Revision of the enigmatic Southeast Asian spider genus Savarna (Araneae, Pholcidae)
Author
Huber, Bernhard A.
Author
Petcharad, Booppa
Author
Bumrungsri, Sara
text
European Journal of Taxonomy
2015
2015-12-10
160
1
23
journal article
http://dx.doi.org/10.5852/ejt.2015.160
65a5e6d8-d655-4949-8f4f-f5fc7930f5f4
830786
AFC4DF73-9767-4929-86F7-328ED9B65FDB
Savarna kraburiensis
Wongprom & Wiwatwitaya, 2015
Figs 5–7
,
44–54
Savarna kraburiensis
Wongprom & Wiwatwitaya, 2015: 2–5
, Fgs 1–2 (
Ƌ
♀
).
Diagnosis
Easily distinguished from known congeners by morphology of male palp (bulb with two long processes; tip of procursus;
Figs 49–50
), and by female external and internal genitalia (epigynal plate with median process;
Fig. 54
). From
S. kaeo
sp. nov.
also distinguished by absence of pair of processes on male clypeus (
Figs 44–45
); from
S. tessellata
and
S. miser
also distinguished by black marks laterally on carapace (
Figs 5–6
).
Type
material
THAILAND
:
Ƌ
,
holotype
;
1 Ƌ
,
2 ♀♀
,
paratypes
;
Ranong
,
Kraburi District
,
Phra Kha Yang Cave
(
10°19.57’ N
,
98°45.91’ E
),
6 m
a.s.l.,
28 Oct. 2014
,
P. Wongprom
leg., Thailand Natural History Museum, Pathum Thani, not examined
.
Material examined
THAILAND
:
8 ƋƋ
,
5 ♀♀
,
Ranong
,
Kraburi District
,
Tham Phra Kayang
[=
Phra Kha Yang Cave
] (
10°19.54’ N
,
98°45.88’ E
),
10–50 m
a.s.l.,
in cave and in forest above cave
,
13 Mar. 2015
,
B.A. Huber & B. Petcharad
leg.,
ZFMK
(
Ar 12991, 12992
)
;
3 ƋƋ
,
3 ♀♀
, same data (
PSUZC
)
;
4 ♀♀
,
2 juvs
, in pure ethanol, same data,
ZFMK
(
Mal 365
)
.
Amendments to original description
Male clypeus without processes but with strong hair brushes not present in female (
Figs 44–46
). Male gonopore without epiandrous spigots (
Fig. 52
). Male and female ALS with only two spigots each (
Fig. 53
). In the palp illustrated in the original description (
Wongprom & Wiwatwitaya 2015: Fg. 1a
) the bulb is rotated about 180° from its natural position. In the natural position, the long pointed process is directed in the opposite direction and the proximal bulbal sclerite is visible in prolateral view. Spines on male legs sometimes absent, sometimes present (each femur 1 with two ventral rows of up to ~25 spines each); tibia 1 L/d: 53; prolateral trichobothrium absent on tibia 1, present on other tibiae; male and female tarsus 4 with single row of ventral comb-hairs (
Fig. 51
). Tibia
1 in
10 males
: 5.7–6.8 (mean: 6.3); in
5 females
: 4.7–5.6 (mean 5.3).
Figs 14–18.
Savarna kaeo
sp. nov.
14–15
. Left male palp, prolateral and retrolateral views (b = genital bulb; bp = bulbal process; pr = procursus; tr = trochanter).
16
. Male chelicerae, frontal view.
17–18
. Cleared female genitalia, ventral and dorsal views. Scale bars: 16 = 0.3 mm; 14–15, 17–18 = 0.5 mm.
Figs 19–27.
Savarna kaeo
sp. nov.
19–20
. Male prosoma, frontal and oblique frontal views.
21
. Right male palpal trochanter (tr) and femur, retrolateral view.
22
. Left procursus (pr) and bulbal process (bp), prolatero-distal view (b = bulb).
23
. Right procursus (pr) and bulbal process (bp), retrolatero-distal view (b = bulb).
24
. Female ALS.
25
. Male palpal tarsal organ.
26
. Male gonopore.
27
. Epigynum, ventrolateral view. Scale bars: 24–25 = 10 µm; 26 = 80 µm; 21–23 = 100 µm; 19–20, 27 = 200 µm.
Natural history
Spiders were found both inside and outside the cave, in domed webs of ~
20 cm
diameter. In the cave, they only occurred in the twilight zone. Outside the cave, they were found to be most abundant in the small forest above the cave. When lightly disturbed, the spiders vibrated in their webs; when disturbed more strongly, spiders dropped to the ground, remaining motionless (
cf.
Fig. 7
).
Distribution
Known only from the
type
locality (
Fig. 1
).