Pre-testical spermathecal pores and unusual setal arrangement in the South African endemic microchaetid earthworms of presumed Gondwanan origin (Oligochaeta: Microchaetidae)
Author
Plisko, Jadwiga Danuta
text
African Invertebrates
2009
2009-12-31
50
2
237
237
http://www.bioone.org/doi/abs/10.5733/afin.050.0202
journal article
10.5733/afin.050.0202
2305-2562
7910427
Proandricus adami
Plisko, 2003
Fig. 6
Proandricus adami
: Plisko 2003: 295
.
Type
locality:
South Africa
:
KZN
,
Royal Natal National Park
,
Bank of Devil’s Hoek River
, approx.
28
°
42'S
:
28
°
54'E
.
Known
from numerous sites located in the
central
Drakensberg
mountain range
.
Material examined:
Holotype
NMSA
/Olig.02530; 3 cl
paratypes
NMSA
/Olig.00918 and
NMSA
/Olig.00919.
New
material:
South Africa
,
KZN
,
Royal Natal National Park
, forest, 13 cl and 5 juv collected during
March–April 2006
from 6 slightly distanced sites by
MDTP
KwaZulu-Natal
University
students group:
NMSA
/Olig.04203,
NMSA
/Olig.04204,
NMSA
/Olig.04215,
NMSA
/Olig.04246,
NMSA
/Olig.04249,
NMSA
/Olig.04252a,
NMSA
/Olig.04254,
NMSA
/Olig.04304,
NMSA
/Olig.4307,
NMSA
/Olig.04308.
The external morphological characters in all examined material agree with the original description. Although, preclitellarly and postclitellarly the setae are closely paired in regular rows, and in the clitellar area some of the pairs of
ab
are shifted up between
bc
, forming an irregular arrangement, some additional abnormality in setal arrangement was noted. On segment 16 and
18 in
inter-setal space between
ab
and
cd
, a pair of additional setae occurred, making five pairs of setae per segment; in the other
two specimens
, two additional pairs of setae were placed above
ab
, giving six pair per segment on three clitellar segments.This abnormal phenomenon has been noted in three clitellate specimens and on
two juvenile
, selected from the population of 18 individuals collected in the Royal Natal National Park forested area. Spermathecae variable in shape and size. Anterior pairs elongated, always evidently smaller than posterior, often containing sperm. Posterior oval, large, with thin, elongated duct. In the majority of specimens examined, the posterior spermathecae are iridescent, filled with sperm and transparent through the body wall. Spermathecal pores in intersegmental furrows 9/10 and 10/11, although the pores of 9/10 are externally difficult to trace. Clitellum slightly resembling this of
amphius
, although on various individuals it was variably developed, unequally encircling segments ventrally.
P. adami
,
although noticeably similar to the species of the
lesothoensis
species-group, manifests irregularity in the setal arrangement extending to abnormality. Similar to
bourquini
,
and
lesothoensis
, but differs by peculiarly arranged setae.