<document id="91DC85213EC2C524902F57462E003E56" ID-CLB-Dataset="182444" ID-DOI="10.11646/zootaxa.5202.1.1" ID-GBIF-Dataset="45ba25c1-47a9-4001-93d9-294815bf37e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7285600" ID-ZooBank="BF4831D2-D98B-4265-9138-03DB8607B826" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1667554680289" checkinUser="plazi" docAuthor="Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta &amp; Riva, Ignacio De La" docDate="2022" docId="03C287E8FFFBFFCAFF4BFF0AFB2BE065" docLanguage="en" docName="zootaxa.5202.1.1.pdf" docOrigin="Zootaxa 5202 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5202.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Grayia caesar" docType="treatment" docVersion="10" lastPageNumber="90" masterDocId="FFFBFF90FFA2FF93FFDCFF9DFFB5E232" masterDocTitle="Synopsis of the terrestrial Reptiles of Equatorial Guinea" masterLastPageNumber="197" masterPageNumber="1" pageNumber="90" updateTime="1732576365292" updateUser="ExternalLinkService">
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<mods:title id="33DC28F9EAE9DA1C9CADDAE7BBA10AF5">Synopsis of the terrestrial Reptiles of Equatorial Guinea</mods:title>
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<mods:namePart id="7698E8B03D2ABC4F57AA7055E2AB8D92">Sánchez-Vialas, Alberto</mods:namePart>
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<mods:namePart id="8B7BC308CBAD3E68782BDCE772ADD88A">Calvo-Revuelta, Marta</mods:namePart>
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<mods:namePart id="8A0A515471B996B9C4737BBF6AF9A16F">Riva, Ignacio De La</mods:namePart>
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<mods:title id="E553BE7483E67E33E2BD40324FFA8EBE">Zootaxa</mods:title>
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<mods:date id="FCCE2FDA5524A020A61A868B57AD7734">2022</mods:date>
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<treatment id="03C287E8FFFBFFCAFF4BFF0AFB2BE065" ID-DOI="http://doi.org/10.5281/zenodo.7293479" ID-GBIF-Taxon="204305273" ID-Zenodo-Dep="7293479" LSID="urn:lsid:plazi:treatment:03C287E8FFFBFFCAFF4BFF0AFB2BE065" httpUri="http://treatment.plazi.org/id/03C287E8FFFBFFCAFF4BFF0AFB2BE065" lastPageNumber="90" pageId="89" pageNumber="90" scope_family="Colubridae" scope_order="Squamata">
<subSubSection id="C3716575FFFBFFCAFF4BFF0AFDB8E283" box="[151,525,151,177]" pageId="89" pageNumber="90" type="nomenclature">
<paragraph id="8BD436FEFFFBFFCAFF4BFF0AFDB8E283" blockId="89.[151,525,151,213]" box="[151,525,151,177]" pageId="89" pageNumber="90">
<heading id="D09C8192FFFBFFCAFF4BFF0AFDB8E283" bold="true" box="[151,525,151,177]" fontSize="11" level="2" pageId="89" pageNumber="90" reason="4">
<taxonomicName id="4C6B4D7DFFFBFFCAFF4BFF0AFDB8E283" ID-CoL="3HBV8" authority="(Gunther, 1863)" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[151,525,151,177]" class="Reptilia" family="Colubridae" genus="Grayia" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="caesar">
<emphasis id="B91FEAECFFFBFFCAFF4BFF0AFDB8E283" bold="true" box="[151,525,151,177]" pageId="89" pageNumber="90">
<emphasis id="B91FEAECFFFBFFCAFF4BFF0AFEF5E283" bold="true" box="[151,320,151,177]" italics="true" pageId="89" pageNumber="90">Grayia caesar</emphasis>
(Günther, 1863)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3716575FFFBFFCAFF4BFF26FEA3E2E7" box="[151,278,187,213]" pageId="89" pageNumber="90" type="description">
<paragraph id="8BD436FEFFFBFFCAFF4BFF26FEA3E2E7" blockId="89.[151,525,151,213]" box="[151,278,187,213]" pageId="89" pageNumber="90">
<figureCitation id="13502A7BFFFBFFCAFF4BFF26FEA3E2E7" box="[151,278,187,213]" captionStart="FIGURE 22" captionStartId="90.[151,250,1901,1925]" captionTargetBox="[166,1420,221,1855]" captionTargetId="figure-19@90.[158,1430,181,1877]" captionTargetPageId="90" captionText="FIGURE 22. From left to right: lateral, dorsal and ventral views of the head of: Toxicodryas blandingii (A) (MNCN 23819), Grayia caesar (B) (EBD 20945), and Grayia ornata (C) (EBD 24950). Photos: AS-V." figureDoi="http://doi.org/10.5281/zenodo.7285691" httpUri="https://zenodo.org/record/7285691/files/figure.png" pageId="89" pageNumber="90">Figure 22B</figureCitation>
