desutter-Grandcolas, 1993: 1 . Type genus: Luzara Walker, 1869 . Remarks: Desutter-Grandcolas (1992: 95) defined the Luzarini based on the morphology of the male genitalia, stating that ‘chez les Luzarinae [ sic ], toute la partie médiane de l’épiphalle est régressée, notamment les lophi médians; d’autre part, les “bras épiphalliques” ainsi formés acquièrent une structure complex, avec généralement une ou deux zones de rupture ou d’articulation et des épines latérales’. The retrograde median region of the epiphallus is unique amongst Luzarini , and is likely to be apomorphic. The complex ‘bras épiphalliques’, or ‘epiphallic arms’, and their lateral lobes are highly variable: their morphology and spatial associations with the epiphallic parameres have proved very useful at lower taxonomic levels (e.g. Desutter-Grandcolas, 1992 , 1993 ; Desutter-Grandcolas & Otte, 1997 ).