A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini) Author Fresneda, Javier Author Faille, Arnaud Author Fery, Hans Author Ribera, Ignacio text Zootaxa 2019 2019-01-04 4543 1 1 36 journal article 27714 10.11646/zootaxa.4543.1.1 22ef92b8-1d9c-4e9f-b01c-ce1f5d3faa7b 1175-5326 2617607 31ABFA6E-6126-4603-B84F-4BEC7632D1E8 Speonemadus pulchellus ( Reitter, 1885 ) stat. rest. Figs 11 , 18 , 34 , 35 Anemadus pulchellus Reitter, 1885 : 59 . Speonemadus orchesioides pulchellus (Reitter) ; Jeannel 1936 : 214 Hormosacus vandalitiae Jeannel ; Blas 1984 : 252 (partim). Speonemadus orchesioides (Fairmaire) ; Giachino & Vailati 1993 : 174 (partim). Type locality : " Sicilien " [= Sicily ], Italy ( Reitter 1885 ) . Type series . Lectotype (by present designation) ( Fig. 34 ): ♂ , " pulchellus / m. [= mihi] Sicilien " [hw Reitter], "coll. Reitter " [printed], " HOLOTYPUS 1884 / Anemadus / pulchellus Reitter " [red label, printed], " LECTOTYPUS / Anemadus / pulchellus Reitter, 1885 / Fresneda et al . des. 2018" [red label, printed] ( HNHM ); aedeagus mounted on a glue card on the same pin . Paralectotypes ( Fig. 35 ): four specimens mounted on a single pin with five cards (one of the specimens is lost), " A. pulchellus / m. Sicilien " [hw Reitter ], "155" [printed], " TYPE " [red label, printed], " MUSEUM PARIS / ( COLL . E. REITTER) / A. GROUVELLE 1913" [printed], " pulchellus / Reitt." [hw], " PARALECTOTYPUS / Anemadus / pulchellus Reitter, 1885 / Fresneda et al . des. 2018" [red label, printed] (coll. générale in MNHN ) . Diagnosis. Length 2.8–2.9 mm; elytra regularly narrowing towards apex; external face of protibiae without tubercles, with setae directly inserted in tegument; antennomeres VIII–XI longer than wide; parameres very robust, slightly longer than apex of median lobe, with rounded apex ( Fig. 18 ); apex of median lobe regularly arched ( Fig. 11 ). Molecular data place S. pulchellus as sister species of S. vandalitiae . Remarks . Speonemadus pulchellus was considered to be a subspecies of S. orchesioides by Jeannel (1936: 214) , and a synonym by Giachino & Vailati (1993) . All the studied and figured specimens of Speonemadus from Sicily (see Appendix) share a more or less similar aedeagus, which is also similar to that of the specimen from Djebel Babor figured by Giachino & Vailati (1993) as S. orchesioides , and that we consider to belong to S. pulchellus . There are some small differences in the apex of the aedeagus of the specimens from Ficuzza (Appendix), which are slightly more rounded than those of the lectotype . We consider these differences as intraspecific variation, although the study of additional material may prove the existence of consistent variation deserving taxonomic recognition. In any case, the aedeagi of the material considered under the name S. pulchellus are clearly different to that of the male identified as S. orchesioides by us (from Ammi-Moussa, see above) ( Figs 5 , 13 and 11 , 18 respectively). Distribution ( Fig. 38 ; Appendix): Italy (including Sicily ), Eastern Algeria and likely Tunisia .