A checklist of Burmanniaceae in Eastern Indochina with a new record from Vietnam, Burmannia itoana
Author
Nuraliev, Maxim S.
0000-0001-8291-2633
Joint Russian-Vietnamese Tropical Scientific and Technological Center, Cau Giay, Hanoi, Vietnam. & Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia. & max. nuraliev @ gmail. com; https: // orcid. org / 0000 - 0001 - 8291 - 2633
max.nuraliev@gmail.com
Author
Yudina, Sophia V.
0000-0003-3053-8357
Joint Russian-Vietnamese Tropical Scientific and Technological Center, Cau Giay, Hanoi, Vietnam. & Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia. & Institute for Information Transmission Problems, 127051, Moscow, Russia. & yudina. sophia @ gmail. com; https: // orcid. org / 0000 - 0003 - 3053 - 8357
yudina.sophia@gmail.com
Author
Truong, Ba Vuong
0000-0003-3452-8455
Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan St., Distr. 3, Ho Chi Minh City, Vietnam. Faculty of Biology, Shenzhen MSU-BIT University, 518172, Shenzhen, China. & bavuong 2019 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 3452 - 8455
bavuong2019@yahoo.com
Author
Dang, Van Son
0000-0001-8681-4141
Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan St., Distr. 3, Ho Chi Minh City, Vietnam. Faculty of Biology, Shenzhen MSU-BIT University, 518172, Shenzhen, China. & dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141
dvsonitb@gmail.com
Author
Kopylov-Guskov, Yury O.
0000-0002-3051-8789
Joint Russian-Vietnamese Tropical Scientific and Technological Center, Cau Giay, Hanoi, Vietnam. & Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia. & yurez-kg @ yandex. ru; https: // orcid. org / 0000 - 0002 - 3051 - 8789
yurez-kg@yandex.ru
Author
Lyskov, Dmitry F.
0000-0003-0818-1662
Joint Russian-Vietnamese Tropical Scientific and Technological Center, Cau Giay, Hanoi, Vietnam. & Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia. & df. lyskov @ yandex. ru; https: // orcid. org / 0000 - 0003 - 0818 - 1662
df.lyskov@yandex.ru
Author
Kuznetsov, Andrey N.
0000-0001-5595-1039
Joint Russian-Vietnamese Tropical Scientific and Technological Center, Cau Giay, Hanoi, Vietnam. & A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninsky 33, Moscow, Russia. & forestkuz @ mail. ru; https: // orcid. org / 0000 - 0001 - 5595 - 1039
forestkuz@mail.ru
Author
Kuznetsova, Svetlana P.
0000-0002-7610-5058
Joint Russian-Vietnamese Tropical Scientific and Technological Center, Cau Giay, Hanoi, Vietnam. & tropcenterhanoi @ mail. ru; https: // orcid. org / 0000 - 0002 - 7610 - 5058
tropcenterhanoi@mail.ru
Author
Zhang, Dianxiang
0000-0001-6549-8872
Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China. & dx-zhang @ scbg. ac. cn; https: // orcid. org / 0000 - 0001 - 6549 - 8872
dx-zhang@scbg.ac.cn
text
Phytotaxa
2022
2022-04-27
544
1
61
70
journal article
55797
10.11646/phytotaxa.544.1.5
f31786c6-80cb-48a4-aea8-e4e15772e82e
1179-3163
6501413
Key to the species of
Burmannia
in
Cambodia
,
Laos
and
Vietnam
Partly based on:
Gagnepain (1908)
,
Zhang (1999)
,
Wu
et al.
(2010)
and species protologues.
Terminology of flower structure follows
Yudina
et al.
(2022)
.
1. Plant chlorophyllous (with green leaves and stem)............................................................................................................................2
- Plant apparently achlorophyllous (lacking green coloration in all plant parts)..................................................................................8
2. Plant robust, perennial, with at least some leaves longer than
3 cm
and wider than
0.3 cm
..............................................
B. disticha
- Plant delicate, annual, with leaves up to
3 cm
long and
0.3 cm
wide ................................................................................................3
3. Both outer and inner tepal lobes with double margins ......................................................................................................
B. coelestis
- Outer tepal lobes with single or double margins; inner tepal lobes with single margins ...................................................................4
4. Perianth wings narrow, ca.
0.5–0.8 mm
wide; outer tepal lobes with single margins..........................................
B. cochinchinensis
- Perianth wings pronounced, at least
1 mm
wide (usually
2–3 mm
wide); outer tepal lobes with single or double margins.............5
5. Outer tepal lobes with single or double margins (single margins only in
China
and
India
); inner tepal lobes ca.
0.4–1.5 mm
long (in Eastern Indochina, ca.
1–1.5 mm
long) .....................................................................................................................
B. chinensis
- Outer tepal lobes with single margins; inner tepal lobes ca.
0.5–0.8 mm
long..................................................................................6
6. Flower
10–13 mm
long; outer tepal lobes ca. (1)
1.8–2.2 mm
long ............................................................................
B. subcoelestis
- Flower
6–11 mm
long; outer tepal lobes ca.
1–1.5 mm
long .............................................................................................................7
7. Outer tepal lobes fleshy; inner tepal lobes oblanceolate or spatulate...............................................................................
B. luteoalba
- Outer tepal lobes thin; inner tepal lobes lanceolate..............................................................................................................
B. pusilla
8(1). Whole plant bluish..............................................................................................................................................................................9
- Whole plant whitish..........................................................................................................................................................................12
9. Perianth wings prominent,
1–2 mm
wide..............................................................................................................................
B. itoana
- Perianth wings narrow (often rib-like), up to
0.8 mm
wide .............................................................................................................10
10. Perianth wings rib-like to semi-elliptic,
0.2–0.8 mm
wide; outer tepal lobes ca.
1 mm
long
...........................................
B. wallichii
- Perianth wings rib-like; outer tepal lobes
2.5–4 mm
long................................................................................................................11
11. Flower larger (outer tepal lobes
3–4 mm
×
1.5–2 mm
, inner tepal lobes
2 mm
×
1.5 mm
, ovary
2.5–3 mm
long); outer tepal lobes at apex slightly attenuate and obtuse.................................................................................................................................
B. coerulea
- Flower smaller (outer tepal lobes
2.5–3 mm
×
1.25–1.5 mm
, inner tepal lobes
1 mm
×
0.8 mm
, ovary
2–2.5 mm
long); outer tepal lobes at apex attenuate into long acute caudate tip ca.
1 mm
long..............................................................................
B. unguiculata
12(8). Inflorescence dense (usually head-like), with 3–13 flowers; flower wingless or with rib-like perianth wings...........
B. championii
- Inflorescence lax (axes usually apparently elongated), with up to 5 (rarely 9) flowers; perianth wings prominent or rarely rib-like ..........................................................................................................................................................................................................13
13. Outer tepal lobes broadly oblong to broadly quadrangular, with emarginate apex; inner tepal lobes absent ....................
B. oblonga
- Outer tepal lobes ovate, triangular or oblong, with acuminate, obtuse, rounded or slightly retuse apex; inner tepal lobes absent or present...............................................................................................................................................................................................14
14. Floral tube
1.5–2.5 mm
long; outer tepal lobes
0.5–1.2 mm
long; inner tepal lobes present ........................................
B. nepalensis
- Floral tube
3.5–5 mm
long; outer tepal lobes
1–1.8 mm
long; inner tepal lobes absent or present.................................
B. lutescens