</paragraph>
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<subSubSection id="C3716575FFFBFFCAFF4BFE9DFD09E304" pageId="89" pageNumber="90" type="reference_group">
<paragraph id="8BD436FEFFFBFFCAFF4BFE9DFBD8E32A" blockId="89.[151,1133,255,310]" box="[151,1133,255,280]" pageId="89" pageNumber="90">
<taxonomicName id="4C6B4D7DFFFBFFCAFF4BFE9DFD96E325" authority="Gunther, 1863: 357" authorityName="Gunther" authorityPageNumber="357" authorityYear="1863" box="[151,547,255,280]" class="Reptilia" family="Colubridae" genus="Xenurophis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="caesar">
<emphasis id="B91FEAECFFFBFFCAFF4BFE9DFEE0E325" box="[151,341,256,280]" italics="true" pageId="89" pageNumber="90">Xenurophis caesar</emphasis>
Günther, 1863: 357
</taxonomicName>
. 
<typeStatus id="54D0885CFFFBFFCAFDF0FE9DFDEBE32A" box="[556,606,256,280]" pageId="89" pageNumber="90">Type</typeStatus>
locality: “Fernando Po”, 
<location id="8EB46025FFFBFFCAFCBEFE9DFC17E32A" LSID="urn:lsid:plazi:treatment:03C287E8FFFBFFCAFF4BFF0AFB2BE065:8EB46025FFFBFFCAFCBEFE9DFC17E32A" box="[866,930,256,280]" country="Equatorial Guinea" name="Bioko" pageId="89" pageNumber="90">Bioko</location>
, 
<collectingCountry id="F37C766EFFFBFFCAFC71FE9DFBDCE32A" box="[941,1129,256,280]" name="Equatorial Guinea" pageId="89" pageNumber="90">Equatorial Guinea</collectingCountry>
.
</paragraph>
<paragraph id="8BD436FEFFFBFFCAFF4BFE82FD09E304" blockId="89.[151,1133,255,310]" box="[151,700,286,310]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="AB7B11D0FFFBFFCAFF4BFE82FD09E304" box="[151,700,286,310]" pageId="89" pageNumber="90">
<taxonomicName id="4C6B4D7DFFFBFFCAFF4BFE82FE60E304" ID-CoL="3HBV8" authority="(Gunther, 1863)" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[151,469,286,310]" class="Reptilia" family="Colubridae" genus="Grayia" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="caesar">
<emphasis id="B91FEAECFFFBFFCAFF4BFE82FE9CE304" box="[151,297,287,310]" italics="true" pageId="89" pageNumber="90">Grayia caesar</emphasis>
(Günther, 1863)
</taxonomicName>
: 
<treatmentCitation id="0ACA10EFFFFBFFCAFE03FE83FD0DE304" author="Boulenger, G. A." box="[479,696,286,310]" page="944" pageId="89" pageNumber="90" year="1909">
<bibRefCitation id="EFFA4B0FFFFBFFCAFE03FE83FD0DE304" author="Boulenger, G. A." box="[479,696,286,310]" pageId="89" pageNumber="90" pagination="944 - 952" refId="ref72366" refString="Boulenger, G. A. (1909) On the ophidian genus Grayia. Proceedings of the Zoological Society of London, 1909, 944 - 952. https: // doi. org / 10.1111 / j. 1469 - 7998.1910. tb 06985. x" type="journal article" year="1909">Boulenger 1909: 944</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFFBFFCAFF4BFEF9FB84E3F5" pageId="89" pageNumber="90" type="distribution">
<paragraph id="8BD436FEFFFBFFCAFF4BFEF9FB84E3F5" blockId="89.[151,1436,356,599]" pageId="89" pageNumber="90">
<emphasis id="B91FEAECFFFBFFCAFF4BFEF9FE9CE34C" bold="true" box="[151,297,356,382]" pageId="89" pageNumber="90">Distribution</emphasis>
. Central Africa, from Bioko and 
<collectingCountry id="F37C766EFFFBFFCAFD48FEF8FCBCE34D" box="[660,777,357,383]" name="Cameroon" pageId="89" pageNumber="90">Cameroon</collectingCountry>
to 
<collectingCountry id="F37C766EFFFBFFCAFCF5FEF9FB1BE34D" box="[809,1198,356,383]" name="Democratic Republic of the Congo" pageId="89" pageNumber="90">Democratic Republic of the Congo</collectingCountry>
; it extends southward to 
<collectingCountry id="F37C766EFFFBFFCAFF69FE15FEA7E390" box="[181,274,392,418]" name="Angola" pageId="89" pageNumber="90">Cabinda</collectingCountry>
