<document id="9A9BAC3282D52207188FBBC98C7D5B98" ID-DOI="10.5281/zenodo.5353792" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353792" ID-ZooBank="88D45C5E-0E49-4F02-87A4-C4FA55A83B2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212659467" checkinUser="felipe" docAuthor="Freese, Edwin L., Veal, Doug A. &amp; Lago, Paul K." docDate="2020" docId="28608792FF9B3666FF76FF3BB919FD5E" docLanguage="en" docName="0787.pdf" docOrigin="Insecta Mundi 2020 (787)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Phanaeus vindex MacLeay 1819" docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="D459FFEAFF86367BFFB6FFEDBB37FF84" masterDocTitle="The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist" masterLastPageNumber="83" masterPageNumber="1" pageNumber="28" updateTime="1699188000474" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="0D8D1CBD0EA3C2E91F886C57CC7042B8">The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist</mods:title>
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<mods:namePart id="894C3FFD993C081DEFA8AF750614032D">Freese, Edwin L.</mods:namePart>
<mods:affiliation id="E26A07554645F5FB06F8EA4BACA61CC8">33493 “ S ” Avenue Adel, Iowa 50003</mods:affiliation>
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<mods:namePart id="D6B2329A935B1F95019C988A8224AC1B">Veal, Doug A.</mods:namePart>
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<mods:namePart id="867D7239AE02A3AF3D340585B1CBA57D">Lago, Paul K.</mods:namePart>
<mods:affiliation id="2BBBCD47706F08C9BB7B6B834D35EE5B">Biology Department, University of Mississippi University, Mississippi 38677</mods:affiliation>
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<mods:identifier id="45F224F4A4C0562208F35CAF69431F48" type="ISSN">1942-1354</mods:identifier>
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<heading id="FB3E81E8FF9B3666FF76FF3BB94EFF6B" bold="true" box="[192,633,214,239]" fontSize="10" level="2" pageId="29" pageNumber="28" reason="3">
<taxonomicName id="67C94D07FF9B3666FF76FF3BB94EFF6B" ID-CoL="4FSL9" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[192,633,214,239]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="vindex">
<emphasis id="92BDEA96FF9B3666FF76FF3BB94EFF6B" bold="true" box="[192,633,214,239]" pageId="29" pageNumber="28">
<emphasis id="92BDEA96FF9B3666FF76FF3BBA9DFF6A" bold="true" box="[192,426,214,238]" italics="true" pageId="29" pageNumber="28">Phanaeus vindex</emphasis>
MacLeay, 1819
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<paragraph id="A0763684FF9B3666FF76FEE9BA58FEBD" blockId="29.[192,1440,260,313]" pageId="29" pageNumber="28">
<bibRefCitation id="C4584B75FF9B3666FF76FEE9BA4AFE98" author="Wickham, H. F." box="[192,381,260,284]" pageId="29" pageNumber="28" pagination="1 - 40" refId="ref34735" refString="Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40." type="journal article" year="1911">Wickham 1911</bibRefCitation>
; 
<bibRefCitation id="C4584B75FF9B3666FE3AFEE9B9AFFE98" author="Blume, R. R. &amp; A. Aga" box="[396,664,260,284]" pageId="29" pageNumber="28" pagination="113 - 120" refId="ref30713" refString="Blume, R. R., and A. Aga. 1978. Observation on ecological and phylogenetic relationships of Phanaeus difformis LeConte and Phanaeus vindex MacLeay (Coleoptera: Scarabaeidae) in North America. Southwestern Entomologist 3 (2): 113 - 120." type="journal article" year="1978">Blume and Aga 1978</bibRefCitation>
; 
<bibRefCitation id="C4584B75FF9B3666FD11FEE9B854FE98" author="Edmonds, W. D." box="[679,867,260,284]" pageId="29" pageNumber="28" pagination="1 - 105" refId="ref31397" refString="Edmonds, W. D. 1994. Revision of Phanaeus MacLeay, a New World Genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae). Natural History Museum of Los Angeles County, Contributions in Science 443: 1 - 105." type="journal article" year="1994">Edmonds 1994</bibRefCitation>
; 
<bibRefCitation id="C4584B75FF9B3666FCC4FEE9BFC7FE98" author="Worthington, R. J. &amp; K. J. Larsen" box="[882,1264,260,284]" pageId="29" pageNumber="28" pagination="77 - 90" refId="ref34793" refString="Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90." type="journal article" year="2010">Worthington and Larsen 2010</bibRefCitation>
; ISIC; IWUC; DAVC; ELFC.
