The genus Harmonia (Coleoptera, Coccinellidae) in the Middle East region Author Biranvand, Amir Young Researchers and Elite Club, Khorramabad Branch, Islamic Azad University, Khorramabad, Iran Author Nedvěd, Oldřich Faculty of Science, University of South Bohemia, CZ- 37005 České Budějovice, Czech Republic & Biology Centre, Czech Academy of Sciences, Institute of Entomology, CZ- 37005 České Budějovice, Czech Republic Author Tomaszewska, Wioletta Museum and Institute of Zoology, Polish Academy of Sciences, PL- 00 - 679 Warsaw, Poland Author Al Ansi, Amin N. King Saud University Museum of Arthropods, Department of Plant Protection, King Saud University, SA- 11451 Riyadh, Saudi Arabia Author Fekrat, Lida Department of Plant Protection, Ferdowsi University of Mashhad, Mashhad, Iran Author Haghghadam, Zahra Mojib Agricultural and Natural Resources Research Center of Gilan Province, Rasht, Iran Author Khormizi, Mehdi Zare Yazd Provincial Office, Department of Environment, Iran Author Noorinahad, Sara Department of Plant Protection, Lorestan University, Iran Author Şenal, Derya Author Shakarami, Jahanshir Department of Plant Protection, Lorestan University, Iran Author Haelewaters, Danny Faculty of Science, University of South Bohemia, CZ- 37005 České Budějovice, Czech Republic text Acta Entomologica Musei Nationalis Pragae 2019 2019-04-17 59 1 163 170 journal article 8093 10.2478/aemnp-2019-0014 02d54075-6c21-4d05-b8f8-40828a70b492 1804-6487 4552749 605E5337-395B-4C88-AC75-E761330518FE Key to species of Harmonia in the Middle East 1 Body strongly convex. Pronotum white with 4 or 5 black dots in M shape, or with large trapezoid black patch in center leaving only lateral margins white. Elytra black, each elytron with 2 or 4 yellow-reddish spots; or elytra yellow to red, each elytron with maximum 10 black spots in 4 rows (2½–3–3–1) ( Figs 1–4 ). Prosternal process carinate ( Fig. 6 ). Male genitalia as in Figs 14–16 . Penis with tip afler constriction about two times longer than wide. Penis guide from lateral view with apex clearly bent upwards. Body length 6.5– 7.2 mm .................................. H. axyridis (Pallas, 1773) – Body slightly convex. Pronotum white to yellowish, with 11 black dots; 5 large in Mshape in center, 6 small laterally. Elytra yellow, each elytron with maximum 8 black spots in 4 rows (1–3–3–1) ( Fig. 5 ). Prosternal process without carinae ( Fig. 7 ). Male genitalia as in Figs 17–19 . Penis with tip afler constriction about four times longer than wide. Penis guide from lateral view straight. Body length 5.1–6.6 mm . .................................. ................. H. quadripunctata ( Pontoppidan, 1763 ) *