(northern 
<collectingCountry id="F37C766EFFFBFFCAFE55FE15FE68E390" box="[393,477,392,418]" name="Angola" pageId="89" pageNumber="90">Angola</collectingCountry>
) (
<bibRefCitation id="EFFA4B0FFFFBFFCAFE28FE15FC90E391" author="Chippaux, J. P. &amp; Jackson, K." box="[500,805,392,419]" pageId="89" pageNumber="90" refId="ref74358" refString="Chippaux, J. P. &amp; Jackson, K. (2019) Snakes of Central and Western Africa. Johns Hopkins University Press, Baltimore, Maryland, 429 pp." type="book" year="2019">Chippaux &amp; Jackson 2019</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFFBFFCAFCEEFE14FBAFE391" author="Marques, M. P. &amp; Ceriiaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A." box="[818,1050,392,419]" pageId="89" pageNumber="90" pagination="1 - 501" refId="ref83759" refString="Marques, M. P., Ceriiaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola - Atlas of Historical and Bibliographic Records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques 
<emphasis id="B91FEAECFFFBFFCAFC40FE14FC60E390" box="[924,981,392,418]" italics="true" pageId="89" pageNumber="90">et al.</emphasis>
2018
</bibRefCitation>
). In 
<collectingCountry id="F37C766EFFFBFFCAFB91FE15FAAFE390" box="[1101,1306,392,418]" name="Equatorial Guinea" pageId="89" pageNumber="90">Equatorial Guinea</collectingCountry>
it has been recorded from Bioko (
<bibRefCitation id="EFFA4B0FFFFBFFCAFE4CFE31FD88E3F4" author="Gunther" box="[400,573,428,454]" pageId="89" pageNumber="90" pagination="348 - 365" refId="ref78893" refString="Gunther. A. (1863 a) Third account of new species of snakes in the collection of the British Museum. Annals &amp; Magazine of Natural History, Series 3, 12 (71), 348 - 365. https: // doi. org / 10.1080 / 00222936308681536" type="journal article" year="1863">Günther 1863a</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFFBFFCAFD94FE31FD4AE3F5" author="Boulenger, G. A." box="[584,767,428,455]" pageId="89" pageNumber="90" refId="ref71762" refString="Boulenger, G. A. (1894) Catalogue of the Snakes in the British Museum (Natural History). Vol. 2. Trustees, London, xi + 382 pp., 20 pls." type="book" year="1894">Boulenger 1894</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFFBFFCAFCD7FE30FC2AE3F5" author="Bocage, J. V. B." box="[779,927,429,455]" pageId="89" pageNumber="90" pagination="25 - 59" refId="ref71106" refString="Bocage, J. V. B. (1903 a) Contribution a la faune des quatre iles du Golfe de Guinee. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 7, 25 - 59." type="journal article" year="1903">Bocage 1903</bibRefCitation>
) (
<figureCitation id="13502A7BFFFBFFCAFC6AFE30FB90E3F5" box="[950,1061,429,455]" captionStart="MAPS 24" captionStartId="87.[151,223,1943,1967]" captionTargetBox="[203,1383,194,1906]" captionTargetId="figure-18@87.[161,1427,181,1919]" captionTargetPageId="87" captionText="MAPS 24 A–C. Distribution maps for Equatorial Guinean records of (A) Toxicodryas adamanteus (question mark represents an imprecise record); (B) Toxicodryas blandingii; (C) Grayia caesar (question mark represents an imprecise record)." figureDoi="http://doi.org/10.5281/zenodo.7304733" httpUri="https://zenodo.org/record/7304733/files/figure.png" pageId="89" pageNumber="90">Map 24C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFFBFFCAFF1BFE4DFB2BE065" pageId="89" pageNumber="90" type="discussion">
<paragraph id="8BD436FEFFFBFFCAFF1BFE4DFE86E03C" blockId="89.[151,1436,356,599]" pageId="89" pageNumber="90">
<emphasis id="B91FEAECFFFBFFCAFF1BFE4DFEFAE3D8" bold="true" box="[199,335,464,490]" pageId="89" pageNumber="90">Comments.</emphasis>
There are no precise records of this species for Bioko. We provide the first records of this species for 
<collectingCountry id="F37C766EFFFBFFCAFF63FE68FE9BE03C" box="[191,302,500,527]" name="Equatorial Guinea" pageId="89" pageNumber="90">Río Muni</collectingCountry>
.