</paragraph>
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<paragraph id="A0763684FF9B3666FF76FEA5B97DFEE4" blockId="29.[192,586,328,352]" box="[192,586,328,352]" pageId="29" pageNumber="28">Collection dates: April–October.</paragraph>
<paragraph id="A0763684FF9B3666FF76FE82B949FE03" blockId="29.[192,638,367,391]" box="[192,638,367,391]" pageId="29" pageNumber="28">Collected from deer and horse dung.</paragraph>
<paragraph id="A0763684FF9B3666FF76FE7ABA61FE6C" blockId="29.[192,1443,407,488]" pageId="29" pageNumber="28">
<materialsCitation id="10A13CD9FF9B3666FF76FE7ABAC4FE48" collectionCode="ISIC" collectorName="Osborn &amp; W. D. Edmonds" county="Story Co." location="Experiment Station" municipality="Specimen" pageId="29" pageNumber="28" specimenCount="1" stateProvince="Iowa">
<collectingMunicipality id="4012ACFEFF9B3666FF76FE7ABA00FE2B" box="[192,311,407,431]" pageId="29" pageNumber="28">Specimen</collectingMunicipality>
label data: 
<location id="A516605FFF9B3666FE78FE7AB9F0FE2B" LSID="urn:lsid:plazi:treatment:28608792FF9B3666FF76FF3BB919FD5E:A516605FFF9B3666FE78FE7AB9F0FE2B" box="[462,711,407,431]" county="Story Co." municipality="Specimen" name="Experiment Station" pageId="29" pageNumber="28" stateProvince="Iowa">Experiment Station</location>
, 
<location id="A516605FFF9B3666FD60FE7AB829FE2B" LSID="urn:lsid:plazi:treatment:28608792FF9B3666FF76FF3BB919FD5E:A516605FFF9B3666FD60FE7AB829FE2B" box="[726,798,407,431]" county="Story Co." municipality="Specimen" name="Ames" pageId="29" pageNumber="28" stateProvince="Iowa">Ames</location>
, 
<collectingCounty id="49174E08FF9B3666FC9DFE7AB895FE2B" box="[811,930,407,431]" pageId="29" pageNumber="28">Story Co.</collectingCounty>
, 
<collectingRegion id="620DF866FF9B3666FC06FE7AB8D8FE2A" box="[944,1007,407,430]" country="United States of America" name="Iowa" pageId="29" pageNumber="28">Iowa</collectingRegion>
, [no date], 
<collectorName id="0D3C5352FF9B3666FBCBFE7ABFEEFE2B" box="[1149,1241,407,431]" pageId="29" pageNumber="28">Osborn</collectorName>
, determined 
<collectorName id="0D3C5352FF9B3666FAC8FE7ABA6DFE48" pageId="29" pageNumber="28">W. D. Edmonds</collectorName>
1983 (
<collectionCode id="C6D8AE41FF9B3666FE1BFE59BADDFE48" box="[429,490,436,460]" pageId="29" pageNumber="28">ISIC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="10A13CD9FF9B3666FE49FE59BA64FE6C" collectingDate="1932-06-15" collectionCode="ISIC" collectorName="Travis &amp; W. D. Edmonds" county="Decatur Co." location="Leon" pageId="29" pageNumber="28" specimenCount="1" stateProvince="Iowa">
<location id="A516605FFF9B3666FE49FE59B908FE4F" LSID="urn:lsid:plazi:treatment:28608792FF9B3666FF76FF3BB919FD5E:A516605FFF9B3666FE49FE59B908FE4F" box="[511,575,436,459]" county="Decatur Co." name="Leon" pageId="29" pageNumber="28" stateProvince="Iowa">Leon</location>
, 
<collectingCounty id="49174E08FF9B3666FDFDFE59B9D6FE48" box="[587,737,436,460]" pageId="29" pageNumber="28">Decatur Co.</collectingCounty>
