Variability of Habroblattula drepanoides gen. et. sp. nov. (Insecta: Blattaria: Blattulidae) from the Yixian Formation in Liaoning, China
Author
Wang, Tian-Tian
Author
Liang, Jun-Hui
Author
Ren, Dong
text
Zootaxa
2007
1443
17
27
journal article
10.5281/zenodo.176061
9705d176-1598-469c-8614-33dc4a7e5af1
1175-5326
176061
Genus
Habroblattula
gen
. nov.
Type
species.
Habroblattula drepanoides
sp. nov.
Diagnosis
. Wings, pronotum and legs with bright and dark colour. Forewing Sc terminally branched, CuP strongly curved, all branches strong and dark, intercalaries and cross-veins distinct. Hindwing with pterostigma (dark macula in the R1), R1 and RS differentiated, CuA with abundant secondary branches.
Description.
Forewings with more or less parallel margins and characteristic forewing coloration (with falciform dapple of the R and M area and with macula along with the basal part of the posterior margin). Costal space very short and narrow; Sc terminally branched; R slightly curved, multiple simple branches, ending before wing apex; apically descending M simplified; CuA with branches long, running parallel to margin; CuP strongly curved; Anal veins simple and with thick cross-veins (visible even in poorly preserved specimens). Hindwing with pterostigma (dark macula in the R1). Sc simple; R1 and RS differentiated; M without tertiary branchings; CuA with abundant secondary branches; CuP simple. Apex distinctly colored. Pronotum with anomalous dapple, wider than long. Legs long with bright dark macula, femur shorter than tibia; all tibiae very short, with numerous spurs. Body soft, terminal segments with multisegmented cerci and styli.
Remarks
.
Habroblattula
is assigned to the family
Blattulidae
, because the forewing has long Sc, regular venation with distinct intercalaries and hindwing has simple CuP, branched A1. (Vishniakova 1983)
Habroblattula
gen
. nov.
is closely related to
Svabula
Vršanský, 2005b
from the Berriasian Lower Cretaceous sediments of Sharin-gol in
Mongolia
. Synapomorphies include wider forewing with margins parallel, branches strong, straight and dark, clavus with cross-veins.
Svabula
differs in small forewing size, different coloured pattern, having less apparent intercalaries and thick cross-veins.
Hindwing of the new genus resembles
Kridla
Vršanský, 2005a
from the Albian or Cenomanian Upper Cretaceous sediments of the Kyndalskaya Formation in having distinct pterostigma and distinct apex coloration, but differs from the latter by secondarily branched R1, and A2, simple M and more abundant cross-veins.
The new genus differs from genus
Tarakanula
Vršanský, 2003
by prominently coloured wings, pronotum and legs of new genus, with more dense veins and with apparent thick cross-veins.
Habroblattula
gen
. nov.
is easily differentiated from
Elisama
Giebel, 1856
and
Blattula
Handlirsch, 1906
–1908 by having coloured wings, pronotum and legs, the forewing with strong, dark branches and apparent cross-veins in clavus and Cu area.
Xonpepetla
Cifuentes-Ruiz et
Vršanský, 2006
from the Campanian of
Mexico
differ in being robust body with very short and very wide forewings.
Etymology
: The name is derived from the Greek prefix
habr
(meaning “elegancy”) and the
type
genus
Blattula
(the
type
genus of this family).