</paragraph>
<paragraph id="8BD436FEFFFBFFCAFF1BFD85FB2BE065" blockId="89.[151,1436,356,599]" pageId="89" pageNumber="90">
<emphasis id="B91FEAECFFFBFFCAFF1BFD85FE7DE000" bold="true" box="[199,456,536,562]" pageId="89" pageNumber="90">Specimens examined.</emphasis>
<materialsCitation id="3B033CA3FFFBFFCAFE0CFD84FB2FE065" ID-GBIF-Occurrence="3962324572" accessionNumber="EBD 25028, EBD 20945" collectingDate="1985-10-14" collectingDateMax="1987-02" collectingDateMin="1985-10-14" collectionCode="EBD" country="Equatorial Guinea" location="Two" municipality="San Joaquin de Ndyiacom" pageId="89" pageNumber="90" specimenCount="1">
<location id="8EB46025FFFBFFCAFE0CFD84FDB4E001" LSID="urn:lsid:plazi:treatment:03C287E8FFFBFFCAFF4BFF0AFB2BE065:8EB46025FFFBFFCAFE0CFD84FDB4E001" box="[464,513,537,563]" country="Equatorial Guinea" municipality="San Joaquin de Ndyiacom" name="Two" pageId="89" pageNumber="90">Two</location>
specimens. 
<collectingCountry id="F37C766EFFFBFFCAFD52FD84FCB5E000" box="[654,768,536,563]" name="Equatorial Guinea" pageId="89" pageNumber="90">Río Muni</collectingCountry>
: 
<collectingMunicipality id="6BB0AC84FFFBFFCAFCD0FD84FB83E000" box="[780,1078,536,563]" pageId="89" pageNumber="90">San Joaquín de Ndyiacom</collectingMunicipality>
(road from Bata to Ayamiken), 
<date id="FFD5103EFFFBFFCAFF4BFDA1FE8EE065" box="[151,315,572,599]" pageId="89" pageNumber="90" value="1987-02">
<collectingDate id="EF91E9D6FFFBFFCAFF4BFDA1FE8EE065" box="[151,315,572,599]" pageId="89" pageNumber="90" value="1987-02">February 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFFBFFCAFE96FDA0FE31E065" box="[330,388,573,599]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="89" pageNumber="90">EBD</collectionCode>
25028); Miboman (
<quantity id="4C939B1BFFFBFFCAFDB9FDA0FD18E064" box="[613,685,572,599]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="89" pageNumber="90" unit="km" value="30.0">30 km</quantity>
from Bata), 
<date id="FFD5103EFFFBFFCAFCE0FDA0FC4DE065" box="[828,1016,572,599]" pageId="89" pageNumber="90" value="1985-10-14">
<collectingDate id="EF91E9D6FFFBFFCAFCE0FDA0FC4DE065" box="[828,1016,572,599]" pageId="89" pageNumber="90" value="1985-10-14">14 October 1985</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFFBFFCAFBDBFDA0FBF4E065" box="[1031,1089,573,599]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="89" pageNumber="90">EBD</collectionCode>
20945)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>