, 
<collectingRegion id="620DF866FF9B3666FD58FE59B81AFE4F" box="[750,813,436,459]" country="United States of America" name="Iowa" pageId="29" pageNumber="28">Iowa</collectingRegion>
, 
<date id="D4771044FF9B3666FC8FFE5EB8D6FE48" box="[825,993,435,460]" pageId="29" pageNumber="28" value="1932-06-15">
<collectingDate id="C433E9ACFF9B3666FC8FFE5EB8D6FE48" box="[825,993,435,460]" pageId="29" pageNumber="28" value="1932-06-15">15 June 1932</collectingDate>
</date>
, 
<collectorName id="0D3C5352FF9B3666FC5BFE59BF08FE48" box="[1005,1087,436,460]" pageId="29" pageNumber="28">Travis</collectorName>
, determined 
<collectorName id="0D3C5352FF9B3666FB57FE59BE94FE48" box="[1249,1443,436,460]" pageId="29" pageNumber="28">W. D. Edmonds</collectorName>
1983 (
<collectionCode id="C6D8AE41FF9B3666FEBAFE3DBA7DFE6C" box="[268,330,464,488]" pageId="29" pageNumber="28">ISIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A0763684FF9B3666FF76FE15BAD0FDE2" blockId="29.[192,1440,504,614]" pageId="29" pageNumber="28">
County records (
<figureCitation id="38F22A01FF9B3666FE20FE15B921FD94" box="[406,534,504,528]" captionStart="Figures 90–97" captionStartId="72.[192,287,1847,1869]" captionTargetBox="[147,1439,217,1801]" captionTargetPageId="72" captionText="Figures 90–97. Distribution of Scarabaeoidea in Iowa. 90) Copris fricator fricator (Fabricius, 1787). 91) Copris minutus (Drury, 1770). 92) Dichotomius carolinus (Linnaeus, 1767). 93) Phanaeus vindex MacLeay, 1819. 94) Canthon chalcites (Haldeman, 1843). 95) Canthon pilularius (Linnaeus, 1758). 96) Canthon vigilans LeConte," figureDoi="http://doi.org/10.5281/zenodo.5353820" httpUri="https://zenodo.org/record/5353820/files/figure.png" pageId="29" pageNumber="28">Figure 93</figureCitation>
): Allamakee, Appanoose, Boone, Clarke, Clayton, Davis, Decatur, Des Moines, Dickinson, Fayette, Fremont, Henry, Jefferson, Keokuk, Lee, Linn, Louisa, Madison, Marion, Mills, Monroe, Muscatine, Page, Pottawattamie, Story, Van Buren, Wapello, Warren, 
<collectingRegion id="620DF866FF9B3666FAB2FDDCBEABFDCD" box="[1284,1436,561,585]" country="United States of America" name="Washington" pageId="29" pageNumber="28">Washington</collectingRegion>
, Winneshiek, Woodbury.
</paragraph>
<paragraph id="A0763684FF9B3666FF76FD4BB9A1FD3A" blockId="29.[192,662,678,730]" box="[192,662,678,702]" pageId="29" pageNumber="28">Tribe CANTHONINI Lansberge, 1874</paragraph>
<paragraph id="A0763684FF9B3666FF76FD2FB919FD5E" blockId="29.[192,662,678,730]" box="[192,558,706,730]" pageId="29" pageNumber="28">
<taxonomicName id="67C94D07FF9B3666FF76FD2FB919FD5E" authority="Hoffmannsegg, 1817" authorityName="Hoffmannsegg" authorityYear="1817" box="[192,558,706,730]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="92BDEA96FF9B3666FF76FD2FBA1DFD5E" box="[192,298,706,730]" italics="true" pageId="29" pageNumber="28">Canthon</emphasis>
Hoffmannsegg, 1817
</taxonomicName>
</paragraph>
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