Four new species of the primitively segmented spider genus Songthela (Mesothelae, Liphistiidae) from Chongqing Municipality, China Author Chen, Zhaoyang 0000-0003-3657-974X College of Life Sciences, Hunan Normal University, Changsha 410081, Hunan Province, China. & 243936410 @ qq. com; https: // orcid. org / 0000 - 0003 - 3657 - 974 X 243936410@qq.com Author Liu, Fengxiang State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China Author Li, Daiqin 0000-0001-8269-7734 Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543, Singapore dbslidq @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 8269 - 7734 dbslidq@nus.edu.sg Author Xu, Xin 0000-0001-5632-6622 College of Life Sciences, Hunan Normal University, Changsha 410081, Hunan Province, China. & xuxin @ hunnu. edu. cn; https: // orcid. org / 0000 - 0001 - 5632 - 6622 xuxin@hunnu.edu.cn text Zootaxa 2022 2022-01-17 5091 4 546 558 journal article 2550 10.11646/zootaxa.5091.4.2 e48c79a8-4d67-4e4c-97a1-468f16bb66c3 1175-5326 5863681 E4D0CDE9-D9CF-4444-8644-263DC1F9EA88 Genus Songthela Ono, 2000 Type species. Heptathela hangzhouensis Chen, Zhang & Zhu, 1981 . Diagnosis. Males of Songthela can be distinguished from all other Heptathelinae genera by contrategulum with dense dentations ( Figs 3A, 3D , 4A, 4D , 5A, 5D , 6A, 6D ); by conductor with one or two apical spines, middle portion smooth or covered with several teeth ( Figs 3A–E , 4A–4E , 5A–5E , 6A–E ); by embolus with a wide and flat opening ( Figs 3A, 3D , 4A, 4D , 5A, 5D , 6A, 6D ). Songthela females differ from all other Heptathelinae genera by two pairs of receptacular clusters, median ones with obvious tubular genital stalks located on anterior margin of bursa copulatrix or slightly situated dorsally, lateral ones located relatively dorsolaterally ( Figs 3H–O , 4H–M , 5H–M , 6H–M ). Distribution. Southern China and northern Vietnam .