A taxonomic revision of Cephalaria subgen. Cephalaria sect. Atrocephalae (Caprifoliaceae) from Iran
Author
Ranjbar, Massoud
Author
Ranjbar, Zahra
text
Phytotaxa
2017
2017-06-27
311
2
101
138
http://dx.doi.org/10.11646/phytotaxa.311.2.1
journal article
10.11646/phytotaxa.311.2.1
1179-3163
13700563
7.
Cephalaria juncea
Boissier (1846: 72)
.
Type:—
IRAN
. Prov. Kohgiluyeh and Boyer-Ahmad:
Dena
Mt.
,
Th. Kotschy 627
(G-Boiss! [G00330272!],
lectotype
designated here;
isolectotypes
G!, E!, FI!, K!, P!, MO!).
Fig. 7
.
Perennial, erect, slender, seemly leafless, looking similar to
Juncus
spp.
Stem up to
40 cm
high, branched, scabrous, green, with opposite branches. Leaves slightly pubescent, linear to filiform, minute, angustate, less than
1 cm
wide, pinnatipartite with decurrent segments, opposite, blades rather symmetrical, leaves green when dried. Lower cauline leaves withered at anthesis. Middle cauline leaves scarcely developed, filiform, pinnatipartite with 8–10 decurrent segments, linear in outline, 4.5–5 ×
0.5–0.8 cm
; lateral segments oblong-elliptic, 3–5 ×
1.5–3 mm
, obtuse at apex, entire, midvein impressed to flat adaxially, rounded abaxially, secondary veins weakly percurrent, oblique, tertiary veins unclear. Peduncle
10–20 cm
long, glabrous to sparsely pilose. Capitula subglobose, 1–1.5 ×
0.7–1 cm
in diameter in flower, subradiant, few-flowered, solitary, 1.2–2 ×
1–1.5 cm
in diameter in fruit. Involucral bracts in one series, ovate-rotund, ca. 5 ×
2.5–3 mm
, pinkish to brown-coloured dorsally, villous, obtuse to slightly acuminate at apex. Receptacular bracts in 4–5 series, sublanceolate-acuminate 5–8 ×
2–2.5 mm
, pinkish with brown strips, abruptly acute or acuminate and recurved at apex, pilose or sericeous, ciliate at margins. Calyx cupuliform, 1–1.5 ×
1.5–2 mm
, green to creamy-coloured, pubescent, with irregular teeth
1–2 mm
long. Corolla ca.
6 mm
long, yellow to lemon-coloured, densely covered with appressed hairs outside, stamens slightly longer than corolla, anthers deeply yellow. Involucel 3–4 ×
1.5 mm
in flower, 4-angled, adpressely hirsute (hairs up to ca.
1 mm
long), 4 long and 4 short teeth at apex.
FIGURE 7.
Lectotype of
Cephalaria juncea
Boiss.
(
Kotschy 627
, G-Boiss! [G00330272]).
Additional specimens examined
:—
IRAN
. Prov.
Fars
:
79 km
NW of Ardakan
,
2300 m
,
Pabo
2420
(
TARI
)
;
Shiraz
,
Kaleh Kalagh Mt.
,
Behbudi
912-E
(
TARI
)
.
Distribution and habitat:
—
Cephalaria juncea
is endemic to western to central
Iran
and is known only from mountainous areas in Chaharmahal va Bakhtiari,
Esfahan
,
Fars
and Kohgiluyeh Boyer Ahmad provinces. The species grows on rocky mountain slopes at elevations of
1500–3100 m
.
Taxonomic notes:
—
Cephalaria juncea
is the only species of
C.
sect.
Atrocephalae
in
Iran
that looks leafless and similar to
Juncus
, and may be an isolated species.
Nomenclatural notes:
—The original material of
Cephalaria juncea
was collected by Kotschy on
13 July 1842
and found at G (G00330272, G00330272-a, G00360038, G00406424, G006437), K (K000762996), FI (FI004499), E (E00690430), MO (MO5259440, MO5259441) and P (P02284056, P02284057, PO2284058).
Boissier (1846)
cited these specimens in the protologue, but he did not designate a type. Szabó (1914) examined
two specimens
of G and E without designating a type. Lack examined some of these specimens at G-Boiss and G, but did not designate any
lectotype
. Gautier examined G-Boiss and G specimens and considered two sheets of G-Boiss (G00330272, G00330272- a) as the
holotype
specimen (on red lable), but apparently he did not publish his choice. Here it is confirmed that the GBoiss specimens have been certainly examined by Boissier and one sheet (G00330272) is considered as the
lectotype
of
C. juncea
(this sheet is perfect and contains morphologically recognisable elements).