Seven new species of cestode parasites (Neodermata, Platyhelminthes) from Australian birds
Author
Mariaux, Jean
B97E611D-EC33-4858-A81C-3E656D0DA1E2
Natural History Museum of Geneva, CP 6434, 1211 Geneva 6, Switzerland. Department of Genetics and Evolution, University of Geneva, Boulevard d’Ivoy 4, 1205 Geneva, Switzerland. Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113 Sofia, Bulgaria.
Author
Georgiev, Boyko B.
88352C92-555A-444D-93F4-97F9A3AC2AEF
bbg@ecolab.bas.bg
text
European Journal of Taxonomy
2018
2018-05-31
440
1
42
journal article
22133
10.5852/ejt.2018.440
1a75a03c-3b98-4f41-821d-10da4406d93e
3827554
DB80A42B-5C53-455B-86A4-2003D6F03522
Notopentorchis musealis
sp. nov.
urn:lsid:zoobank.org:act:
99114137-DA7B-47EE-996F-49005C2B8A07
Figs 29–33
,
Table 5
Etymology
The species name refers to the fact that this species is described from museum material.
Material examined
Holotype
AUSTRALIA
:
New South Wales
,
Sydney
,
38°8′ S
,
144°22′ E
, 2005,
Karie Rose
leg. (AHC 36478).
Paratype
AUSTRALIA
: same data as for
holotype
(AHC 36478).
Intensity
Two specimens and a few small fragments. Co-infection with
Neoliga
sp.
Description
Body of medium size, with maximum length
34 mm
for 404 proglottides; maximum width 590, at level of gravid proglottides. Proglottides craspedote, all but the few most developed (pre-gravid) ones wider than long. Ventral osmoregulatory canals connected posteriorly in each proglottis with transverse anastomosis. Scolex slightly rounded, not distinctly separated from neck, with diameter 248–345 (n = 2). Suckers elongated
170–215 in
diameter (182, n = 7).
Rostellum
very muscular, sucker-like, slightly concave anteriorly and conical posteriorly,
114–139 in
diameter and 107–125 long (n = 2) (
Fig. 29
). Rostellar hooks mostly lost (probably several tens, only 7 remaining), in two rows, subequal,
21–23 in
length (22, n = 5). Posterior hooks with slightly more curved handle than anterior hooks (
Fig. 30
); guard bifid. Small epiphyses on both guards and handles. Neck long; proglottization distinct at 475 from posterior margin of suckers. Genital pores at mid-length of lateral proglottis margin, irregularly alternating in short series, maximum 5 consecutive pores observed on one side. Ventral osmoregulatory canals connected posteriorly in each proglottis by single transverse anastomosis. Genital ducts passing ventrally to osmoregulatory canals. Genital atrium cup-shaped, voluminous.
Testes large, spherical, 5–8 (6.5, n = 11) in number, in single posterior and dorsal layer, often overlapping vitellarium and posterior parts of ovary, occasionally overlapping longitudinal osmoregulatory canals. External vas deferens convoluted antero-porally, poral to developing uterus surrounded by intensely-stained cells and forming together with them dense mass in anteroporal part of median field. Cirrus-sac 91–112 × 54–64 (105 × 59, n = 15), consisting of narrower, almost cylindrical, distal part and spherical thick-walled proximal part, often overlapping poral osmoregulatory canals; internal vas deferens coiled in proximal part of cirrus-sac. Cirrus cylindrical, unarmed,
10–12 in
diameter.
Figs 29–33.
Notopentorchis musealis
sp. nov.
29
. Scolex.
30
. Hooks: A, C. Anterior hooks, B. Posterior hook.
31
. Mature proglottis.
32
. Postmature proglottis.
33
. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm.
Vitellarium oval, massive, mostly irregular, 55–76 × 40–50 (68 × 46, n = 10), situated adjacent to posterior proglottis margin. Ovary central, transversely elongate, initially slightly bilobate but rapidly becoming more compact and irregular (
Figs 31–32
). Mehlis’ gland overlapping or just anterior to vitellarium. Seminal receptacle with rounded distal end and tubular and gradually tapering proximal end. Vaginal pore dorsal to pore of cirrus-sac; most of copulatory part of vagina passing posteriorly to cirrus-sac, straight; lumen encircled by thin cellular sleeve; connected with seminal receptacle by thin and short conductive part (
Fig. 32
).
Young uterus in mature proglottides well marked, with strong walls, developing antero-dorsally to ovary, initially globular (
Fig. 32
), often with some irregular incomplete septa. With increasing in size, uterus becoming more irregular, with densification surrounding its anterior wall (
Fig. 33
); with further development, uterus increasing in size; paruterine organ developing mostly anterior to it but also embracing uterus laterally. No fully developed eggs available.
Remarks
The present material unambiguously belongs to
Notopentorchis
Burt, 1938
as diagnosed by
Georgiev & Kornyushin (1994)
. This genus has recently been thoroughly reviewed and presently comprises eight species, all of them found in swifts from the Old World (
Georgiev & Bray 1991
;
Dimitrova
et al.
2017
).
Our material is unfortunately in a rather poor state of conservation and limited as neither a scolex with a full set of hooks nor fully developed gravid proglottides are available. It is, however, sufficient to allow examination of most diagnostic characters. These characters were proved to be sufficiently unique to distinguish it from congeners and to allow for a species description.
Of the eight known species, three belong to a group with rostellar hooks of clearly distinct length in each of the two rostellar rows (
Dimitrova
et al.
2017
). The present material differs from all remaining species by a larger cirrus-sac and larger suckers (
Table 5
). In addition, it differs from
N. collocaliae
Burt, 1938
by a larger scolex and
rostellum
; from
N. javanica
(Hübscher, 1937)
by fewer testes; from
N. iduncula
(Spasskii, 1946)
,
N. caffrapi
(
Mokhehle, 1951
)
, and
N. micropus
Singh, 1952
by smaller rostellar hooks (
Mokhehle 1951
;
Singh 1952
;
Baer 1959
;
Dimitrova
et al.
2017
).
So far, no species of
Notopentorchis
has been reported from
Australia
. The white-throated needletail
Hirundapus caudacutus
, hosting the worms described here, however occurs in Central and
East Asia
and winters in Eastern
Australia
(
Lepage 2017
), making this range extension logical. The single occurrence of
Notopentorchis
in this host was reported as
N. vesiculigera
(Krabbe, 1882)
by
Oshmarin (1963)
in 15 out 29 birds in the Primorye Region (South-Eastern
Russia
). Besides its European and African distribution,
N. vesiculigera
clearly differs from our material by its much larger hooks and higher number of testes. We conclude that the present material belongs to an undescribed species that we name
Notopentorchis musealis
sp. nov.