A taxonomic revision of seed harvester ants of the Tetramorium solidum group (Hymenoptera: Formicidae) in southern Africa
Author
Mbanyana, Nokuthula
157A51ED-7C85-4CA3-84E4-B7BD1E2A7E8F
Iziko Museums of South Africa, P. O. Box 61, Cape Town, 8000 South Africa. Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna-son, Okinawa, Japan. Department of Botany and Zoology, Stellenbosch University, Private Bag X 1, Matieland, 7602, South Africa.
nmbanyana@iziko.org.za
Author
Garcia, Francisco Hita
B7ADF56F-935D-4BD8-ADB3-50E96F8BB463
fhitagarcia@gmail.com
Author
Robertson, Hamish Gibson
972FAB6C-C5FA-45EA-9A17-9F156845BE4F
Iziko Museums of South Africa, P. O. Box 61, Cape Town, 8000 South Africa. Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna-son, Okinawa, Japan. Department of Botany and Zoology, Stellenbosch University, Private Bag X 1, Matieland, 7602, South Africa.
hrobertson@iziko.org.za
Author
Roux, Johannes Jacobus Le
2995D443-01FA-4F65-BFEE-ECA07B73504A
jleroux@sun.ac.za
text
European Journal of Taxonomy
2018
2018-08-07
454
1
59
journal article
22298
10.5852/ejt.2018.454
654c9653-fbef-4d0d-9e07-619084ac1162
3817358
07F6B28D-ECA4-4F14-8132-0A58DE156057
Genus
Tetramorium
Mayr, 1855
Diagnosis of
T. solidum
species-group workers
The following diagnosis, which is based on
Bolton
(1980)
, distinguishes the group from all other groups in the Afrotropical region: relatively large
Tetramorium
(HW
0.875
–1.279
mm
, WL
0.915
–1.498
mm
) with very well-developed, massively constructed heads (HW>
0.80 mm
and in most cases exceeding 1.00 mm) equipped with strong mandibles; 12-segmented antennae; anterior clypeal margin with deep and wide median indentation, sometimes occupying half of the anterior clypeal margin, except for
T. barbigerum
Bolton
, 1980
, in which it is more shallowly impressed; large eyes (OI 24–26); frontal carinae absent or very short, if present ending before anterior eye level; frontal scrobes absent; ventral face of head (usually) with J-shaped ammocheate hairs; base of first gastral tergite always sculptured, even if weakly so.
Overview of species of
Tetramorium solidum
group
The species of the
Tetramorium solidum
group can be distinguished from other groups of
Tetramorium
by the presence of a median notch or impression on the anterior clypeal margin, which is often broad and deeply indented, combined with the presence of ammocheate hairs on the ventral side of the head (
Bolton
1980
).
Bolton
(1980
,
1985
) grouped species of the
T
.
solidum
group into three complexes based on the form of their pilosity. The first complex comprises
T
.
dichroum
Santschi, 1932
,
T
.
peringueyi
Arnold, 1926
and
T
.
rothschildi
(
Forel, 1907
)
. These three species are characterized by the presence of abundant hairs on the dorsal surfaces of the head and body; these hairs are branched in
T. rothschildi
and simple in the other two species. The second complex includes
T
.
clunum
Forel, 1913
,
T
.
galoasanum
Santschi, 1910
and
T
.
setuliferum
Emery, 1895
; they all lack erect hairs on the dorsal surface of the antennae, mesosoma, petiole, postpetiole and first gastral tergite. They have a bizarre pilosity, which consists of short, broad, blunted, flattened and strongly appressed glittering silvery hairs. The third complex consists of
T
.
barbigerum
,
T
.
glabratum
Stitz, 1923
,
T
.
grandinode
Santschi, 1913
,
T
.
jordani
Santschi, 1937
,
T
.
pogonion
Bolton
, 1980
,
T
.
rufescens
Stitz, 1923
,
T
.
signatum
Emery, 1895
and
T
.
solidum
Emery, 1886
, in which the body is hairless, or nearly so. Only
T
.
solidum
has a few hairs on the mesosoma (
Bolton
1980
).
Synopsis of species of the Afrotropical region
Tetramorium aisha
Mbanyana, Robertson & Hita Garcia
sp. nov.
Tetramorium barbigerum
Bolton
, 1980
Tetramorium brigitteae
Mbanyana, Robertson & Hita Garcia
sp. nov.
Tetramorium clunum
Forel, 1913
Tetramorium dichroum
Santschi, 1932
Tetramorium duncani
Mbanyana, Robertson & Hita Garcia
sp. nov.
Tetramorium galoasanum
Santschi, 1910
Tetramorium glabratum
Stitz, 1923
=
Tetramorium solidum
st.
glabratum
Stitz, 1923
=
Tetramorium solidum
race
glabratum
var.
aciculatum
Stitz, 1923
[name unavailable]
=
Tetramorium rutilum
Prins, 1973
Tetramorium grandinode
Santschi, 1913
=
Tetramorium grandinode
var.
hopensis
Forel, 1914
Tetramorium jordani
Santschi, 1937
=
Tetramorium aspinatum
Prins, 1973
Tetramorium lerouxi
Mbanyana, Robertson & Hita Garcia
sp. nov.
Tetramorium margueriteae
Mbanyana, Robertson & Hita Garcia
sp. nov.
Tetramorium peringueyi
Arnold, 1926
Tetramorium pogonion
Bolton
, 1980
Tetramorium rothschildi
(
Forel, 1907
)
Tetramorium rufescens
Stitz, 1923
=
Tetramorium solidum
st.
rufescens
Stitz, 1923
Tetramorium setuliferum
Emery, 1895
=
Tetramorium squamiferum
Forel, 1894
[name unavailable]
=
Tetramorium setuliferum
var.
cucalense
Santschi, 1910
=
Tetramorium setuliferum
var.
triptolemus
Arnold, 1917
Tetramorium signatum
Emery, 1895
=
Tetramorium solidum
var.
signatum
Emery, 1895
=
Tetramorium solidum
subsp.
lugubre
Forel, 1910
=
Tetramorium solidum
var.
grootensis
Forel, 1913
=
Tetramorium solidum
var.
tuckeri
Arnold, 1926
Tetramorium solidum
Emery, 1886